Krummholz
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''Krummholz'' (german: krumm, "crooked, bent, twisted" and ''Holz'', "wood") — also called ''knieholz'' ("knee timber") — is a type of stunted, deformed vegetation encountered in the
subarctic The subarctic zone is a region in the Northern Hemisphere immediately south of the true Arctic, north of humid continental regions and covering much of Alaska, Canada, Iceland, the north of Scandinavia, Siberia, and the Cairngorms. Genera ...
and subalpine
tree line The tree line is the edge of the habitat at which trees are capable of growing. It is found at high elevations and high latitudes. Beyond the tree line, trees cannot tolerate the environmental conditions (usually cold temperatures, extreme snow ...
landscapes, shaped by continual exposure to fierce, freezing winds. Under these conditions, trees can only survive where they are sheltered by rock formations or
snow Snow comprises individual ice crystals that grow while suspended in the atmosphere—usually within clouds—and then fall, accumulating on the ground where they undergo further changes. It consists of frozen crystalline water throughou ...
cover. As the lower portion of these trees continues to grow, the coverage becomes extremely dense near the ground. In
Newfoundland and Labrador Newfoundland and Labrador (; french: Terre-Neuve-et-Labrador; frequently abbreviated as NL) is the easternmost province of Canada, in the country's Atlantic region. The province comprises the island of Newfoundland and the continental region ...
, the formation is known as tuckamore. ''Krummholz'' trees are also found on beaches such as the Oregon coast, where trees can become much taller than their subalpine cousins.


Species

Common trees showing ''krummholz'' formation include
European spruce ''Picea abies'', the Norway spruce or European spruce, is a species of spruce native to Northern, Central and Eastern Europe. It has branchlets that typically hang downwards, and the largest cones of any spruce, 9–17 cm long. It is very close ...
,
mountain pine ''Pinus mugo'', known as bog pine, creeping pine, dwarf mountain pine, mugo pine, mountain pine, scrub mountain pine, or Swiss mountain pine, is a species of conifer, native to high elevation habitats from southwestern to Central Europe and So ...
, balsam fir, red spruce,
black spruce ''Picea mariana'', the black spruce, is a North American species of spruce tree in the pine family. It is widespread across Canada, found in all 10 provinces and all 3 territories. It is the official tree of the province of Newfoundland and Lab ...
, subalpine fir,
subalpine larch ''Larix lyallii'', the subalpine larch, or simply alpine larch, is a deciduous, coniferous tree native to northwestern North America. It lives at high altitudes, from , in the Rocky Mountains of Idaho, Montana, British Columbia, and Alberta. Ther ...
, Engelmann spruce,
whitebark pine ''Pinus albicaulis'', known by the common names whitebark pine, white bark pine, white pine, pitch pine, scrub pine, and creeping pine, is a conifer tree native to the mountains of the western United States and Canada, specifically subalpine ...
, limber pine, Bristlecone pine, and lodgepole pine. Instances of the ''krummholz'' form of black spruce, '' Picea mariana'', are found in the northern Canadian
boreal forest Taiga (; rus, тайга́, p=tɐjˈɡa; relates to Mongolic and Turkic languages), generally referred to in North America as a boreal forest or snow forest, is a biome characterized by coniferous forests consisting mostly of pines, spruc ...
s.C. Michael Hogan. 2008
''Black Spruce: Picea mariana'', GlobalTwitcher.com, ed. N. Stromberg
''Krummholz''-form black spruce and balsam fir are abundant in the alpine transition zone of the White Mountains of Maine and New Hampshire and of the same zone in the
Green Mountains The Green Mountains are a mountain range in the U.S. state of Vermont. The range runs primarily south to north and extends approximately from the border with Massachusetts to the border with Quebec, Canada. The part of the same range that is i ...
of
Vermont Vermont () is a U.S. state, state in the northeast New England region of the United States. Vermont is bordered by the states of Massachusetts to the south, New Hampshire to the east, and New York (state), New York to the west, and the Provin ...
. Subalpine fir is the most common associate of spruce in ''krummholz'' vegetation. Other associated
coniferous Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All ext ...
species include alpine larch ('' Larix lyallii''), whitebark pine ('' Pinus albicaulis''), limber pine ('' Pinus flexilis''), and western white pine (''
Pinus monticola Western white pine (''Pinus monticola''), also called silver pine and California mountain pine, is a species of pine in the family Pinaceae. It occurs in mountain ranges of northwestern North America. It is the state tree of Idaho. Description ...
'') in southern British Columbia, and mountain hemlock ('' Tsuga mertensiana'') on the eastern slopes of the Coast Range and in the Revelstoke area of
British Columbia British Columbia (commonly abbreviated as BC) is the westernmost province of Canada, situated between the Pacific Ocean and the Rocky Mountains. It has a diverse geography, with rugged landscapes that include rocky coastlines, sandy beaches, for ...
. Lodgepole pine ( ''Pinus contorta'' var. ''contorta'') is a minor associate in most of the British Columbia interior, except in dry alpine areas of the southwest Cariboo/Chilcotin district where it is abundant (Pojar 1985).Pojar, J. 1985. Ecological classification of lodgepole pine in Canada. pp. 77–88 ''in'' D.M. Baumgartner, R.G. Krebill, J.T. Arnott and G.F. Weeman (compilers). Lodgepole pine the species and its management. Symp. Proc.. Cooperative Extension, Washington State University, Pullman, WA. Ericaceous species (''Vaccinium scoparium, V. membranaceum, V. caespitosum, Cassiope mertensiana, Phyllodoce empetriformis'') are common in the snow accumulation zone around the base of ''krummholz'' colonies.


Flag tree

A variation of ''krummholz'' formation is a flag tree or banner tree. Branches on the windward side are killed or deformed by the almost constant strong winds, giving the tree a characteristic flag-like appearance. Where the lower portion of the tree is protected by snow cover or rocks, only the exposed upper portion may have this appearance. This is a rather common occurrence in red spruce trees of the highest peaks of the central and even the southern
Appalachian Mountains The Appalachian Mountains, often called the Appalachians, (french: Appalaches), are a system of mountains in eastern to northeastern North America. The Appalachians first formed roughly 480 million years ago during the Ordovician Period. The ...
, and is most often seen in the windswept high peaks and plateaus of the
Allegheny Mountains The Allegheny Mountain Range (; also spelled Alleghany or Allegany), informally the Alleghenies, is part of the vast Appalachian Mountain Range of the Eastern United States and Canada and posed a significant barrier to land travel in less devel ...
. This formation most notably occurs with high frequency in the Dolly Sods and Roaring Plains West Wilderness areas along the
Allegheny Front The Allegheny Front is the major southeast- or east-facing escarpment in the Allegheny Mountains in southern Pennsylvania, western Maryland, eastern West Virginia, and western Virginia, USA. The Allegheny Front forms the boundary between the ...
in eastern
West Virginia West Virginia is a state in the Appalachian, Mid-Atlantic and Southeastern regions of the United States.The Census Bureau and the Association of American Geographers classify West Virginia as part of the Southern United States while the ...
, typically occurring at elevations of and higher. Trade winds in tropical regions near the equator can also shape trees in a similar manner. The southernmost reaches of the Magellanic subpolar forests of Chile contain many flag trees also.


See also

* Dwarf forest * ''
Libidibia coriaria ''Libidibia coriaria'', synonym ''Caesalpinia coriaria'', is a leguminous tree or large shrub native to the Caribbean, Central America, Mexico, and northern and western South America. Common names include divi-divi, cascalote, guaracabuya, guat ...
'' (divi-divi) * '' Pinus mugo'' (scrub mountain pine) * Reaction wood *
Windthrow In forestry, windthrow refers to trees uprooted by wind. Breakage of the tree bole (trunk) instead of uprooting is called windsnap. Blowdown refers to both windthrow and windsnap. Causes Windthrow is common in all forested parts of the w ...


References


Further reading


Recent history of subalpine Krummholz

Maintaining Krummholz above treeline


External links

*{{commons-inline

Trees Conifers Forest ecology Montane ecology