Ant Ants are eusocial insects of the family Formicidae and, along with the related wasps and bees, belong to the order Hymenoptera. Ants evolved from vespoid wasp ancestors in the Cretaceous period. More than 13,800 of an estimated total of 22 ...

s (

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Formicidae in the order

Hymenoptera Hymenoptera is a large order (biology), order of insects, comprising the sawfly, sawflies, wasps, bees, and ants. Over 150,000 living species of Hymenoptera have been described, in addition to over 2,000 extinct ones. Many of the species are Par ...

) are the most species-rich of all

social insect Eusociality (from Greek εὖ ''eu'' "good" and social), the highest level of organization of sociality, is defined by the following characteristics: cooperative brood care (including care of offspring from other individuals), overlapping generat ...

s, with more than 12,000 described

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

and many others awaiting description. Formicidae is divided into 21

subfamilies In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...

, of which 17 are

extant Extant is the opposite of the word extinct. It may refer to: * Extant hereditary titles * Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English * Extant taxon, a taxon which is not extinct, ...

and four subfamilies are

extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...

, described from

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

s. In total more than 300 genera have been described. Ants have come to occupy virtually all major terrestrial habitats, with the exception of tundra and cold ever-wet forests. They display a wide range of social behaviors, foraging habits and associations with other organisms, which has generated scientific and public interest. The following is a list of worldwide ant genera organised by subfamily.

Key

Subfamilies

Agroecomyrmecinae

The subfamily

Agroecomyrmecinae Agroecomyrmecinae is a subfamily of ants containing two extant and two fossil genera. The subfamily was originally classified in 1930 by Carpenter as Agroecomyrmecini, a Myrmicinae tribe. Bolton raised the tribe to subfamily status in 2003, sugg ...

represents two extant genera and two fossil genera, once widespread in both hemispheres during the early

Tertiary Tertiary ( ) is a widely used but obsolete term for the geologic period from 66 million to 2.6 million years ago. The period began with the demise of the non-avian dinosaurs in the Cretaceous–Paleogene extinction event, at the start ...

. The subfamily was originally classified as Agroecomyrmecini, a

Myrmicinae Myrmicinae is a subfamily of ants, with about 140 extant genera; their distribution is cosmopolitan. The pupae lack cocoons. Some species retain a functional sting. The petioles of Myrmicinae consist of two nodes. The nests are permanent and ...

tribe until English

myrmecologist Myrmecology (; from Greek: wikt:μύρμηξ, μύρμηξ, ''myrmex'', "ant" and λόγος, ''logos'', "study") is a branch of entomology focusing on the science, scientific study of ants. Some early myrmecologists considered ant society as th ...

Barry Bolton Barry Bolton is an English myrmecologist, an expert on the classification, systematics, and taxonomy of ants, who long worked at the Natural History Museum, London. He is known especially for monographs on African and Asian ants, and for encyclo ...

raised the tribe to subfamily status in 2003. Most specimens collected are from

Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...

and

Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...

, although one

type specimen In biology, a type is a particular wiktionary:en:specimen, specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to a ...

of an Agroecomyrmecinae species was collected from

Ghana Ghana (; tw, Gaana, ee, Gana), officially the Republic of Ghana, is a country in West Africa. It abuts the Gulf of Guinea and the Atlantic Ocean to the south, sharing borders with Ivory Coast in the west, Burkina Faso in the north, and To ...

Amblyoponinae

The subfamily

Amblyoponinae Amblyoponinae is a subfamily of ants in the poneromorph subfamilies group containing 13 extant genera and one extinct genus. The ants in this subfamily are mostly specialized subterranean predators. Adult workers pierce the integument (non letha ...

represents nine extant genera and one fossil genus. Established by Swiss myrmecologist

Auguste Forel Auguste-Henri Forel (1 September 1848 – 27 July 1931) was a Swiss myrmecologist, neuroanatomist, psychiatrist and eugenicist, notable for his investigations into the structure of the human brain and that of ants. For example, he is considered a ...

in 1893, these ants are specialist predators, distributed worldwide in the tropics.

Aneuretinae

The subfamily

Aneuretinae Aneuretinae is a subfamily of ants consisting of a single extant species, ''Aneuretus simoni'' ( Sri Lankan relict ant), and 9 fossil species. Earlier, the phylogenetic position of ''A. simoni'' was thought to be intermediate between primitive a ...

represents one extant genus and eight fossil genera established by Italian entomologist Carlo Emery in 1913. Only a single species of this subfamily is extant, the Sri Lankan relict ant (''Aneuretus simoni''), endemic to

Sri Lanka Sri Lanka (, ; si, ශ්‍රී ලංකා, Śrī Laṅkā, translit-std=ISO (); ta, இலங்கை, Ilaṅkai, translit-std=ISO ()), formerly known as Ceylon and officially the Democratic Socialist Republic of Sri Lanka, is an ...

Apomyrminae

The subfamily Apomyrminae contains the single genus '' Apomyrma'' which only has one species, the rare subterranean ant ''Apomyrma stygia'' from West Africa. Several undescribed species are known to exist, all from tropical Africa.

Brownimeciinae

The subfamily Brownimeciinae contains the single genus '' Brownimecia'' which only has one species, ''Brownimecia clavata''. It was described in 1997 after a fossilised specimen was collected from

Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...

amber from

New Jersey New Jersey is a state in the Mid-Atlantic and Northeastern regions of the United States. It is bordered on the north and east by the state of New York; on the east, southeast, and south by the Atlantic Ocean; on the west by the Delaware ...

, and was initially placed in the subfamily

Ponerinae Ponerinae is a subfamily of ants in the Poneromorph subfamilies group, with about 1,600 species in 47 extant genera, including ''Dinoponera gigantea'' - one of the world's largest species of ant. Mated workers have replaced the queen as the fun ...

. The species was later classified into its own subfamily in 2003 by

Dolichoderinae

The subfamily

Dolichoderinae Dolichoderinae is a subfamily of ants, which includes species such as the Argentine ant (''Linepithema humile''), the erratic ant, the odorous house ant, and the cone ant. The subfamily presents a great diversity of species throughout the wor ...

was established by Forel in 1878, which represents 28 extant genera and 20 fossil genera. The subfamily presents a great diversity of species throughout the world, mainly in the tropics. Most species are generalised scavengers, but some are predacious.

Dorylinae

The subfamily

Dorylinae Dorylinae is an ant subfamily, with distributions in both the Old World and New World. Brady ''et al.'' (2014) synonymized the previous dorylomorph subfamilies (Aenictinae, Aenictogitoninae, Cerapachyinae, Ecitoninae, and Leptanilloidinae) unde ...

was established by Leach in 1815, which represents 27 extant genera and one fossil genus. Many species of ant in this subfamily are known as

army ants The name army ant (or legionary ant or ''marabunta'') is applied to over 200 ant species in different lineages. Because of their aggressive predatory foraging groups, known as "raids", a huge number of ants forage simultaneously over a limit ...

that are distributed in the Old World and New World.

Ectatomminae

The subfamily Ectatomminae represents four extant genera and three fossil genera, established in 1895 by Carlo Emery. They are distributed in tropical and warm climates in the New World and Old World, as well as the

Indo-Australian all of this entry re immigration is a complete fabrication Indian Australians or Indo-Australians are Australians of Indian ancestry. This includes both those who are Australian by birth, and those born in India or elsewhere in the Indian d ...

regions.

Formiciinae

The subfamily

Formiciinae Formiciinae is an extinct subfamily of ants known from Eocene deposits in Europe and North America. Genera *Formiciinae Lutz, 1986 **Formiciini Lutz, 1986 ***''Titanomyrma'' Archibald, ''et al.'', 2011 ****''Titanomyrma gigantea'' (Lutz, 1986 ...

represents one extinct genus of ants dating back to the

Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...

. Ants of the genus ''

Titanomyrma ''Titanomyrma'' is a genus of prehistoric giant ant. The latest species to be discovered, ''T. lubei'', was described in 2011, when a 49.5-million-year-old fossilized winged queen ant, comparable in size to hummingbirds, was found in Wyoming, Uni ...

'' are the largest ants ever known, with queen specimens the size of small

hummingbirds Hummingbirds are birds native to the Americas and comprise the biological family Trochilidae. With about 361 species and 113 genera, they occur from Alaska to Tierra del Fuego, but the vast majority of the species are found in the tropics arou ...

. Fossils have been collected from the state of

Wyoming Wyoming () is a U.S. state, state in the Mountain states, Mountain West subregion of the Western United States. It is bordered by Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho to the west, Utah to the south ...

and in

Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...

Formicinae

The subfamily

Formicinae The Formicinae are a subfamily within the Formicidae containing ants of moderate evolutionary development. Formicines retain some primitive features, such as the presence of cocoons around pupae, the presence of ocelli in workers, and little ...

represents 51 extant genera and 30 fossil genera that are globally distributed. Established by French zoologist

Pierre André Latreille Pierre André Latreille (; 29 November 1762 – 6 February 1833) was a French zoologist, specialising in arthropods. Having trained as a Roman Catholic priest before the French Revolution, Latreille was imprisoned, and only regained his freedom ...

in 1809, the subfamily has more than 3,000 described species, placing it as the second largest ant subfamily. Despite this, the hyperdiverse genus '' Camponotus'' is the most diverse group of ants in the world, with more than 1,100 species described.

Haidomyrmecinae

The subfamily

Haidomyrmecinae Haidomyrmecinae, occasionally called Hell ants, are an extinct subfamily of ants (Formicidae) known from Cretaceous fossils found in ambers of North America, Europe, and Asia, spanning the late Albian to Campanian, around 100 to 79 million years ...

contains 9 fossil genera of specialized ants described from Cretaceous ambers. The Subfamily was previously treated as the tribe Haidomyrmecini and placed within

Sphecomyrminae Sphecomyrminae is an extinct subfamily of ants in family Formicidae known from a series of Cretaceous fossils found in North America, Europe, and Asia. Sphecomyrminae contains eight genera, divided into two tribes Sphecomyrmini and Zigrasimeciin ...

. The tribe was elevated to a subfamily in 2020.

Heteroponerinae

The subfamily

Heteroponerinae Heteroponerinae is a subfamily of ants in the poneromorph subfamilies group containing three genera in one tribe. The subfamily was created in 2003 when Barry Bolton divided the Ponerinae Ponerinae is a subfamily of ants in the Poneromo ...

represents three extant genera of ants, established in 2003 when

divided the subfamily

into six subfamilies. These ants are known from the

Neotropics The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropics, tropical Ecoregion#Terrestrial, terrestrial ecoregions of the Americas and the entire South American temperat ...

of Central America and South America while '' Aulacopone relicta'' is from

Azerbaijan Azerbaijan (, ; az, Azərbaycan ), officially the Republic of Azerbaijan, , also sometimes officially called the Azerbaijan Republic is a transcontinental country located at the boundary of Eastern Europe and Western Asia. It is a part of th ...

Leptanillinae

The subfamily

Leptanillinae Leptanillinae is a subfamily of ants. They are further divided into the tribes Anomalomyrmini and Leptanillini. In all Leptanillini, the larvae feed their hemolymph Hemolymph, or haemolymph, is a fluid, analogous to the blood in vertebrat ...

represents nine extant genera of ants, established in 1910 by Carlo Emery. They are subterranean ants from Africa, Europe and a single species known from Australia. Studies about their biology is minimal.

Martialinae

The subfamily

Martialinae ''Martialis heureka'' is a species of ant discovered in 2000 from the Amazon rainforest near Manaus, Brazil. It was described as a new species and placed as the sole member of a new subfamily, Martialinae. The generic name means "from Mars" ...

contains the single genus ''Martialis'' which only has one species, ''Martialis heureka''. The ant was discovered in 2000

Amazon rainforest The Amazon rainforest, Amazon jungle or ; es, Selva amazónica, , or usually ; french: Forêt amazonienne; nl, Amazoneregenwoud. In English, the names are sometimes capitalized further, as Amazon Rainforest, Amazon Forest, or Amazon Jungle. ...

near

Manaus, Brazil Manaus () is the capital and largest city of the Brazilian state of Amazonas. It is the seventh-largest city in Brazil, with an estimated 2020 population of 2,219,580 distributed over a land area of about . Located at the east center of the st ...

. Described in 2008, the ant belongs to the oldest known distinct lineage to have diverged from the ancestors of all other ants.

Myrmeciinae

The subfamily

Myrmeciinae Myrmeciinae is a subfamily of the Formicidae, ants once found worldwide but now restricted to Australia and New Caledonia. This subfamily is one of several ant subfamilies which possess gamergates, female worker ants which are able to mate and ...

represents two extant genera and five fossil genera that were once found worldwide. Established by Carlo Emery in 1877, the extant genera are restricted to Australia,

New Caledonia ) , anthem = "" , image_map = New Caledonia on the globe (small islands magnified) (Polynesia centered).svg , map_alt = Location of New Caledonia , map_caption = Location of New Caledonia , mapsize = 290px , subdivision_type = Sovereign st ...

and

New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...

. The notorious ant genus ''

Myrmecia Myrmecia can refer to: * ''Myrmecia'' (alga), genus of algae associated with lichens * ''Myrmecia'' (ant), genus of ants called bulldog ants * Myrmecia (skin), a kind of deep wart on the human hands or feet See also * '' Copromorpha myrmecias'' ...

'' is known for their venomous stings and aggression, which has caused several human deaths in sensitive people.

Myrmicinae

The subfamily

was established by Lepeletier de Saint-Fargeau in 1835. It represents 142 extant genera and 36 fossil genera that are distributed globally. It is the largest subfamily of the Formicidae, with more than 6,758 species described. The seed-harvesting ants and

fungus-growing ants Fungus-growing ants (tribe Attini) comprise all the known fungus-growing ant species participating in ant–fungus mutualism. They are known for cutting grasses and leaves, carrying them to their colonies' nests, and using them to grow fungus o ...

are well known among the Myrmicines.

Paraponerinae

The subfamily

Paraponerinae ''Paraponera'' is a genus of ants and the only genus in the subfamily Paraponerinae. The name means "near-''Ponera''". It consists of two species: the extant ''Paraponera clavata'', also known as a bullet ant, found in the Neotropics, and the v ...

contains a single genus '' Paraponera''. This genus has two species, one of which was found in Dominican amber from the

Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...

. The extant species, ''

Paraponera clavata ''Paraponera clavata'' is a species of ant, commonly known as the bullet ant, named for its extremely painful sting. It inhabits humid lowland rainforests in Central and South America. Etymology The specific epithet of the ant, ', means "club- ...

'', is found in Central America and South America, and the pain from its sting is said to be greater than any other insect sting on earth.

Ponerinae

The subfamily

was established by Lepeletier de Saint-Fargeau in 1835, which represents 47 extant genera and 12 fossil genera. The subfamily is among the most diverse in the family Formicidae, with more than 1,000 species described. They are mostly distributed in the tropics and subtropics.

Proceratiinae

The subfamily

Proceratiinae Proceratiinae is a subfamily of ants in the poneromorph subfamilies group, with three extant genera, of which most are tropical or subtropical, although overall distribution is worldwide. Identification The ants are relatively small to medium i ...

was established by Italian entomologist

Carlo Emery Carlo Emery (25 October 1848, Naples – 11 May 1925) was an Italian entomologist. He is remembered for Emery's rule, which states that insect social parasitism (biology), social parasites are often closely related to their hosts. Early in hi ...

in 1895, which represents three extant genera and one extinct genus. Found worldwide, these ants are mainly encountered in tropical and subtropical areas. Little is known about their biology.

Pseudomyrmecinae

The subfamily

Pseudomyrmecinae Pseudomyrmecinae is a small subfamily of ants containing only three genera of slender, large-eyed arboreal ants, predominantly tropical or subtropical in distribution. In the course of adapting to arboreal conditions (unlike the predominantly gro ...

was established by M.R. Smith in 1952, which represents three genera of ants that are primarily

arboreal Arboreal locomotion is the Animal locomotion, locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally, but others are exclusively arboreal. Th ...

nesting ants in the

tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...

and

subtropical The subtropical zones or subtropics are geographical zone, geographical and Köppen climate classification, climate zones to the Northern Hemisphere, north and Southern Hemisphere, south of the tropics. Geographically part of the Geographical z ...

regions. They are found in

Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...

Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area ...

, Australia, North America and South America.

Sphecomyrminae

The subfamily

contains 9 fossil genera of stem ants. Most fossilized ants from Cretaceous amber were placed in this subfamily, however revisions in 2017 and 2020 removed several genera and added former members of the subfamily

Armaniinae Armaniidae was a name formerly given to a group of extinct ant-like hymenopterans known from a series of Cretaceous fossils found in Asia and Africa. Armaniidae has been suggested by several authors to belong to the family Formicidae as one of t ...

''Incertae sedis''

There are several ant genera where their

taxonomic Taxonomy is the practice and science of categorization or classification. A taxonomy (or taxonomical classification) is a scheme of classification, especially a hierarchical classification, in which things are organized into groups or types. ...

placement is uncertain (''

incertae sedis ' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertainty ...

''). These genera have not yet been assigned to any

subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...

within Formicidae; 16 genera are currently listed as ''incertae sedis''.

Formerly included in Formicidae

There are several genera which were formerly placed in Formicidae, but have subsequently been removed. These genera are now placed in other families, are considered ''incertae sedis'' within Hymenoptera taxonomy, or are considered invalid.

Notes

References

{{Formicidae subfamilies

Genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclat ...