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passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped'), which includes more than half of all bird species. Sometimes known as perching birds, passerines are distinguished from other orders of birds by th ...
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
s of the family Alaudidae. Larks have a
cosmopolitan distribution In biogeography, cosmopolitan distribution is the term for the range of a taxon that extends across all or most of the world in appropriate habitats. Such a taxon, usually a species, is said to exhibit cosmopolitanism or cosmopolitism. The extr ...
with the largest number of species occurring in Africa. Only a single species, the horned lark, occurs in North America, and only Horsfield's bush lark occurs in Australia. Habitats vary widely, but many species live in dry regions. When the word "lark" is used without specification, it often refers to the Eurasian skylark ''(Alauda arvensis)''.


Taxonomy and systematics

The family Alaudidae was introduced in 1825 by the Irish zoologist
Nicholas Aylward Vigors Nicholas Aylward Vigors (1785 – 26 October 1840) was an Irish zoologist and politician. He popularized the classification of birds on the basis of the quinarian system. Early life Vigors was born at Old Leighlin, County Carlow on 1785 a ...
as a subfamily Alaudina of the finch family Fringillidae. Larks are a well-defined family, partly because of the shape of their . They have multiple scutes on the hind side of their tarsi, rather than the single plate found in most songbirds. They also lack a pessulus, the bony central structure in the syrinx of songbirds. They were long placed at or near the beginning of the songbirds or oscines (now often called
Passeri A songbird is a bird belonging to the suborder Passeri of the perching birds (Passeriformes). Another name that is sometimes seen as the scientific or vernacular name is Oscines, from Latin ''oscen'', "songbird". The Passeriformes contains 5000 ...
), just after the
suboscines The Tyranni (suboscines) are a suborder of passerine birds that includes more than 1,000 species, the large majority of which are South American. It is named after the type genus ''Tyrannus''. These have a different anatomy of the syrinx musc ...
and before the swallows, for example in the
American Ornithologists' Union The American Ornithological Society (AOS) is an ornithological organization based in the United States. The society was formed in October 2016 by the merger of the American Ornithologists' Union (AOU) and the Cooper Ornithological Society. Its ...
's first check-list. Some authorities, such as the
British Ornithologists' Union The British Ornithologists' Union (BOU) aims to encourage the study of birds ("ornithology") and around the world, in order to understand their biology and to aid their conservation. The BOU was founded in 1858 by Professor Alfred Newton, Henr ...
and the ''
Handbook of the Birds of the World The ''Handbook of the Birds of the World'' (HBW) is a multi-volume series produced by the Spanish publishing house Lynx Edicions in partnership with BirdLife International. It is the first handbook to cover every known living species of bird. ...
'', adhere to that placement. However, many other classifications follow the Sibley-Ahlquist taxonomy in placing the larks in a large oscine subgroup Passerida (which excludes crows,
shrike Shrikes () are passerine birds of the family Laniidae. The family is composed of 34 species in four genera. The family name, and that of the largest genus, '' Lanius'', is derived from the Latin word for "butcher", and some shrikes are also know ...
s and their allies, vireos, and many groups characteristic of Australia and southeastern Asia). For instance, the American Ornithologists' Union places larks just after the crows, shrikes, and vireos. At a finer level of detail, some now place the larks at the beginning of a
superfamily SUPERFAMILY is a database and search platform of structural and functional annotation for all proteins and genomes. It classifies amino acid sequences into known structural domains, especially into SCOP superfamilies. Domains are functional, str ...
Sylvioidea with the swallows, various "
Old World warbler Old World warblers are a large group of birds formerly grouped together in the bird family Sylviidae. The family held over 400 species in over 70 genera, and were the source of much taxonomic confusion. Two families were split out initially, the ci ...
" and "
babbler Babbler may refer to: * Old World babbler, a large family of mostly Old World passerine birds * Australo-Papuan babbler, passerine birds endemic to Australia-New Guinea * Babbler (software), a French software company * Babbler (''Dungeons & Dr ...
" groups, and others.
Molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...
studies have shown that within the Sylvioidea the larks form a sister
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English ter ...
to the family Panuridae which contains a single species, the
bearded reedling The bearded reedling (''Panurus biarmicus'') is a small, sexually dimorphic reed-bed passerine bird. It is frequently known as the bearded tit, due to some similarities to the long-tailed tit, or the bearded parrotbill. It is the only species ...
(''Panurus biarmicus''). The phylogeny of larks (Alaudidae) was reviewed in 2013, leading to the recognition of the arrangement below.


Extant genera

The family Alaudidae contains 100 extant species which are divided into 21
genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclat ...
: For more detail, see list of lark species.


Extinct genera

* Genus '' Eremarida'' — (''Eremarida xerophila'')


Description

Larks, or the family Alaudidae, are small- to medium-sized birds, in length and in mass. The smallest larks are likely the '' Spizocorys'' species, which can weigh only around in species like the pink-billed lark and the Obbia lark, while the largest lark is the
Tibetan lark The Tibetan lark (''Melanocorypha maxima'') is a species of lark in the family Alaudidae found on the Tibetan plateau The Tibetan Plateau (, also known as the Qinghai–Tibet Plateau or the Qing–Zang Plateau () or as the Himalayan Plateau ...
. Like many ground birds, most lark species have long hind claws, which are thought to provide stability while standing. Most have streaked brown plumage, some boldly marked with black or white. Their dull appearance
camouflage Camouflage is the use of any combination of materials, coloration, or illumination for concealment, either by making animals or objects hard to see, or by disguising them as something else. Examples include the leopard's spotted coat, the b ...
s them on the ground, especially when on the nest. They feed on
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pa ...
s and
seeds A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiosperm ...
; though adults of most species eat seeds primarily, all species feed their young insects for at least the first week after hatching. Many species dig with their bills to uncover food. Some larks have heavy bills (reaching an extreme in the thick-billed lark) for cracking seeds open, while others have long, down-curved bills, which are especially suitable for digging. Larks are the only passerines that lose all their feathers in their first
moult In biology, moulting (British English), or molting (American English), also known as sloughing, shedding, or in many invertebrates, ecdysis, is the manner in which an animal routinely casts off a part of its body (often, but not always, an outer ...
(in all species whose first moult is known). This may result from the poor quality of the chicks' feathers, which in turn may result from the benefits to the parents of switching the young to a lower-quality diet (seeds), which requires less work from the parents. In many respects, including long tertial feathers, larks resemble other ground birds such as pipits. However, in larks the tarsus (the lowest leg bone, connected to the toes) has only one set of scales on the rear surface, which is rounded. Pipits and all other
songbird A songbird is a bird belonging to the suborder Passeri of the perching birds (Passeriformes). Another name that is sometimes seen as the scientific or vernacular name is Oscines, from Latin ''oscen'', "songbird". The Passeriformes contains 5000 ...
s have two plates of scales on the rear surface, which meet at a protruding rear edge.


Calls and song

Larks have more elaborate calls than most birds, and often extravagant songs given in display flight. These melodious sounds (to human ears), combined with a willingness to expand into
anthropogenic Anthropogenic ("human" + "generating") is an adjective that may refer to: * Anthropogeny, the study of the origins of humanity Counterintuitively, anthropogenic may also refer to things that have been generated by humans, as follows: * Human i ...
habitats — as long as these are not too intensively managed — have ensured larks a prominent place in literature and music, especially the Eurasian skylark in northern Europe and the crested lark and
calandra lark The calandra lark (''Melanocorypha calandra'') or European calandra-lark breeds in warm temperate countries around the Mediterranean and eastwards through Turkey into northern Iran and southern Russia. It is replaced further east by its relative ...
in southern Europe.


Behaviour


Breeding

Male larks use song flights to defend their breeding territory and attract a mate. Most species build nests on the ground, usually
cups CUPS (formerly an acronym for Common UNIX Printing System) is a modular printing system for Unix-like computer operating systems which allows a computer to act as a print server. A computer running CUPS is a host that can accept print job ...
of dead grass, but in some species the nests are more complicated and partly domed. A few desert species nest very low in bushes, perhaps so circulating air can cool the nest. Larks' eggs are usually speckled. The size of the clutch is very variable and ranges from the single egg laid by Sclater's lark up to 6-8 eggs laid by the
calandra lark The calandra lark (''Melanocorypha calandra'') or European calandra-lark breeds in warm temperate countries around the Mediterranean and eastwards through Turkey into northern Iran and southern Russia. It is replaced further east by its relative ...
and the black lark. Larks incubate for 11 to 16 days.


In culture


Larks as food

Larks, commonly consumed with bones intact, have historically been considered wholesome, delicate, and light game. They can be used in a number of dishes; for example, they can be stewed, broiled, or used as filling in a meat pie. Lark's tongues are reputed to have been particularly highly valued as a delicacy. In modern times, shrinking habitats made lark meat rare and hard to come by, though it can still be found in restaurants in Italy and elsewhere in southern Europe.


Symbolism

The lark in mythology and literature stands for daybreak, as in Chaucer's "
The Knight's Tale "The Knight's Tale" ( enm, The Knightes Tale) is the first tale from Geoffrey Chaucer's '' The Canterbury Tales''. The Knight is described by Chaucer in the " General Prologue" as the person of highest social standing amongst the pilgrims, t ...
", "the bisy larke, messager of day", and Shakespeare's Sonnet 29, "the lark at break of day arising / From sullen earth, sings hymns at heaven's gate" (11–12). The lark is also (often simultaneously) associated with "lovers and lovers' observance" (as in
Bernart de Ventadorn Bernart de Ventadorn (also Bernard de Ventadour or Bernat del Ventadorn; – ) was a French poet-composer troubadour of the classical age of troubadour poetry. Generally regarded as the most important troubadour in both poetry and music, his ...
's
Can vei la lauzeta mover Can vei la lauzeta mover (PC 70.43) is a song written in the Occitan language by Bernart de Ventadorn, a 12th-century troubadour. It is among both the oldest and best known of the troubadour songs. Both the lyrics and the melody of the song survive ...
) and with "church services", and often those the meanings of daybreak and religious reference are combined (in Blake's ''
Visions of the Daughters of Albion ''Visions of the Daughters of Albion'' is a 1793 poem by William Blake, produced as a book with his own illustrations. It is a short and early example of his prophetic books, and a sequel of sorts to ''The Book of Thel''. Plot The central narra ...
'', into a "spiritual daybreak" to signify "passage from Earth to Heaven and from Heaven to Earth". In Renaissance painters such as
Domenico Ghirlandaio Domenico di Tommaso Curradi di Doffo Bigordi (, , ; 2 June 1448 – 11 January 1494), professionally known as Domenico Ghirlandaio, also spelled as Ghirlandajo, was an Italian Renaissance painter born in Florence. Ghirlandaio was part of t ...
the lark symbolizes
Christ Jesus, likely from he, יֵשׁוּעַ, translit=Yēšūaʿ, label= Hebrew/ Aramaic ( AD 30 or 33), also referred to as Jesus Christ or Jesus of Nazareth (among other names and titles), was a first-century Jewish preacher and relig ...
, in reference to
John John is a common English name and surname: * John (given name) * John (surname) John may also refer to: New Testament Works * Gospel of John, a title often shortened to John * First Epistle of John, often shortened to 1 John * Secon ...
16:16.


Literature

Percy Bysshe Shelley's famed 1820 poem
To a Skylark "To a Skylark" is a poem completed by Percy Bysshe Shelley in late June 1820 and published accompanying his lyrical drama '' Prometheus Unbound'' by Charles and James Collier in London. It was inspired by an evening walk in the country near ...
was inspired by the melodious song of a skylark during an evening walk. English poet George Meredith wrote a poem '' The Lark Ascending'' in 1881.


Music

English composer
Ralph Vaughan Williams Ralph Vaughan Williams, (; 12 October 1872– 26 August 1958) was an English composer. His works include operas, ballets, chamber music, secular and religious vocal pieces and orchestral compositions including nine symphonies, written over ...
wrote a musical setting of George Meredith's poem, completed in 1914. It was composed for violin and piano, and entitled '' The Lark Ascending - A Romance''. The work received its first performance in December 1920. Soon afterwards the composer arranged it for violin and orchestra, in which version it was first performed in June 1921, and this is how the work remains best-known today.


Pet

Traditionally larks are kept as pets in China. In Beijing, larks are taught to mimic the voice of other songbirds and animals. It is an old-fashioned habit of the Beijingers to teach their larks 13 kinds of sounds in a strict order (called "the 13 songs of a lark", Chinese: 百灵十三套). The larks that can sing the full 13 sounds in the correct order are highly valued, while any disruption in the songs will decrease its value significantly.


"On a lark" Phrase

In english, the term "on a lark" describes something that is done on the spur of the moment, something that is done spontaneously and for fun. The word lark in the term on a lark is used to mean frolicking or playing. The use of the word lark to mean frolicking first appeared in the early 1800s. It may stem from the word skylark, slang used by sailors to mean roughhousing in the rigging. It may date back to the Old Norse word leika which means to play. A third possibility is an Old English dialect word lake, which means to leap and play.


Early awakening

Larks sing early in the day, often before dawn, leading to the expression "up with the lark" for a person who is awake early in the day, and the term '' lark'' being applied to someone who habitually rises early in the morning.


See also

* Lark bunting * Lark sparrow * Magpie-lark (Neither a lark nor a magpie, but a giant monarch flycatcher) *
Meadowlark Meadowlarks are New World grassland birds belonging to genera '' Sturnella'' and '' Leistes''. This group includes seven species of largely insectivorous grassland birds. In all species the male at least has a black or brown back and extensivel ...
* Titlark, a synonym for meadow pipit * Songlark * "Alouette" (a song)


References


Further reading

*


External links


Lark videos, photos and sounds
- Internet Bird Collection {{Taxonbar, from=Q29858 * Taxa named by Nicholas Aylward Vigors