''Juniperus macrocarpa'' (large-fruited juniper, syn. ''J. oxycedrus'' subsp. ''macrocarpa'' (Sibth. & Sm.) Ball) is a species of

juniper Junipers are coniferous trees and shrubs in the genus ''Juniperus'' () of the cypress family Cupressaceae. Depending on the taxonomy, between 50 and 67 species of junipers are widely distributed throughout the Northern Hemisphere, from the Arc ...

, native across the northern

Mediterranean The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on ...

region from southwestern

Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = '' Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , ...

east to western

Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a transcontinental country located mainly on the Anatolian Peninsula in Western Asia, with a small portion on the Balkan Peninsula ...

and

Cyprus Cyprus ; tr, Kıbrıs (), officially the Republic of Cyprus,, , lit: Republic of Cyprus is an island country located south of the Anatolian Peninsula in the eastern Mediterranean Sea. Its continental position is disputed; while it is ...

, growing on

coast The coast, also known as the coastline or seashore, is defined as the area where land meets the ocean, or as a line that forms the boundary between the land and the coastline. The Earth has around of coastline. Coasts are important zones in n ...

sand dune A dune is a landform composed of wind- or water-driven sand. It typically takes the form of a mound, ridge, or hill. An area with dunes is called a dune system or a dune complex. A large dune complex is called a dune field, while broad, f ...

s from sea level up to in altitude.Adams, R. P. (2004). ''Junipers of the World''. Trafford. Farjon, A. (2005). ''Monograph of Cupressaceae and Sciadopitys''. Royal Botanic Gardens, Kew. It is a spreading

shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from tree ...

tall, rarely a small

tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...

up to tall. The

leaves A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, st ...

are broad lanceolate, produced in whorls of three, green, long and 2–3 mm broad, with a double white

stoma In botany, a stoma (from Greek ''στόμα'', "mouth", plural "stomata"), also called a stomate (plural "stomates"), is a pore found in the epidermis of leaves, stems, and other organs, that controls the rate of gas exchange. The pore is bo ...

tal band split by a green midrib on the inner surface. It is

dioecious Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproducti ...

, with separate male and female plants. The seed cones are

berry-like In botany, a berry is a fleshy fruit without a stone (pit) produced from a single flower containing one ovary. Berries so defined include grapes, currants, and tomatoes, as well as cucumbers, eggplants (aubergines) and bananas, but exclude ...

, green ripening in 18 months to orange-red with a variable pink waxy coating; they are spherical, 12–18 mm diameter, and have six fused scales in two whorls, three of the scales with a single

seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiosper ...

. The seeds are dispersed when

bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...

s eat the cones, digesting the fleshy scales and passing the hard seeds in their droppings. The

pollen Pollen is a powdery substance produced by seed plants. It consists of pollen grains (highly reduced microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametop ...

cones are yellow, 2–3 mm long, and fall soon after shedding their pollen in late winter.Arboretum de Villardebelle
photos of cones and shoots
/ref> Despite its distinct morphology with large cones and broad leaves more like those of '' Juniperus drupacea'', it has often been treated as a

subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. Not all specie ...

of '' Juniperus oxycedrus'', though recent genetic studiesAdams, R. P. (2000). Systematics of ''Juniperus'' section ''Juniperus'' based on leaf essential oils and RAPD DNA fingerprinting. ''Biochem. Syst. Ecol''. 28: 515-528
available online (pdf file)
/ref>Adams, R. P., Morris, J. A., Pandey, R. N., & Schwarzbach, A. E. (2005). Cryptic speciation between Juniperus deltoides and J. oxycedrus (Cupressaceae) in the Mediterranean. ''Biochem. Syst. Ecol''. 33: 771-787
available online (pdf file)
/ref> have shown its DNA is distinct from that of ''J. oxycedrus''.

References

{{Taxonbar, from=Q138307 macrocarpa Trees of Mediterranean climate Trees of Europe Trees of Turkey Flora of Cyprus Dioecious plants Flora of the Mediterranean Basin