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biology Biology is the scientific study of life. It is a natural science with a broad scope but has several unifying themes that tie it together as a single, coherent field. For instance, all organisms are made up of cells that process hereditary i ...
, mating is the pairing of either opposite-
sex Sex is the trait that determines whether a sexually reproducing animal or plant produces male or female gametes. Male plants and animals produce smaller mobile gametes (spermatozoa, sperm, pollen), while females produce larger ones ( ova, of ...
or
hermaphroditic In reproductive biology, a hermaphrodite () is an organism that has both kinds of reproductive organs and can produce both gametes associated with male and female sexes. Many taxonomic groups of animals (mostly invertebrates) do not have s ...
organism In biology, an organism () is any living system that functions as an individual entity. All organisms are composed of cells (cell theory). Organisms are classified by taxonomy into groups such as multicellular animals, plants, and ...
s for the purposes of
sexual reproduction Sexual reproduction is a type of reproduction that involves a complex life cycle in which a gamete ( haploid reproductive cells, such as a sperm or egg cell) with a single set of chromosomes combines with another gamete to produce a zygote th ...
. ''
Fertilization Fertilisation or fertilization (see spelling differences), also known as generative fertilisation, syngamy and impregnation, is the fusion of gametes to give rise to a new individual organism or offspring and initiate its development. Proce ...
'' is the fusion of two
gametes A gamete (; , ultimately ) is a haploid cell that fuses with another haploid cell during fertilization in organisms that reproduce sexually. Gametes are an organism's reproductive cells, also referred to as sex cells. In species that produce ...
. ''
Copulation Sexual intercourse (or coitus or copulation) is a sexual activity typically involving the insertion and thrusting of the penis into the vagina for sexual pleasure or reproduction.Sexual intercourse most commonly means penile–vaginal penetra ...
'' is the union of the
sex organ A sex organ (or reproductive organ) is any part of an animal or plant that is involved in sexual reproduction. The reproductive organs together constitute the reproductive system. In animals, the testis in the male, and the ovary in the female, a ...
s of two sexually reproducing animals for
insemination Insemination is the introduction of sperm into a female’s reproductive system for the purpose of impregnating, also called fertilizing, the female for sexual reproduction. The sperm is introduced into the uterus of a mammal or the oviduct of a ...
and subsequent
internal fertilization Internal fertilization is the union of an egg and sperm cell during sexual reproduction inside the female body. Internal fertilization, unlike its counterpart, external fertilization, brings more control to the female with reproduction. For inte ...
. Mating may also lead to
external fertilization External fertilization is a mode of reproduction in which a male organism's sperm fertilizes a female organism's egg outside of the female's body. It is contrasted with internal fertilization, in which sperm are introduced via insemination and then ...
, as seen in amphibians,
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of ...
es and plants. For most species, mating is between two individuals of opposite sexes. However, for some hermaphroditic species, copulation is not required because the parent organism is capable of self-fertilization (
autogamy Autogamy, or self-fertilization, refers to the fusion of two gametes that come from one individual. Autogamy is predominantly observed in the form of self-pollination, a reproductive mechanism employed by many flowering plants. However, species o ...
); for example,
banana slug Banana slugs are North American terrestrial slugs comprising the genus ''Ariolimax''. MolluscaBase eds. (2021). MolluscaBase. Ariolimax Mörch, 1859. Accessed through: World Register of Marine Species at: http://www.marinespecies.org/aphia.php?p= ...
s. The term ''mating'' is also applied to related processes in
bacteria Bacteria (; singular: bacterium) are ubiquitous, mostly free-living organisms often consisting of one Cell (biology), biological cell. They constitute a large domain (biology), domain of prokaryotic microorganisms. Typically a few micrometr ...
, archaea and
virus A virus is a submicroscopic infectious agent that replicates only inside the living cells of an organism. Viruses infect all life forms, from animals and plants to microorganisms, including bacteria and archaea. Since Dmitri Ivanovsk ...
es. Mating in these cases involves the pairing of individuals, accompanied by the pairing of their
homologous chromosomes A couple of homologous chromosomes, or homologs, are a set of one maternal and one paternal chromosome that pair up with each other inside a cell during fertilization. Homologs have the same genes in the same loci where they provide points alon ...
and then exchange of genomic information leading to formation of recombinant progeny (see
mating systems A mating system is a way in which a group is structured in relation to sexual behaviour. The precise meaning depends upon the context. With respect to animals, the term describes which males and females mate under which circumstances. Recognised ...
).


Animals

For animals, mating strategies include
random mating Panmixia (or panmixis) means random mating. A panmictic population is one where all individuals are potential partners. This assumes that there are no mating restrictions, neither genetic nor behavioural, upon the population and that therefore all ...
,
disassortative mating Disassortative mating (also known as negative assortative mating or heterogamy) is a mating pattern in which individuals with dissimilar phenotypes mate with one another more frequently than would be expected under random mating. Disassortative mat ...
,
assortative mating Assortative mating (also referred to as positive assortative mating or homogamy) is a mating pattern and a form of sexual selection in which individuals with similar phenotypes or genotypes mate with one another more frequently than would be ex ...
, or a
mating pool A mating pool is a concept used in evolutionary computation, which refers to a family of algorithms used to solve optimization and search problems.Regupathi, R. “Cost Optimization Of Multistoried Rc Framed Structure Using Hybrid Genetic Algorithm ...
. In some birds, it includes behaviors such as
nest A nest is a structure built for certain animals to hold eggs or young. Although nests are most closely associated with birds, members of all classes of vertebrates and some invertebrates construct nests. They may be composed of organic materi ...
-building and feeding offspring. The human practice of mating and artificially inseminating domesticated animals is part of animal husbandry. In some
terrestrial Terrestrial refers to things related to land or the planet Earth. Terrestrial may also refer to: * Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on or near the ground, as opposed to ...
arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and cuticle made of chiti ...
s, including
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three ...
s representing basal (primitive)
phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
clades, the male deposits
spermatozoa A spermatozoon (; also spelled spermatozoön; ; ) is a motile sperm cell, or moving form of the haploid cell that is the male gamete. A spermatozoon joins an ovum to form a zygote. (A zygote is a single cell, with a complete set of chromos ...
on the substrate, sometimes stored within a special structure.
Courtship Courtship is the period wherein some couples get to know each other prior to a possible marriage. Courtship traditionally may begin after a betrothal and may conclude with the celebration of marriage. A courtship may be an informal and private m ...
involves inducing the female to take up the sperm package into her genital opening without actual copulation. Courtship is often facilitated through forming groups, called
leks A lek is an aggregation of male animals gathered to engage in competitive displays and courtship rituals, known as lekking, to entice visiting females which are surveying prospective partners with which to mate. A lek can also indicate an avail ...
, in flies and many other insects. For example, male ''
Tokunagayusurika akamusi ''Tokunagayusurika akamusi'' is a midge fly species of the family ''Chironomidae'', commonly called "nonbiting midges" or "lake flies." Discovered by Tokunaga in 1938, the species is common to eutrophic lakes in Japan. The midge family is very ...
'' forms swarms dancing in the air to attract females. In groups such as
dragonflies A dragonfly is a flying insect belonging to the infraorder Anisoptera below the order Odonata. About 3,000 extant species of true dragonfly are known. Most are tropical, with fewer species in temperate regions. Loss of wetland habitat threa ...
and many
spider Spiders ( order Araneae) are air-breathing arthropods that have eight legs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species ...
s, males extrude sperm into secondary copulatory structures removed from their genital opening, which are then used to inseminate the female (in dragonflies, it is a set of modified sternites on the second abdominal segment; in spiders, it is the male
pedipalp Pedipalps (commonly shortened to palps or palpi) are the second pair of appendages of chelicerates – a group of arthropods including spiders, scorpions, horseshoe crabs, and sea spiders. The pedipalps are lateral to the chelicerae ("jaws") an ...
s). In advanced groups of insects, the male uses its aedeagus, a structure formed from the terminal segments of the abdomen, to deposit sperm directly (though sometimes in a capsule called a "
spermatophore A spermatophore or sperm ampulla is a capsule or mass containing spermatozoa created by males of various animal species, especially salamanders and arthropods, and transferred in entirety to the female's ovipore during reproduction. Spermatophores ...
") into the female's reproductive tract. Other animals reproduce sexually with external fertilization, including many basal
vertebrate Vertebrates () comprise all animal taxa within the subphylum Vertebrata () (chordates with backbones), including all mammals, birds, reptiles, amphibians, and fish. Vertebrates represent the overwhelming majority of the phylum Chordata, with c ...
s. Vertebrates (such as reptiles, some
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of ...
, and most
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
s) reproduce with internal fertilization through cloacal copulation (see also
hemipenis A hemipenis (plural hemipenes) is one of a pair of intromittent organs of male squamates (snakes, lizards and worm lizards). Hemipenes are usually held inverted within the body, and are everted for reproduction via erectile tissue, much like ...
), while mammals copulate vaginally. In domesticated animals there are various type of mating methods being employed to mate animals like Pen Mating (when female is moved to the desired male into a pen) or paddock mating (where one male is let loose in the paddock with several females). File:Bagmati River, Pashupatinath, Nepal Animal sex バグマティ川とパシュパティナート火葬場 サルの交尾 5835.JPG,
Macaque monkey The macaques () constitute a genus (''Macaca'') of gregarious Old World monkeys of the subfamily Cercopithecinae. The 23 species of macaques inhabit ranges throughout Asia, North Africa, and (in one instance) Gibraltar. Macaques are principall ...
s mating File:オオカミ(Gray wolf) (5339403526).jpg, Gray wolves mating File:LionsMating.jpg, Lions mating File:Snails mating.jpg,
Hermaphroditic In reproductive biology, a hermaphrodite () is an organism that has both kinds of reproductive organs and can produce both gametes associated with male and female sexes. Many taxonomic groups of animals (mostly invertebrates) do not have s ...
snails (''
Cornu aspersum ''Cornu aspersum'' (syn. ''Cryptomphalus aspersus''), known by the common name garden snail, is a species of land snail in the family Helicidae, which includes some of the most familiar land snails. Of all terrestrial molluscs, this species may ...
'')
mating In biology, mating is the pairing of either opposite- sex or hermaphroditic organisms for the purposes of sexual reproduction. ''Fertilization'' is the fusion of two gametes. ''Copulation'' is the union of the sex organs of two sexually reprod ...
File:Tortoise mating.jpg, African spurred tortoises (''
Centrochelys sulcata The African spurred tortoise (''Centrochelys sulcata''), also called the sulcata tortoise, is a species of tortoise inhabiting the southern edge of the Sahara desert in Africa. It is the largest mainland species of tortoise in the world, and the ...
'') mating File:Chalkhill blue butterflies (Polyommatus coridon) mating 1.jpg, Chalkhill blue butterflies ('' Lysandra coridon'') mating File:Hoverflies mating midair.jpg,
Hoverflies Hover flies, also called flower flies or syrphid flies, make up the insect family Syrphidae. As their common name suggests, they are often seen hovering or nectaring at flowers; the adults of many species feed mainly on nectar and pollen, while ...
('' Simosyrphus grandicornis'') mating in midair File:Joined moths.JPG, Poplar hawk-moths ('' Laothoe populi'') mating File:Ladybird-Coccinellidae-mating.jpg,
Ladybug Coccinellidae () is a widespread family of small beetles ranging in size from . They are commonly known as ladybugs in North America and ladybirds in Great Britain. Some entomologists prefer the names ladybird beetles or lady beetles as they ...
s mating File:Aphrophora alni mating.jpg, Spittlebugs ('' Aphrophora alni'') mating File:Dogs mating.jpg, Dogs mating File:Goats mating.jpg,
Goats The goat or domestic goat (''Capra hircus'') is a domesticated species of goat-antelope typically kept as livestock. It was domesticated from the wild goat (''C. aegagrus'') of Southwest Asia and Eastern Europe. The goat is a member of th ...
mating


Plants and fungi

Like in animals, mating in other Eukaryotes, such as
plant Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae exclu ...
s and
fungi A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from ...
, denotes . However, in vascular plants this is mostly achieved without physical contact between mating individuals (see
pollination Pollination is the transfer of pollen from an anther of a plant to the stigma of a plant, later enabling fertilisation and the production of seeds, most often by an animal or by wind. Pollinating agents can be animals such as insects, birds, a ...
), and in some cases, e.g., in fungi no distinguishable male or female organs exist (see
isogamy Isogamy is a form of sexual reproduction that involves gametes of the same morphology (indistinguishable in shape and size), found in most unicellular eukaryotes. Because both gametes look alike, they generally cannot be classified as male or ...
); however,
mating type Mating types are the microorganism equivalent to sexes in multicellular lifeforms and are thought to be the ancestor to distinct sexes. They also occur in macro-organisms such as fungi. Definition Mating types are the microorganism equivalent to ...
s in some fungal species are somewhat analogous to
sexual dimorphism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most an ...
in animals, and determine whether or not two individual isolates can mate. ''
Yeasts Yeasts are eukaryotic, single-celled microorganisms classified as members of the fungus kingdom. The first yeast originated hundreds of millions of years ago, and at least 1,500 species are currently recognized. They are estimated to constitut ...
'' are
eukaryotic Eukaryotes () are organisms whose Cell (biology), cells have a cell nucleus, nucleus. All animals, plants, fungi, and many unicellular organisms, are Eukaryotes. They belong to the group of organisms Eukaryota or Eukarya, which is one of the ...
microorganism A microorganism, or microbe,, ''mikros'', "small") and ''organism'' from the el, ὀργανισμός, ''organismós'', "organism"). It is usually written as a single word but is sometimes hyphenated (''micro-organism''), especially in olde ...
s classified in the
kingdom Kingdom commonly refers to: * A monarchy ruled by a king or queen * Kingdom (biology), a category in biological taxonomy Kingdom may also refer to: Arts and media Television * ''Kingdom'' (British TV series), a 2007 British television drama s ...
Fungi A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from ...
, with 1,500
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
currently described. In general, under high stress conditions like nutrient starvation, haploid cells will die; under the same conditions, however, diploid cells of ''
Saccharomyces cerevisiae ''Saccharomyces cerevisiae'' () (brewer's yeast or baker's yeast) is a species of yeast (single-celled fungus microorganisms). The species has been instrumental in winemaking, baking, and brewing since ancient times. It is believed to have b ...
'' can undergo sporulation, entering sexual reproduction (
meiosis Meiosis (; , since it is a reductional division) is a special type of cell division of germ cells in sexually-reproducing organisms that produces the gametes, such as sperm or egg cells. It involves two rounds of division that ultimately r ...
) and produce a variety of haploid spores, which can go on to mate (conjugate) and reform the diploid.


Protists

Protists are a large group of diverse
eukaryotic Eukaryotes () are organisms whose Cell (biology), cells have a cell nucleus, nucleus. All animals, plants, fungi, and many unicellular organisms, are Eukaryotes. They belong to the group of organisms Eukaryota or Eukarya, which is one of the ...
microorganism A microorganism, or microbe,, ''mikros'', "small") and ''organism'' from the el, ὀργανισμός, ''organismós'', "organism"). It is usually written as a single word but is sometimes hyphenated (''micro-organism''), especially in olde ...
s, mainly
unicellular A unicellular organism, also known as a single-celled organism, is an organism that consists of a single cell, unlike a multicellular organism that consists of multiple cells. Organisms fall into two general categories: prokaryotic organisms and ...
animals and plants, that do not form tissues. Eukaryotes emerged in evolution more than 1.5 billion years ago. The earliest eukaryotes were likely protists. Mating and sexual reproduction are widespread among extant eukaryotes including protists such as ''
Paramecium '' ''Paramecium'' ( , ; also spelled ''Paramoecium'') is a genus of eukaryotic, unicellular ciliates, commonly studied as a representative of the ciliate group. ''Paramecia'' are widespread in freshwater, brackish, and marine environments and a ...
'' and '' Chlamydomonas''. In many eukaryotic species, mating is promoted by
sex pheromones Sex pheromones are pheromones released by an organism to attract an individual of the same species, encourage them to mate with them, or perform some other function closely related with sexual reproduction. Sex pheromones specifically focus on ind ...
including the protist '' Blepharisma japonicum.'' Based on a phylogenetic analysis, Dacks and Roger proposed that facultative sex was present in the common ancestor of all eukaryotes. However, to many biologists it seemed unlikely until recently, that mating and sex could be a primordial and fundamental characteristic of eukaryotes. A principal reason for this view was that mating and sex appeared to be lacking in certain
pathogen In biology, a pathogen ( el, πάθος, "suffering", "passion" and , "producer of") in the oldest and broadest sense, is any organism or agent that can produce disease. A pathogen may also be referred to as an infectious agent, or simply a germ ...
ic protists whose ancestors branched off early from the eukaryotic family tree. However, several of these protists are now known to be capable of, or to recently have had, the capability for
meiosis Meiosis (; , since it is a reductional division) is a special type of cell division of germ cells in sexually-reproducing organisms that produces the gametes, such as sperm or egg cells. It involves two rounds of division that ultimately r ...
and hence mating. To cite one example, the common intestinal parasite ''
Giardia intestinalis ''Giardia duodenalis'', also known as ''Giardia intestinalis'' and ''Giardia lamblia'', is a flagellated parasitic microorganism of the genus ''Giardia'' that colonizes the small intestine, causing a diarrheal condition known as giardiasis. The ...
'' was once considered to be a descendant of a protist lineage that predated the emergence of meiosis and sex. However, ''G. intestinalis'' was recently found to have a core set of genes that function in meiosis and that are widely present among sexual eukaryotes. These results suggested that ''G. intestinalis'' is capable of meiosis and thus mating and sexual reproduction. Furthermore, direct evidence for meiotic recombination, indicative of mating and sexual reproduction, was also found in ''G. intestinalis''. Other protists for which evidence of mating and sexual reproduction has recently been described are parasitic protozoa of the genus '' Leishmania'', ''
Trichomonas vaginalis ''Trichomonas vaginalis'' is an anaerobic, flagellated protozoan parasite and the causative agent of a sexually transmitted disease called trichomoniasis. It is the most common pathogenic protozoan that infects humans in industrialized countri ...
'', and
acanthamoeba ''Acanthamoeba'' is a genus of amoebae that are commonly recovered from soil, fresh water, and other habitats. ''Acanthamoeba'' has two evolutive forms, the metabolically active trophozoite and a dormant, stress-resistant cyst. Trophozoites are ...
. Protists generally reproduce asexually under favorable environmental conditions, but tend to reproduce sexually under stressful conditions, such as starvation or heat shock.


See also

* Heterosexuality * Animal husbandry *
Breeding in the wild Breeding in the wild is the natural process of animal reproduction occurring in the natural habitat of a given species. This terminology is distinct from animal husbandry or breeding of species in captivity. Breeding locations are often chosen fo ...
* Breeding season *
Evolution of sex Sexual reproduction is an adaptive feature which is common to almost all multicellular organisms and various unicellular organisms, with some organisms being incapable of asexual reproduction. Currently the adaptive advantage of sexual repro ...
*
Lordosis behavior Lordosis behavior (), also known as mammalian lordosis (Greek lordōsis, from ''lordos'' "bent backward") or presenting, is the naturally occurring body posture for sexual receptivity to copulation present in females of most mammals including rod ...
* Mate choice copying *
Mating system A mating system is a way in which a group is structured in relation to sexual behaviour. The precise meaning depends upon the context. With respect to animals, the term describes which males and females mate under which circumstances. Recognised ...
* Reproduction * Sex determination system *
Sexual conflict Sexual conflict or sexual antagonism occurs when the two sexes have conflicting optimal fitness strategies concerning reproduction, particularly over the mode and frequency of mating, potentially leading to an evolutionary arms race between ma ...
* Sexual intercourse


References


External links


Introduction to Animal Reproduction

Advantages of Sexual Reproduction
{{Authority control Animal developmental biology Reproduction in animals Sexology Sexuality Ethology Fertility