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Hopliinae Systellopodinae

Mating Rhizotrogon marginipes (Melolonthini/Rhizotrogini), male on top – note sexually dimorphic antennae

Melolonthinae
Melolonthinae
is a subfamily of the scarab beetles (family Scarabaeidae). It is a very diverse group; distributed over most of the world, it contains many familiar species. Some authors include the scarab subfamilies Euchirinae
Euchirinae
and Pachypodinae as tribes in the Melolonthinae. Unlike some of their relatives, their habitus is usually not bizarre. They resemble the Rutelinae
Rutelinae
in being fairly plesiomorphic in outward appearance. Like in many Scarabaeidae, males have large fingered antennae, while those of the females are smaller and somewhat knobby. In the Melolonthinae, this sexual dimorphism is particularly pronounced. Many species have striking – though rarely brilliant or iridescent – hues and bold patterns of hairs. Being often quite sizeable and swarming in numbers at certain times, for example the Amphimallon, Phyllophaga and Polyphylla
Polyphylla
"june beetles" or the Melolontha
Melolontha
cockchafers – all from tribe Melolonthini
Melolonthini
– feature widely in folklore. Some Melolonthinae
Melolonthinae
are economically significant pests. Other than the Melolonthini, the most diverse tribes are the Ablaberini, Liparetrini, Macrodactylini, Pachydemini and Sericini.

Systematics[edit]

Phyllotocus sp. (Sericini)

Stethaspis sp. (Stethaspini/Xylonychini)

According to various authors, the living Melolonthinae
Melolonthinae
are divided into about 20-30 tribes. Some notable genera and species are also listed here:

Ablaberini Burmeister, 1855 – including Camentini Automoliini Britton, 1978 Chasmatopterini Lacordaire, 1856 Colymbomorphini – sometimes in Xylonychini Comophorinini Britton, 1957 – including Comophini Dichelonychini Burmeister 1855 Diphucephalini Britton, 1957 Diphycerini – sometimes in Macrodactylini Diplotaxini – sometimes in Melolonthini Heteronychini Britton, 1957

Heteronyx Guérin-Ménéville, 1838

Hopliini Lichniini Liparetrini Burmeister, 1855 – including Allarini, Colpochilini Macrodactylini Kirby, 1837[verification needed] – including Dichelonyciini Maechidiini Burmeister, 1855 Melolonthini
Melolonthini
Samouelle, 1819[verification needed]

Amphimallon Leucopholis Melolontha
Melolontha
– cockchafers, May bugs Miridiba Phyllophaga – May beetles Polyphylla Sophrops

Oncerini Pachydemini Reitter, 1902 Pachytrichini Burmeister, 1855 Phyllotocidiini Britton, 1957 Podolasiini – sometimes in Hopliini Rhizotrogini – sometimes in Melolonthini

Holotrichia Rhizotrogus

Scitalini Britton, 1957 Sericini Dalla Torre, 1912

Maladera

Sericoidini Burmeister, 1855 Stethaspini – sometimes in Xylonychini Systellopini Sharp, 1877 Tanyproctini – sometimes in Pachydemini Xylonychini Britton, 1957

In addition, a prehistoric tribe, the Cretomelolonthini, is only known from fossils. Several genera are of unclear relations; they are not yet firmly placed in a tribe:

Acoma Conebius Fuavel, 1903 Costelytra – Liparetrini? Hemictenius – Pachydemini? Metascelis Westwood, 1842 Mycernus – Colymbomorphini, Stethaspini, Xylonychini?

Odontria – Liparetrini? Prodontria – Liparetrini? Psilodontria – Colymbomorphini, Stethaspini, Xylonychini? Scythrodes – Liparetrini? Sericospilus – Liparetrini?

"Anonetus" and "Tryssus", both used by Erichson in 1847, are nomina nuda. Holophylla and Hoplorida are of uncertain validity. Whether the scarab beetle Xenaclopus belongs in the present subfamily is doubtful. Alosimus, generally held to be a blister beetle of the Lyttini
Lyttini
and not at all close to the scarabs, was placed in the Melolonthinae
Melolonthinae
by entomologist Werner Heinz Muche. References[edit] Data related to Melolonthinae
Melolonthinae
at Wikispecies Media related to Melolonthinae
Melolonthinae
at Wikimedia Commons

http://insects.tamu.edu/research/collection/hallan/test/Arthropoda/Insects/Coleoptera/Family/Scarabaeidae.txt

Taxon identifiers

Wd: Q785080 BugGuide: 2951 Fauna Europaea: 257338 Fossilworks: 70051 ITIS: 678508 NCBI: 7059 NZOR: 70817595-f753-46ac

.
Melolonthinae
HOME
The Info List - Melolonthinae


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Hopliinae Systellopodinae

Mating Rhizotrogon marginipes (Melolonthini/Rhizotrogini), male on top – note sexually dimorphic antennae

Melolonthinae
Melolonthinae
is a subfamily of the scarab beetles (family Scarabaeidae). It is a very diverse group; distributed over most of the world, it contains many familiar species. Some authors include the scarab subfamilies Euchirinae
Euchirinae
and Pachypodinae as tribes in the Melolonthinae. Unlike some of their relatives, their habitus is usually not bizarre. They resemble the Rutelinae
Rutelinae
in being fairly plesiomorphic in outward appearance. Like in many Scarabaeidae, males have large fingered antennae, while those of the females are smaller and somewhat knobby. In the Melolonthinae, this sexual dimorphism is particularly pronounced. Many species have striking – though rarely brilliant or iridescent – hues and bold patterns of hairs. Being often quite sizeable and swarming in numbers at certain times, for example the Amphimallon, Phyllophaga and Polyphylla
Polyphylla
"june beetles" or the Melolontha
Melolontha
cockchafers – all from tribe Melolonthini
Melolonthini
– feature widely in folklore. Some Melolonthinae
Melolonthinae
are economically significant pests. Other than the Melolonthini, the most diverse tribes are the Ablaberini, Liparetrini, Macrodactylini, Pachydemini and Sericini.

Systematics[edit]

Phyllotocus sp. (Sericini)

Stethaspis sp. (Stethaspini/Xylonychini)

According to various authors, the living Melolonthinae
Melolonthinae
are divided into about 20-30 tribes. Some notable genera and species are also listed here:

Ablaberini Burmeister, 1855 – including Camentini Automoliini Britton, 1978 Chasmatopterini Lacordaire, 1856 Colymbomorphini – sometimes in Xylonychini Comophorinini Britton, 1957 – including Comophini Dichelonychini Burmeister 1855 Diphucephalini Britton, 1957 Diphycerini – sometimes in Macrodactylini Diplotaxini – sometimes in Melolonthini Heteronychini Britton, 1957

Heteronyx Guérin-Ménéville, 1838

Hopliini Lichniini Liparetrini Burmeister, 1855 – including Allarini, Colpochilini Macrodactylini Kirby, 1837[verification needed] – including Dichelonyciini Maechidiini Burmeister, 1855 Melolonthini
Melolonthini
Samouelle, 1819[verification needed]

Amphimallon Leucopholis Melolontha
Melolontha
– cockchafers, May bugs Miridiba Phyllophaga – May beetles Polyphylla Sophrops

Oncerini Pachydemini Reitter, 1902 Pachytrichini Burmeister, 1855 Phyllotocidiini Britton, 1957 Podolasiini – sometimes in Hopliini Rhizotrogini – sometimes in Melolonthini

Holotrichia Rhizotrogus

Scitalini Britton, 1957 Sericini Dalla Torre, 1912

Maladera

Sericoidini Burmeister, 1855 Stethaspini – sometimes in Xylonychini Systellopini Sharp, 1877 Tanyproctini – sometimes in Pachydemini Xylonychini Britton, 1957

In addition, a prehistoric tribe, the Cretomelolonthini, is only known from fossils. Several genera are of unclear relations; they are not yet firmly placed in a tribe:

Acoma Conebius Fuavel, 1903 Costelytra – Liparetrini? Hemictenius – Pachydemini? Metascelis Westwood, 1842 Mycernus – Colymbomorphini, Stethaspini, Xylonychini?

Odontria – Liparetrini? Prodontria – Liparetrini? Psilodontria – Colymbomorphini, Stethaspini, Xylonychini? Scythrodes – Liparetrini? Sericospilus – Liparetrini?

"Anonetus" and "Tryssus", both used by Erichson in 1847, are nomina nuda. Holophylla and Hoplorida are of uncertain validity. Whether the scarab beetle Xenaclopus belongs in the present subfamily is doubtful. Alosimus, generally held to be a blister beetle of the Lyttini
Lyttini
and not at all close to the scarabs, was placed in the Melolonthinae
Melolonthinae
by entomologist Werner Heinz Muche. References[edit] Data related to Melolonthinae
Melolonthinae
at Wikispecies Media related to Melolonthinae
Melolonthinae
at Wikimedia Commons

http://insects.tamu.edu/research/collection/hallan/test/Arthropoda/Insects/Coleoptera/Family/Scarabaeidae.txt

Taxon identifiers

Wd: Q785080 BugGuide: 2951 Fauna Europaea: 257338 Fossilworks: 70051 ITIS: 678508 NCBI: 7059 NZOR: 70817595-f753-46ac

.
Melolonthinae
HOME
The Info List - Melolonthinae


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Hopliinae Systellopodinae

Mating Rhizotrogon marginipes (Melolonthini/Rhizotrogini), male on top – note sexually dimorphic antennae

Melolonthinae
Melolonthinae
is a subfamily of the scarab beetles (family Scarabaeidae). It is a very diverse group; distributed over most of the world, it contains many familiar species. Some authors include the scarab subfamilies Euchirinae
Euchirinae
and Pachypodinae as tribes in the Melolonthinae. Unlike some of their relatives, their habitus is usually not bizarre. They resemble the Rutelinae
Rutelinae
in being fairly plesiomorphic in outward appearance. Like in many Scarabaeidae, males have large fingered antennae, while those of the females are smaller and somewhat knobby. In the Melolonthinae, this sexual dimorphism is particularly pronounced. Many species have striking – though rarely brilliant or iridescent – hues and bold patterns of hairs. Being often quite sizeable and swarming in numbers at certain times, for example the Amphimallon, Phyllophaga and Polyphylla
Polyphylla
"june beetles" or the Melolontha
Melolontha
cockchafers – all from tribe Melolonthini
Melolonthini
– feature widely in folklore. Some Melolonthinae
Melolonthinae
are economically significant pests. Other than the Melolonthini, the most diverse tribes are the Ablaberini, Liparetrini, Macrodactylini, Pachydemini and Sericini.

Systematics[edit]

Phyllotocus sp. (Sericini)

Stethaspis sp. (Stethaspini/Xylonychini)

According to various authors, the living Melolonthinae
Melolonthinae
are divided into about 20-30 tribes. Some notable genera and species are also listed here:

Ablaberini Burmeister, 1855 – including Camentini Automoliini Britton, 1978 Chasmatopterini Lacordaire, 1856 Colymbomorphini – sometimes in Xylonychini Comophorinini Britton, 1957 – including Comophini Dichelonychini Burmeister 1855 Diphucephalini Britton, 1957 Diphycerini – sometimes in Macrodactylini Diplotaxini – sometimes in Melolonthini Heteronychini Britton, 1957

Heteronyx Guérin-Ménéville, 1838

Hopliini Lichniini Liparetrini Burmeister, 1855 – including Allarini, Colpochilini Macrodactylini Kirby, 1837[verification needed] – including Dichelonyciini Maechidiini Burmeister, 1855 Melolonthini
Melolonthini
Samouelle, 1819[verification needed]

Amphimallon Leucopholis Melolontha
Melolontha
– cockchafers, May bugs Miridiba Phyllophaga – May beetles Polyphylla Sophrops

Oncerini Pachydemini Reitter, 1902 Pachytrichini Burmeister, 1855 Phyllotocidiini Britton, 1957 Podolasiini – sometimes in Hopliini Rhizotrogini – sometimes in Melolonthini

Holotrichia Rhizotrogus

Scitalini Britton, 1957 Sericini Dalla Torre, 1912

Maladera

Sericoidini Burmeister, 1855 Stethaspini – sometimes in Xylonychini Systellopini Sharp, 1877 Tanyproctini – sometimes in Pachydemini Xylonychini Britton, 1957

In addition, a prehistoric tribe, the Cretomelolonthini, is only known from fossils. Several genera are of unclear relations; they are not yet firmly placed in a tribe:

Acoma Conebius Fuavel, 1903 Costelytra – Liparetrini? Hemictenius – Pachydemini? Metascelis Westwood, 1842 Mycernus – Colymbomorphini, Stethaspini, Xylonychini?

Odontria – Liparetrini? Prodontria – Liparetrini? Psilodontria – Colymbomorphini, Stethaspini, Xylonychini? Scythrodes – Liparetrini? Sericospilus – Liparetrini?

"Anonetus" and "Tryssus", both used by Erichson in 1847, are nomina nuda. Holophylla and Hoplorida are of uncertain validity. Whether the scarab beetle Xenaclopus belongs in the present subfamily is doubtful. Alosimus, generally held to be a blister beetle of the Lyttini
Lyttini
and not at all close to the scarabs, was placed in the Melolonthinae
Melolonthinae
by entomologist Werner Heinz Muche. References[edit] Data related to Melolonthinae
Melolonthinae
at Wikispecies Media related to Melolonthinae
Melolonthinae
at Wikimedia Commons

http://insects.tamu.edu/research/collection/hallan/test/Arthropoda/Insects/Coleoptera/Family/Scarabaeidae.txt

Taxon identifiers

Wd: Q785080 BugGuide: 2951 Fauna Europaea: 257338 Fossilworks: 70051 ITIS: 678508 NCBI: 7059 NZOR: 70817595-f753-46ac

.
Melolonthinae
HOME
The Info List - Melolonthinae


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Hopliinae Systellopodinae

Mating Rhizotrogon marginipes (Melolonthini/Rhizotrogini), male on top – note sexually dimorphic antennae

Melolonthinae
Melolonthinae
is a subfamily of the scarab beetles (family Scarabaeidae). It is a very diverse group; distributed over most of the world, it contains many familiar species. Some authors include the scarab subfamilies Euchirinae
Euchirinae
and Pachypodinae as tribes in the Melolonthinae. Unlike some of their relatives, their habitus is usually not bizarre. They resemble the Rutelinae
Rutelinae
in being fairly plesiomorphic in outward appearance. Like in many Scarabaeidae, males have large fingered antennae, while those of the females are smaller and somewhat knobby. In the Melolonthinae, this sexual dimorphism is particularly pronounced. Many species have striking – though rarely brilliant or iridescent – hues and bold patterns of hairs. Being often quite sizeable and swarming in numbers at certain times, for example the Amphimallon, Phyllophaga and Polyphylla
Polyphylla
"june beetles" or the Melolontha
Melolontha
cockchafers – all from tribe Melolonthini
Melolonthini
– feature widely in folklore. Some Melolonthinae
Melolonthinae
are economically significant pests. Other than the Melolonthini, the most diverse tribes are the Ablaberini, Liparetrini, Macrodactylini, Pachydemini and Sericini.

Systematics[edit]

Phyllotocus sp. (Sericini)

Stethaspis sp. (Stethaspini/Xylonychini)

According to various authors, the living Melolonthinae
Melolonthinae
are divided into about 20-30 tribes. Some notable genera and species are also listed here:

Ablaberini Burmeister, 1855 – including Camentini Automoliini Britton, 1978 Chasmatopterini Lacordaire, 1856 Colymbomorphini – sometimes in Xylonychini Comophorinini Britton, 1957 – including Comophini Dichelonychini Burmeister 1855 Diphucephalini Britton, 1957 Diphycerini – sometimes in Macrodactylini Diplotaxini – sometimes in Melolonthini Heteronychini Britton, 1957

Heteronyx Guérin-Ménéville, 1838

Hopliini Lichniini Liparetrini Burmeister, 1855 – including Allarini, Colpochilini Macrodactylini Kirby, 1837[verification needed] – including Dichelonyciini Maechidiini Burmeister, 1855 Melolonthini
Melolonthini
Samouelle, 1819[verification needed]

Amphimallon Leucopholis Melolontha
Melolontha
– cockchafers, May bugs Miridiba Phyllophaga – May beetles Polyphylla Sophrops

Oncerini Pachydemini Reitter, 1902 Pachytrichini Burmeister, 1855 Phyllotocidiini Britton, 1957 Podolasiini – sometimes in Hopliini Rhizotrogini – sometimes in Melolonthini

Holotrichia Rhizotrogus

Scitalini Britton, 1957 Sericini Dalla Torre, 1912

Maladera

Sericoidini Burmeister, 1855 Stethaspini – sometimes in Xylonychini Systellopini Sharp, 1877 Tanyproctini – sometimes in Pachydemini Xylonychini Britton, 1957

In addition, a prehistoric tribe, the Cretomelolonthini, is only known from fossils. Several genera are of unclear relations; they are not yet firmly placed in a tribe:

Acoma Conebius Fuavel, 1903 Costelytra – Liparetrini? Hemictenius – Pachydemini? Metascelis Westwood, 1842 Mycernus – Colymbomorphini, Stethaspini, Xylonychini?

Odontria – Liparetrini? Prodontria – Liparetrini? Psilodontria – Colymbomorphini, Stethaspini, Xylonychini? Scythrodes – Liparetrini? Sericospilus – Liparetrini?

"Anonetus" and "Tryssus", both used by Erichson in 1847, are nomina nuda. Holophylla and Hoplorida are of uncertain validity. Whether the scarab beetle Xenaclopus belongs in the present subfamily is doubtful. Alosimus, generally held to be a blister beetle of the Lyttini
Lyttini
and not at all close to the scarabs, was placed in the Melolonthinae
Melolonthinae
by entomologist Werner Heinz Muche. References[edit] Data related to Melolonthinae
Melolonthinae
at Wikispecies Media related to Melolonthinae
Melolonthinae
at Wikimedia Commons

http://insects.tamu.edu/research/collection/hallan/test/Arthropoda/Insects/Coleoptera/Family/Scarabaeidae.txt

Taxon identifiers

Wd: Q785080 BugGuide: 2951 Fauna Europaea: 257338 Fossilworks: 70051 ITIS: 678508 NCBI: 7059 NZOR: 70817595-f753-46ac

.
l> Melolonthinae


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Hopliinae Systellopodinae

Mating Rhizotrogon marginipes (Melolonthini/Rhizotrogini), male on top – note sexually dimorphic antennae

Melolonthinae
Melolonthinae
is a subfamily of the scarab beetles (family Scarabaeidae). It is a very diverse group; distributed over most of the world, it contains many familiar species. Some authors include the scarab subfamilies Euchirinae
Euchirinae
and Pachypodinae as tribes in the Melolonthinae. Unlike some of their relatives, their habitus is usually not bizarre. They resemble the Rutelinae
Rutelinae
in being fairly plesiomorphic in outward appearance. Like in many Scarabaeidae, males have large fingered antennae, while those of the females are smaller and somewhat knobby. In the Melolonthinae, this sexual dimorphism is particularly pronounced. Many species have striking – though rarely brilliant or iridescent – hues and bold patterns of hairs. Being often quite sizeable and swarming in numbers at certain times, for example the Amphimallon, Phyllophaga and Polyphylla
Polyphylla
"june beetles" or the Melolontha
Melolontha
cockchafers – all from tribe Melolonthini
Melolonthini
– feature widely in folklore. Some Melolonthinae
Melolonthinae
are economically significant pests. Other than the Melolonthini, the most diverse tribes are the Ablaberini, Liparetrini, Macrodactylini, Pachydemini and Sericini.

Systematics[edit]

Phyllotocus sp. (Sericini)

Stethaspis sp. (Stethaspini/Xylonychini)

According to various authors, the living Melolonthinae
Melolonthinae
are divided into about 20-30 tribes. Some notable genera and species are also listed here:

Ablaberini Burmeister, 1855 – including Camentini Automoliini Britton, 1978 Chasmatopterini Lacordaire, 1856 Colymbomorphini – sometimes in Xylonychini Comophorinini Britton, 1957 – including Comophini Dichelonychini Burmeister 1855 Diphucephalini Britton, 1957 Diphycerini – sometimes in Macrodactylini Diplotaxini – sometimes in Melolonthini Heteronychini Britton, 1957

Heteronyx Guérin-Ménéville, 1838

Hopliini Lichniini Liparetrini Burmeister, 1855 – including Allarini, Colpochilini Macrodactylini Kirby, 1837[verification needed] – including Dichelonyciini Maechidiini Burmeister, 1855 Melolonthini
Melolonthini
Samouelle, 1819[verification needed]

Amphimallon Leucopholis Melolontha
Melolontha
– cockchafers, May bugs Miridiba Phyllophaga – May beetles Polyphylla Sophrops

Oncerini Pachydemini Reitter, 1902 Pachytrichini Burmeister, 1855 Phyllotocidiini Britton, 1957 Podolasiini – sometimes in Hopliini Rhizotrogini – sometimes in Melolonthini

Holotrichia Rhizotrogus

Scitalini Britton, 1957 Sericini Dalla Torre, 1912

Maladera

Sericoidini Burmeister, 1855 Stethaspini – sometimes in Xylonychini Systellopini Sharp, 1877 Tanyproctini – sometimes in Pachydemini Xylonychini Britton, 1957

In addition, a prehistoric tribe, the Cretomelolonthini, is only known from fossils. Several genera are of unclear relations; they are not yet firmly placed in a tribe:

Acoma Conebius Fuavel, 1903 Costelytra – Liparetrini? Hemictenius – Pachydemini? Metascelis Westwood, 1842 Mycernus – Colymbomorphini, Stethaspini, Xylonychini?

Odontria – Liparetrini? Prodontria – Liparetrini? Psilodontria – Colymbomorphini, Stethaspini, Xylonychini? Scythrodes – Liparetrini? Sericospilus – Liparetrini?

"Anonetus" and "Tryssus", both used by Erichson in 1847, are nomina nuda. Holophylla and Hoplorida are of uncertain validity. Whether the scarab beetle Xenaclopus belongs in the present subfamily is doubtful. Alosimus, generally held to be a blister beetle of the Lyttini
Lyttini
and not at all close to the scarabs, was placed in the Melolonthinae
Melolonthinae
by entomologist Werner Heinz Muche. References[edit] Data related to Melolonthinae
Melolonthinae
at Wikispecies Media related to Melolonthinae
Melolonthinae
at Wikimedia Commons

http://insects.tamu.edu/research/collection/hallan/test/Arthropoda/Insects/Coleoptera/Family/Scarabaeidae.txt

Taxon identifiers

Wd: Q785080 BugGuide: 2951 Fauna Europaea: 257338 Fossilworks: 70051 ITIS: 678508 NCBI: 7059 NZOR: 70817595-f753-46ac

.

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