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Upland and lowland are conditional descriptions of a
plain In geography, a plain is a flat expanse of land that generally does not change much in elevation, and is primarily treeless. Plains occur as lowlands along valleys or at the base of mountains, as coastal plains, and as plateaus or uplands ...
based on
elevation The elevation of a geographic location is its height above or below a fixed reference point, most commonly a reference geoid, a mathematical model of the Earth's sea level as an equipotential gravitational surface (see Geodetic datum § ...
above
sea level Mean sea level (MSL, often shortened to sea level) is an average surface level of one or more among Earth's coastal bodies of water from which heights such as elevation may be measured. The global MSL is a type of vertical datuma standardis ...
. In studies of the
ecology Ecology () is the study of the relationships between living organisms, including humans, and their physical environment. Ecology considers organisms at the individual, population, community, ecosystem, and biosphere level. Ecology overl ...
of
freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does incl ...
river A river is a natural flowing watercourse, usually freshwater, flowing towards an ocean, sea, lake or another river. In some cases, a river flows into the ground and becomes dry at the end of its course without reaching another body of ...
s,
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
s are classified as upland or lowland.


Definitions

Upland and lowland are portions of plain that are conditionally categorized by their elevation above the
sea level Mean sea level (MSL, often shortened to sea level) is an average surface level of one or more among Earth's coastal bodies of water from which heights such as elevation may be measured. The global MSL is a type of vertical datuma standardis ...
. Lowlands are usually no higher than , while uplands are somewhere around to . On unusual occasions, certain lowlands such as the Caspian Depression lie below sea level. Upland habitats are cold, clear and rocky whose rivers are fast-flowing in mountainous areas; lowland habitats are warm with slow-flowing rivers found in relatively flat lowland areas, with water that is frequently colored by
sediment Sediment is a naturally occurring material that is broken down by processes of weathering and erosion, and is subsequently transported by the action of wind, water, or ice or by the force of gravity acting on the particles. For example, sand ...
and organic matter. These classifications overlap with the geological definitions of "upland" and "lowland". In
geology Geology () is a branch of natural science concerned with Earth and other Astronomical object, astronomical objects, the features or rock (geology), rocks of which it is composed, and the processes by which they change over time. Modern geology ...
an "upland" is generally considered to be land that is at a higher elevation than the
alluvial plain An alluvial plain is a largely flat landform created by the deposition of sediment over a long period of time by one or more rivers coming from highland regions, from which alluvial soil forms. A floodplain is part of the process, being the smal ...
or
stream terrace Fluvial terraces are elongated terraces that flank the sides of floodplains and fluvial valleys all over the world. They consist of a relatively level strip of land, called a "tread", separated from either an adjacent floodplain, other fluvial te ...
, which are considered to be "lowlands". The term "bottomland" refers to low-lying alluvial land near a river. Much freshwater fish and invertebrate communities around the world show a pattern of specialization into upland or lowland river habitats. Classifying rivers and streams as upland or lowland is important in
freshwater ecology Freshwater ecosystems are a subset of Earth's aquatic ecosystems. They include lakes, ponds, rivers, streams, springs, bogs, and wetlands. They can be contrasted with marine ecosystems, which have a larger salt content. Freshwater habitats c ...
, as the two types of river habitat are very different, and usually support very different populations of fish and invertebrate species.


Upland

In freshwater ecology, upland rivers and
stream A stream is a continuous body of surface water flowing within the bed and banks of a channel. Depending on its location or certain characteristics, a stream may be referred to by a variety of local or regional names. Long large streams ...
s are the fast-flowing rivers and streams that drain elevated or mountainous country, often onto broad alluvial plains (where they become lowland rivers). However, elevation is not the sole determinant of whether a river is upland or lowland. Arguably the most important determinants are those of stream power and stream gradient. Rivers with a course that drops rapidly in elevation will have faster water flow and higher stream power or "force of water". This in turn produces the other characteristics of an upland river—an
incised course Incision may refer to: * Cutting, the separation of an object, into two or more portions, through the application of an acutely directed force * A type of open wound caused by a clean, sharp-edged object such as a knife, razor, or glass splinter * ...
, a
river bed A stream bed or streambed is the bottom of a stream or river (bathymetry) or the physical confine of the normal water flow ( channel). The lateral confines or channel margins are known as the stream banks or river banks, during all but flood ...
dominated by
bedrock In geology, bedrock is solid rock that lies under loose material ( regolith) within the crust of Earth or another terrestrial planet. Definition Bedrock is the solid rock that underlies looser surface material. An exposed portion of be ...
and coarse sediments, a riffle and pool structure and cooler water temperatures. Rivers with a course that drops in elevation very slowly will have slower water flow and lower force. This in turn produces the other characteristics of a lowland river—a meandering course lacking
rapids Rapids are sections of a river where the river bed has a relatively steep gradient, causing an increase in water velocity and turbulence. Rapids are hydrological features between a ''run'' (a smoothly flowing part of a stream) and a ''cascade' ...
, a river bed dominated by fine sediments and higher water temperatures. Lowland rivers tend to carry more suspended sediment and organic matter as well, but some lowland rivers have periods of high water clarity in seasonal low-flow periods. The generally clear, cool, fast-flowing waters and bedrock and coarse sediment beds of upland rivers encourage fish species with limited temperature tolerances, high oxygen needs, strong swimming ability and specialised reproductive strategies to prevent eggs or larvae being swept away. These characteristics also encourage invertebrate species with limited temperature tolerances, high oxygen needs and ecologies revolving around coarse sediments and interstices or "gaps" between those coarse sediments. The term "upland" is also used in wetland ecology, where "upland" plants indicate an area that is not a wetland.


Lowland

The generally more
turbid Turbidity is the cloudiness or haziness of a fluid caused by large numbers of individual particles that are generally invisible to the naked eye, similar to smoke in air. The measurement of turbidity is a key test of water quality. Fluids c ...
, warm, slow-flowing waters and fine sediment beds of lowland rivers encourage fish species with broad temperature tolerances and greater tolerances to low oxygen levels, and life history and breeding strategies adapted to these and other traits of lowland rivers. These characteristics also encourage invertebrate species with broad temperature tolerances and greater tolerances to low oxygen levels and ecologies revolving around fine sediments or alternative habitats such as submerged woody debris ("snags") or submergent macrophytes ("water weed").


Lowland alluvial plains

*
American Bottom The American Bottom is the flood plain of the Mississippi River in the Metro-East region of Southern Illinois, extending from Alton, Illinois, south to the Kaskaskia River. It is also sometimes called "American Bottoms". The area is about , mo ...
—flood plain of the Mississippi River in Southern Illinois *
Bois Brule Bottom The Bois Brule Bottom (French: Bois Brûlé) is an alluvial floodplain in Bois Brule Township in Perry County, Missouri stretching between Bois Brule Creek to the west and the Mississippi River to the east. The American Bottom stretches from St ...
*
Bottomland hardwood forest The bottomland hardwood forest is a type of deciduous and evergreen hardwood forest found in broad lowland floodplains along large rivers and lakes in the United States and elsewhere. They are occasionally flooded, which builds up the alluvial s ...
—deciduous hardwood forest found in broad lowland floodplains of the United States


See also

* Freshwater biology *
Highland Highlands or uplands are areas of high elevation such as a mountainous region, elevated mountainous plateau or high hills. Generally speaking, upland (or uplands) refers to ranges of hills, typically from up to while highland (or highlands) is ...
*
Mountain river A mountain river is a river that runs usually in mountains, in narrow, deep valley with steep banks, rocky stream bed, and accumulated rock debris. Mountain rivers are characterized by high slope and flow velocity, insignificant depth, frequent r ...
*
River reclamation Stream restoration or river restoration, also sometimes referred to as river reclamation, is work conducted to improve the environmental health of a river or stream, in support of biodiversity, recreation, flood management and/or landscape developm ...
*
Riparian zone A riparian zone or riparian area is the interface between land and a river or stream. Riparian is also the proper nomenclature for one of the terrestrial biomes of the Earth. Plant habitats and communities along the river margins and banks a ...


References

{{Authority control Freshwater ecology Water and the environment Riparian zone Rivers *