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The jaguar ( Panthera
Panthera
onca), is a wild cat species and the only extant member of the genus Panthera
Panthera
native to the Americas. The jaguar's present range extends from Southwestern United States
Southwestern United States
and Mexico across much of Central America
Central America
and south to Paraguay
Paraguay
and northern Argentina. Though there are single cats now living within the western United States, the species has largely been extirpated from the United States since the early 20th century. It is listed as Near Threatened on the IUCN Red List; and its numbers are declining. Threats include loss and fragmentation of habitat. The jaguar is the largest cat species in the Americas
Americas
and the third-largest after the tiger and the lion. This spotted cat closely resembles the leopard, but is usually larger and sturdier. It ranges across a variety of forested and open terrains, but its preferred habitat is tropical and subtropical moist broadleaf forest, swamps and wooded regions. The jaguar enjoys swimming and is largely a solitary, opportunistic, stalk-and-ambush predator at the top of the food chain. As a keystone species it plays an important role in stabilizing ecosystems and regulating prey populations. While international trade in jaguar or its body parts is prohibited, the cat is still frequently killed, particularly in conflicts with ranchers and farmers in South America. Although reduced, its range remains large. Given its historical distribution, the jaguar has featured prominently in the mythology of numerous indigenous American cultures, including those of the Maya and Aztec.

Contents

1 Etymology 2 Evolution 3 Taxonomy 4 Characteristics

4.1 Color variation

5 Distribution and habitat 6 Ecology and behavior

6.1 Ecological role 6.2 Reproduction and life cycle 6.3 Social activity 6.4 Hunting and diet

6.4.1 Man-eating

7 Threats 8 Conservation

8.1 Jaguar
Jaguar
Conservation Units 8.2 Approaches 8.3 The United States

9 In mythology and culture

9.1 Pre-Columbian Americas 9.2 Contemporary culture

10 References

10.1 Bibliography

11 Further reading 12 External links

Etymology The word 'jaguar' is thought to derive from the Tupian word yaguara, meaning "beast of prey".[2] The word entered English presumably via the Amazonian trade language Tupinambá, via Portuguese jaguar.[3][4] The specific word for jaguar is yaguareté, with the suffix -eté meaning "real" or "true".[5] The word 'panther' derives from classical Latin panthēra, itself from the ancient Greek pánthēr (πάνθηρ).[6] The Greek pan- (πάν), meaning "all", and thēr (θήρ), meaning "prey" bears the meaning of "predator of all animals". Use of the word for a beast originated in antiquity in the Orient, probably from India to Persia to Greece.[7] In Mexican Spanish, its nickname is el tigre: 16th century Spaniards had no native word in their language for the jaguar, which is smaller than a lion, but bigger than a leopard, nor had ever encountered it in the Old World, and so named it after the tiger, since its ferocity would have been known to them through Roman writings and popular literature during the Renaissance.[8] Onca is the Portuguese onça, with the cedilla dropped for typographical reasons, found in English as ounce for the snow leopard, Panthera
Panthera
uncia. It derives from the Latin lyncea lynx, with the letter L confused with the definite article (Italian lonza, Old French l'once).[9] Evolution

Fossil
Fossil
skull of a Pleistocene
Pleistocene
North American jaguar
North American jaguar
( Panthera
Panthera
onca augusta)

The jaguar is the only extant New World
New World
member of the genus Panthera. Results of DNA
DNA
analysis shows the lion, tiger, leopard, jaguar, snow leopard, and clouded leopard share a common ancestor, and that this group is between six and ten million years old; the fossil record points to the emergence of Panthera
Panthera
just two to 3.8 million years ago.[10] The Panthera
Panthera
are thought to have evolved in Asia.[11] The jaguar is thought to have diverged from a common ancestor of the Panthera
Panthera
species at least 1.5 million years ago and to have entered the American continent in the Early Pleistocene
Pleistocene
via Beringia, the land bridge that once spanned the Bering Strait. Results of jaguar mitochondrial DNA
DNA
analysis indicates that the species' lineage evolved between 280,000 and 510,000 years ago.[12] Phylogenetic
Phylogenetic
studies generally have shown the clouded leopard ( Neofelis
Neofelis
nebulosa) is basal to this group.[13][14][15][11] The position of the remaining species varies between studies and is effectively unresolved. Based on morphological evidence, British zoologist Reginald Innes Pocock concluded the jaguar is most closely related to the leopard.[13] However, DNA
DNA
evidence is inconclusive and the position of the jaguar relative to the other species varies between studies.[13][14][15][11] Fossils of extinct Panthera
Panthera
species, such as the European jaguar ( Panthera
Panthera
gombaszoegensis) and the American lion ( Panthera
Panthera
atrox), show characteristics of both the lion and the jaguar.[13] Taxonomy In 1758, Carl Linnaeus
Carl Linnaeus
described the jaguar in his work Systema Naturae and gave it the scientific name Felis
Felis
onca.[16] In the 19th and 20th centuries, several jaguar type specimens formed the basis for descriptions of subspecies.[17] In 1939, Reginald Innes Pocock recognized eight subspecies based on geographic origins and skull morphology of these specimens.[18] Pocock did not have access to sufficient zoological specimens to critically evaluate their subspecific status, but expressed doubt about the status of several. Later consideration of his work suggested only three subspecies should be recognized. The description of P. o. palustris was based on a fossil skull.[19] The author of Mammal
Mammal
Species
Species
of the World listed nine subspecies and both P. o. palustris or P. o. paraguensis separately.[17] Results of morphologic and genetic research indicate a clinal north–south variation between populations, but no evidence for subspecific differentiation.[12][20] A subsequent, more detailed study confirmed the predicted population structure within jaguar populations in Colombia.[21] IUCN Red List
IUCN Red List
assessors for the species and the Cat
Cat
Classification Taskforce of the Cat Specialist Group
Cat Specialist Group
do not recognize any jaguar subspecies as valid.[1][22] The following table is based on the classification of the species provided in Mammal
Mammal
Species
Species
of the World.[17]

Formerly recognised subspecies Region Image

P. o. onca (Linnaeus, 1758) Peruvian jaguar (P. o. peruviana) (De Blainville, 1843) Pantanal
Pantanal
jaguar (P. o. palustris) or Paraguayan jaguar (P. o. paraguensis) (Nelson and Goldman, 1933)

South America: Venezuela
Venezuela
to the Amazon rainforest, coastal Peru, the Pantanal
Pantanal
regions of Mato Grosso
Mato Grosso
and Mato Grosso
Mato Grosso
do Sul, Brazil, along the Paraguay
Paraguay
River into Paraguay
Paraguay
and northeastern Argentina

Female South American jaguar
South American jaguar
at Piquiri River, Mato Grosso
Mato Grosso
state, Brazil

P. o. hernandesii (Gray, 1857) Central American jaguar (P. o. centralis) (Mearns, 1901) Arizona
Arizona
jaguar (P. o. arizonensis) (Goldman, 1932) P. o. goldmani (Goldman, 1932) P. o. veraecrucis (Nelson and Goldman, 1933)

Central and North America: Colombia, Guatemala, Belize, El Salvador
El Salvador
to Yucatán Peninsula, Mexico
Mexico
and southern Arizona, United States
United States
of America.

El Jefe in Arizona, the United States
United States
of America

Characteristics

The head of the jaguar is robust and the jaw extremely powerful. The size of jaguars tends to increase the farther south they are located.

The jaguar, a compact and well-muscled animal, is the largest cat in the New World
New World
and the largest carnivorous mammal in Central and South America.[23] Its coat is generally a tawny yellow, but ranges to reddish-brown, for most of the body. The ventral areas are white.[24] The fur is covered with rosettes for camouflage in the dappled light of its forest habitat. The spots and their shapes vary between individual jaguars: rosettes may include one or several dots. The spots on the head and neck are generally solid, as are those on the tail, where they may merge to form a band.[19] Forest jaguars are frequently darker and considerably smaller than those in open areas, possibly due to the smaller numbers of large, herbivorous prey in forest areas.[25] Its size and weight vary considerably: weights are normally in the range of 56–96 kg (123–212 lb). Larger males have been recorded to weigh as much as 158 kg (348 lb)[26][27] The smallest females weigh about 36 kg (79 lb).[26] Females are typically 10–20 percent smaller than males. The length, from the nose to the base of the tail, varies from 1.12 to 1.85 m (3.7 to 6.1 ft). The tail is the shortest of any big cat, at 45 to 75 cm (18 to 30 in) in length.[26][28] Legs are also short, but thick and powerful, considerably shorter when compared to a small tiger or lion in a similar weight range. The jaguar stands 63 to 76 cm (25 to 30 in) tall at the shoulders.[24] Compared to the similarly colored leopard, the jaguar is bigger, heavier and relatively stocky in build.[19] Further variations in size have been observed across regions and habitats, with size tending to increase from north to south. Jaguars in the Chamela-Cuixmala Biosphere Reserve
Chamela-Cuixmala Biosphere Reserve
on the Mexican Pacific coast weighed just about 50 kg (110 lb), about the size of a female cougar.[29] Jaguars in Venezuela
Venezuela
or Brazil
Brazil
are much larger with average weights of about 95 kg (209 lb) in males and of about 56–78 kg (123–172 lb) in females.[19] In the Brazilian Pantanal, weights of 136 kg (300 lb) or more are not uncommon in old males.[30] The highest recorded weight was 148 kg (326 lb) of a jaguar with an empty stomach. A short and stocky limb structure makes the jaguar adept at climbing, crawling, and swimming.[24] The head is robust and the jaw extremely powerful, it has the third highest bite force of all felids, after the tiger and lion.[31] A 100 kg (220 lb) jaguar can bite with a force of 503.6 kgf (1,110 lbf) at canine teeth and705.8 kgf (1,556 lbf) at carnassial notch.[32] This allows it to pierce the shells of armored reptiles and turtles.[33] A comparative study of bite force adjusted for body size ranked it as the top felid, alongside the clouded leopard and ahead of the tiger and lion.[34] It has been reported that "an individual jaguar can drag an 800 lb (360 kg) bull 25 ft (7.6 m) in its jaws and pulverize the heaviest bones".[35] While the jaguar closely resembles the leopard, it is sturdier and heavier, and the two animals can be distinguished by their rosettes: the rosettes on a jaguar's coat are larger, fewer in number, usually darker, and have thicker lines and small spots in the middle that the leopard lacks. Jaguars also have rounder heads and shorter, stockier limbs compared to leopards.[36] Color variation

A melanistic jaguar is a color morph which occurs at about 6 percent frequency in populations.

Color morphism occurs in the species. A near-black melanistic form occurs regularly. Jaguars with melanism appear entirely black, although their spots are still visible on close examination. The black morph is less common than the spotted form, but at about six percent of the population,[37] it is several orders of magnitude above the mutation rate. Hence, it is being supported by selection. Some evidence indicates the melanism allele is dominant.[38] The black form may be an example of heterozygote advantage; breeding in captivity is not yet conclusive on this. Melanistic jaguars (or “black” jaguars) occur primarily in parts of South America, and are virtually unknown in wild populations residing in the subtropical and temperate regions of North America; they have never been documented north of Mexico’s Isthmus of Tehuantepec.[39] Melanistic jaguars are informally known as black panthers, but (as with all forms of polymorphism) they do not form a separate species. Extremely rare albino individuals, sometimes called white panthers, also occur among jaguars, as with the other big cats.[25] As usual with albinos in the wild, selection keeps the frequency close to the rate of mutation. Distribution and habitat

The jaguar inhabits a variety of forested and open habitat, but is strongly associated with the presence of water.

At present, the jaguar's range extends from Mexico
Mexico
through Central America to South America, including much of Amazonian Brazil. The countries included in this range are Argentina, Belize, Bolivia, Colombia, Costa Rica (particularly on the Osa Peninsula), Ecuador, French Guiana, Guatemala, Guyana, Honduras, Nicaragua, Panama, Paraguay, Peru, Suriname, the United States
United States
and Venezuela. It is now locally extinct in El Salvador
El Salvador
and Uruguay.[1] The jaguar has been an American cat since crossing the Bering Land Bridge during the Pleistocene
Pleistocene
epoch; the immediate ancestor of modern animals is Panthera
Panthera
onca augusta, which was larger than the contemporary cat.[21] It occurs in the 400 km² Cockscomb Basin Wildlife Sanctuary in Belize, the 5,300 km² Sian Ka'an
Sian Ka'an
Biosphere Reserve in Mexico, the approximately 15,000 km2 Manú National Park in Peru, the approximately 26,000 km2 Xingu National Park
Xingu National Park
in Brazil, and numerous other reserves throughout its range. The inclusion of the United States
United States
in the list is based on occasional sightings in the southwest, particularly in Arizona,[40] New Mexico and Texas. In the early 20th century, the jaguar's range extended as far north as the Grand Canyon
Grand Canyon
and possibly Colorado, and as far west as Monterey in Northern California.[41][42][43] The jaguar is a protected species in the United States
United States
under the Endangered Species Act, which has stopped the shooting of the animal for its pelt. In 1996 and from 2004 on, hunting guides and wildlife officials in Arizona
Arizona
photographed and documented jaguars in the southern part of the state.[44] Between 2004 and 2007, two or three jaguars have been reported by researchers around Buenos Aires National Wildlife Refuge in southern Arizona. One of them, called 'Macho B', had been previously photographed in 1996 in the area.[45] For any permanent population in the USA to thrive, protection from killing, an adequate prey base, and connectivity with Mexican populations are essential.[46] In February 2009, a 53.5 kg (118 lb) jaguar was caught, radio-collared and released in an area southwest of Tucson, Arizona; this is farther north than had previously been expected and represents a sign there may be a permanent breeding population of jaguars within southern Arizona. The animal was later confirmed to be indeed the same male individual ('Macho B') that was photographed in 2004.[47] On 2 March 2009, Macho B was recaptured and euthanized after he was found to be suffering from kidney failure; the animal was thought to be 16 years old, older than any known wild jaguar.[48] Completion of the United States– Mexico
Mexico
barrier as currently proposed will reduce the viability of any population currently residing in the United States, by reducing gene flow with Mexican populations, and prevent any further northward expansion for the species.[49] The historic range of the species included much of the southern half of the United States, and in the south extended much farther to cover most of the South American continent. In total, its northern range has receded 1,000 km (621 mi) southward and its southern range 2,000 km (1243 mi) northward. Ice age fossils of the jaguar, dated between 40,000 and 11,500 years ago, have been discovered in the United States, including some at an important site as far north as Missouri. Fossil
Fossil
evidence shows jaguars of up to 190 kg (420 lb), much larger than the contemporary average for the animal.[50] The habitat of the cat includes the rain forests of South and Central America, open, seasonally flooded wetlands, and dry grassland terrain. Of these habitats, the jaguar much prefers dense forest;[25] the cat has lost range most rapidly in regions of drier habitat, such as the Argentine pampas, the arid grasslands of Mexico, and the southwestern United States.[1] The cat will range across tropical, subtropical, and dry deciduous forests (including, historically, oak forests in the United States). The jaguar prefers to live by rivers, swamps, and in dense rainforest with thick cover for stalking prey. Jaguars have been found at elevations as high as 3,800 m, but they typically avoid montane forest and are not found in the high plateau of central Mexico or in the Andes.[25] The jaguars preferred habitats are usually swamps and wooded regions, but jaguars also live in scrublands and deserts.[51] Ecology and behavior Ecological role The adult jaguar is an apex predator, meaning it exists at the top of its food chain and is not preyed on in the wild. The jaguar has also been termed a keystone species, as it is assumed, through controlling the population levels of prey such as herbivorous and granivorous mammals, apex felids maintain the structural integrity of forest systems.[29][52] However, accurately determining what effect species like the jaguar have on ecosystems is difficult, because data must be compared from regions where the species is absent as well as its current habitats, while controlling for the effects of human activity. It is accepted that mid-sized prey species undergo population increases in the absence of the keystone predators, and this has been hypothesized to have cascading negative effects.[53] However, field work has shown this may be natural variability and the population increases may not be sustained. Thus, the keystone predator hypothesis is not accepted by all scientists.[54] The jaguar also has an effect on other predators. The jaguar and the cougar, the next-largest feline of the Americas, are often sympatric (related species sharing overlapping territory) and have often been studied in conjunction. Where sympatric with the jaguar, the cougar is smaller than normal and is smaller than the local jaguars. The jaguar tends to take larger prey, usually over 22 kg (49 lb) and the cougar smaller, usually between 2 and 22 kg (4.4 and 48.5 lb), reducing the latter's size.[55][56] This situation may be advantageous to the cougar. Its broader prey niche, including its ability to take smaller prey, may give it an advantage over the jaguar in human-altered landscapes;[29] while both are classified as near-threatened species, the cougar has a significantly larger current distribution. Reproduction and life cycle

4-month-old cub at the Salzburg Zoo

Jaguar
Jaguar
females reach sexual maturity at about two years of age, and males at three or four. The cat is believed to mate throughout the year in the wild, although births may increase when prey is plentiful.[57] Research on captive male jaguars supports the year-round mating hypothesis, with no seasonal variation in semen traits and ejaculatory quality; low reproductive success has also been observed in captivity.[58] Female estrus is 6–17 days out of a full 37-day cycle, and females will advertise fertility with urinary scent marks and increased vocalization.[57] Both sexes will range more widely than usual during courtship. Pairs separate after mating, and females provide all parenting. The gestation period lasts 93–105 days; females give birth to up to four cubs, and most commonly to two. The mother will not tolerate the presence of males after the birth of cubs, given a risk of infanticide; this behavior is also found in the tiger.[59] The young are born blind, gaining sight after two weeks. Cubs are weaned at three months, but remain in the birth den for six months before leaving to accompany their mother on hunts.[60] They will continue in their mother's company for one to two years before leaving to establish a territory for themselves. Young males are at first nomadic, jostling with their older counterparts until they succeed in claiming a territory. Typical lifespan in the wild is estimated at around 12–15 years; in captivity, the jaguar lives up to 23 years, placing it among the longest-lived cats.[30] Social activity

Jaguars at Cameron Park Zoo
Cameron Park Zoo
in Waco, Texas

Like most cats, the jaguar is solitary outside mother–cub groups. Adults generally meet only to court and mate (though limited noncourting socialization has been observed anecdotally[59]) and carve out large territories for themselves. Female territories, which range from 25 to 40 km2 in size, may overlap, but the animals generally avoid one another. Male ranges cover roughly twice as much area, varying in size with the availability of game and space, and do not overlap. The territory of a male can contain those of several females.[59][61] The jaguar uses scrape marks, urine, and feces to mark its territory.[62][63] Like the other big cats, the jaguar is capable of roaring[64][65] and does so to warn territorial and mating competitors away; intensive bouts of counter-calling between individuals have been observed in the wild.[33] Their roar often resembles a repetitive cough, and they may also vocalize mews and grunts.[30] Mating
Mating
fights between males occur, but are rare, and aggression avoidance behavior has been observed in the wild.[62] When it occurs, conflict is typically over territory: a male's range may encompass that of two or three females, and he will not tolerate intrusions by other adult males.[59] The jaguar is often described as nocturnal, but is more specifically crepuscular (peak activity around dawn and dusk). Both sexes hunt, but males travel farther each day than females, befitting their larger territories. The jaguar may hunt during the day if game is available and is a relatively energetic feline, spending as much as 50–60 percent of its time active.[25] The jaguar's elusive nature and the inaccessibility of much of its preferred habitat make it a difficult animal to sight, let alone study. Hunting and diet

The jaguar has an exceptionally powerful bite that allows it to pierce the shells of armored prey

Like all cats, the jaguar is an obligate carnivore, feeding only on meat. It is an opportunistic hunter and its diet encompasses at least 87 species.[25] It employ an unusual killing method: it bites directly through the skull of prey between the ears to deliver a fatal bite to the brain.[66] The jaguar can take virtually any terrestrial or riparian vertebrate found in Central or South America, except for large crocodilians such as black caiman. The jaguar is more of a dietary generalist than its Old World cousins: the American tropics have a high diversity of small animals but relatively low populations and diversity of the large ungulates which this genus favors.[55] They regularly take adult caimans, except for black caimans,[67] deer, capybaras, tapirs, peccaries, dogs, zorros, and sometimes even anacondas.[19] However, it preys on any small species available, including frogs, mice, birds (mainly ground-based species such as cracids), fish, sloths, monkeys, and turtles. A study conducted in Cockscomb Basin Wildlife Sanctuary
Cockscomb Basin Wildlife Sanctuary
in Belize
Belize
revealed that the diet of jaguars there consisted primarily of armadillos and pacas.[62] Some jaguars will also take domestic livestock.[68] El Jefe, one of the few jaguars that were reported in the United States,[69][70] has also been found to kill and eat American black bears, as deduced from hairs found within his scats and the partly consumed carcass of a black bear sow with the distinctive puncture marks to the skull left by jaguars. This indicates that jaguars might have once preyed on black bears when the species was still present in the area. Spectacled bears
Spectacled bears
are also known to avoid jaguars, possibly because they may constitute occasional prey items.[71] There is evidence that jaguars in the wild consume the roots of Banisteriopsis caapi.[72] While the jaguar often employs the deep throat-bite and suffocation technique typical among Panthera, it sometimes uses a killing method unique amongst cats: it pierces directly through the temporal bones of the skull between the ears of prey (especially the capybara) with its canine teeth, piercing the brain.[73] This may be an adaptation to "cracking open" turtle shells; following the late Pleistocene extinctions, armored reptiles such as turtles would have formed an abundant prey base for the jaguar.[33][25] The skull bite is employed with mammals in particular; with reptiles such as the caiman, the jaguar may leap onto the back of the prey and sever the cervical vertebrae, immobilizing the target. When attacking sea turtles, including the huge leatherback sea turtle which weighs about 385 kg (849 lb) on average, as they try to nest on beaches, the jaguar will bite at the head, often beheading the prey, before dragging it off to eat.[74] Reportedly, while hunting horses, a jaguar may leap onto their back, place one paw on the muzzle and another on the nape and then twist, dislocating the neck. Local people have anecdotally reported that when hunting a pair of horses bound together, the jaguar will kill one horse and then drag it while the other horse, still living, is dragged in their wake.[75] With prey such as smaller dogs, a paw swipe to the skull may be sufficient to kill it.

Illustration of a jaguar killing a tapir, the largest native land animal in its range

The jaguar is a stalk-and-ambush rather than a chase predator. The cat will walk slowly down forest paths, listening for and stalking prey before rushing or ambushing. The jaguar attacks from cover and usually from a target's blind spot with a quick pounce; the species' ambushing abilities are considered nearly peerless in the animal kingdom by both indigenous people and field researchers, and are probably a product of its role as an apex predator in several different environments. The ambush may include leaping into water after prey, as a jaguar is quite capable of carrying a large kill while swimming; its strength is such that carcasses as large as a heifer can be hauled up a tree to avoid flood levels.[59] On killing prey, the jaguar will drag the carcass to a thicket or other secluded spot. It begins eating at the neck and chest, rather than the midsection. The heart and lungs are consumed, followed by the shoulders.[59] The daily food requirement of a 34 kg (75 lb) animal, at the extreme low end of the species' weight range, has been estimated at 1.4 kg (3.1 lb).[41] For captive animals in the 50–60 kg (110–130 lb) range, more than 2 kg (4.4 lb) of meat daily are recommended.[76] In the wild, consumption is naturally more erratic; wild cats expend considerable energy in the capture and kill of prey, and they may consume up to 25 kg (55 lb) of meat at one feeding, followed by periods of famine.[77] Man-eating Further information: Man-eater Unlike all other Panthera
Panthera
species, jaguars very rarely attack humans. However, jaguar attacks appear to be on the rise with increased human encroachment on their habitat and a decrease in prey populations.[78] Sometimes jaguars in captivity attack zookeepers.[79] In addition, it appears that attacks on humans had been more common in the past, at least after conquistadors arrived in the Americas, to the extent that the jaguar had a fearsome reputation in the Americas, akin to the lion and tiger in the Old World. Nevertheless, even in those times, the jaguar's chief prey was the capybara, not the human, and Charles Darwin reported a saying of Native Americans that people would not have to fear the jaguar much, as long as capybaras were abundant.[80] Threats

A South American jaguar
South American jaguar
killed by Theodore Roosevelt

jaguar hunting scene with dogs – MHNT

Jaguar
Jaguar
populations are rapidly declining. The species is listed as Near Threatened
Near Threatened
on the IUCN Red List. The loss of parts of its range, including its virtual elimination from its historic northern areas and the increasing fragmentation of the remaining range, have contributed to this status.[1] Particularly significant declines occurred in the 1960, when more than 15,000 jaguars were killed for their skins in the Brazilian Amazon
Brazilian Amazon
yearly; the Convention on International Trade in Endangered Species
Species
of 1973 brought about a sharp decline in the pelt trade.[81] Detailed work performed under the auspices of the Wildlife Conservation Society revealed the species has lost 37% of its historic range, with its status unknown in an additional 18% of the global range. More encouragingly, the probability of long-term survival was considered high in 70% of its remaining range, particularly in the Amazon basin
Amazon basin
and the adjoining Gran Chaco and Pantanal.[82] The major risks to the jaguar include deforestation across its habitat, increasing competition for food with human beings, especially in dry and unproductive habitat,[1][83] poaching, hurricanes in northern parts of its range, and the behavior of ranchers who will often kill the cat where it preys on livestock. When adapted to the prey, the jaguar has been shown to take cattle as a large portion of its diet; while land clearance for grazing is a problem for the species, the jaguar population may have increased when cattle were first introduced to South America, as the animals took advantage of the new prey base. This willingness to take livestock has induced ranch owners to hire full-time jaguar hunters.[30] The skins of wild cats and other mammals have been highly valued by the fur trade for many decades. From the beginning of the 20th-century Jaguars were hunted in large numbers, but over-harvest and habitat destruction reduced the availability and induced hunters and traders to gradually shift to smaller species by the 1960s. The international trade of jaguar skins had its largest boom between the end of the Second World War
Second World War
and the early 1970, due to the growing economy and lack of regulations. From 1967 onwards, the regulations introduced by national laws and international agreements diminished the reported international trade from as high as 13000 skins in 1967, through 7000 skins in 1969, until it became negligible after 1976, although illegal trade and smuggling continue to be a problem. During this period, the biggest exporters were Brazil
Brazil
and Paraguay, and the biggest importers were the USA and Germany.[84] Conservation

A melanistic jaguar

Adult jaguar in Cameron Park Zoo, Waco, Texas

The jaguar is listed on CITES
CITES
Appendix I, which means that all international trade in jaguars or their parts is prohibited. Hunting jaguars is prohibited in Argentina, Brazil, Colombia, French Guiana, Honduras, Nicaragua, Panama, Paraguay, Suriname, the United States, and Venezuela. Hunting of jaguars is restricted in Brazil, Costa Rica, Guatemala, Mexico
Mexico
and Peru.[1] Trophy hunting
Trophy hunting
is still permitted in Bolivia, and it is not protected in Ecuador
Ecuador
or Guyana. In the US, it is listed as endangered under the Endangered Species
Species
Act. Jaguar
Jaguar
Conservation Units Jaguar
Jaguar
conservation is complicated because of the species' large range spaning 18 countries with different policies and regulations. Specific areas of high importance for jaguar conservation, so-called "Jaguar Conservation Units" (JCU) were determined in 2000. These are large areas inhabited by at least 50 jaguars. Each unit was assessed and evaluated on the basis of size, connectivity, habitat quality for both jaguar and prey, and jaguar population status. That way, 51 Jaguar Conservation Units were determined in 36 geographic regions as priority areas for jaguar conservation including:[82]

the Sierra Madre of Mexico the Selva Maya tropical forests extending over Mexico, Belize, and Guatemala the Chocó-Darién moist forests
Chocó-Darién moist forests
from Honduras, Panama
Panama
to Colombia Sierra de Tamaulipas Venezuelan Llanos northern Cerrado
Cerrado
and Amazon basin
Amazon basin
in Brazil Misiones Province
Misiones Province
in Argentina

Recent studies underlined that to maintain the robust exchange across the jaguar gene pool necessary for maintaining the species, it is important that jaguar habitats are interconnected. To facilitate this, a new project, the Paseo del Jaguar, has been established to connect several jaguar hotspots.[85] In 1986, the Cockscomb Basin Wildlife Sanctuary
Cockscomb Basin Wildlife Sanctuary
was established in Belize
Belize
as the world's first protected area for jaguar conservation.[86] Given the inaccessibility of much of the species' range, particularly the central Amazon, estimating jaguar numbers is difficult. Researchers typically focus on particular bioregions, thus species-wide analysis is scant. In 1991, 600–1,000 (the highest total) were estimated to be living in Belize. A year earlier, 125–180 jaguars were estimated to be living in Mexico's 4,000-km2 (2400-mi2) Calakmul Biosphere
Biosphere
Reserve, with another 350 in the state of Chiapas. The adjoining Maya Biosphere
Biosphere
Reserve in Guatemala, with an area measuring 15,000 km2 (9,000 mi2), may have 465–550 animals.[87] Work employing GPS telemetry in 2003 and 2004 found densities of only six to seven jaguars per 100 km2 in the critical Pantanal
Pantanal
region, compared with 10 to 11 using traditional methods; this suggests the widely used sampling methods may inflate the actual numbers of cats.[88] Approaches In setting up protected reserves, efforts generally also have to be focused on the surrounding areas, as jaguars are unlikely to confine themselves to the bounds of a reservation, especially if the population is increasing in size. Human attitudes in the areas surrounding reserves and laws and regulations to prevent poaching are essential to make conservation areas effective.[89] To estimate population sizes within specific areas and to keep track of individual jaguars, camera trapping and wildlife tracking telemetry are widely used, and feces may be sought out with the help of detector dogs to study jaguar health and diet.[90][91] Current conservation efforts often focus on educating ranch owners and promoting ecotourism.[92] The jaguar is generally defined as an umbrella species – its home range and habitat requirements are sufficiently broad that, if protected, numerous other species of smaller range will also be protected.[93] Umbrella species serve as "mobile links" at the landscape scale, in the jaguar's case through predation. Conservation organizations may thus focus on providing viable, connected habitat for the jaguar, with the knowledge other species will also benefit.[92] Ecotourism
Ecotourism
setups are being used to generate public interest in charismatic animals such as the jaguar, while at the same time generating revenue that can be used in conservation efforts. Audits done in Africa have shown that ecotourism has helped in African cat conservation. As with large African cats, a key concern in jaguar ecotourism is the considerable habitat space the species requires, so if ecotourism is used to aid in jaguar conservation, some considerations need to be made as to how existing ecosystems will be kept intact, or how new ecosystems that are large enough to support a growing jaguar population will be put into place.[94] The United States Main article: North American jaguar The only extant cat native to North America that roars,[95] the jaguar was recorded as an animal of the Americas
Americas
by Thomas Jefferson
Thomas Jefferson
in 1799.[96] Jaguars are still occasionally sighted in Arizona
Arizona
and New Mexico, such as El Jefe,[97][98] prompting actions for its conservation by authorities.[95] For example, on the 20th of August, 2012, the USFWS proposed setting aside 838,232 acres in Arizona
Arizona
and New Mexico
Mexico
— an area larger than Rhode Island — as critical jaguar habitat.[99] In mythology and culture Pre-Columbian Americas

Jaguar warrior
Jaguar warrior
in the Aztec
Aztec
culture

Copy of the Book of Chilam Balam
Chilam Balam
of Ixil in the National Museum of Anthropology (Mexico)

Statuette of Karajà in the Museum of Toulouse

Moche jaguar figurine dating to 300 CE, at the Larco Museum
Larco Museum
in Lima, Peru

See also: Jaguars in Mesoamerican cultures In pre-Columbian Central and South America, the jaguar was a symbol of power and strength. Among the Andean cultures, a jaguar cult disseminated by the early Chavín culture
Chavín culture
became accepted over most of what is today Peru
Peru
by 900 BC. The later Moche culture of northern Peru used the jaguar as a symbol of power in many of their ceramics.[100][101][102] In the religion of the Muisca, who inhabited the cool Altiplano Cundiboyacense
Altiplano Cundiboyacense
in the Colombian Andes, the jaguar was considered a sacred animal and during their religious rituals the people dressed in jaguar skins.[103] The skins were traded with the lowland peoples of the tropical Llanos Orientales.[104] The name of zipa Nemequene
Nemequene
was derived from the Muysccubun words nymy and quyne, meaning "force of the jaguar".[105][106] In Mesoamerica, the Olmec—an early and influential culture of the Gulf Coast region roughly contemporaneous with the Chavín—developed a distinct "were-jaguar" motif of sculptures and figurines showing stylised jaguars or humans with jaguar characteristics. In the later Maya civilization, the jaguar was believed to facilitate communication between the living and the dead and to protect the royal household. The Maya saw these powerful felines as their companions in the spiritual world, and a number of Maya rulers bore names that incorporated the Mayan word for jaguar (b'alam in many of the Mayan languages). Balam (Jaguar) remains a common Maya surname, and it is also the name of Chilam Balam, a legendary author to whom are attributed 17th and 18th-centuries Maya miscellanies preserving much important knowledge. The Aztec
Aztec
civilization shared this image of the jaguar as the representative of the ruler and as a warrior. The Aztecs formed an elite warrior class known as the Jaguar
Jaguar
Knights. In Aztec mythology, the jaguar was considered to be the totem animal of the powerful deity Tezcatlipoca.[59][107] Contemporary culture The jaguar and its name are widely used as a symbol in contemporary culture. It is the national animal of Guyana, and is featured in its coat of arms.[108] The flag of the Department of Amazonas, a Colombian department, features a black jaguar silhouette pouncing towards a hunter.[109] The jaguar also appears in banknotes of Brazilian real. The jaguar is also a common fixture in the mythology of many contemporary native cultures in South America,[110] usually being portrayed as the creature which gave humans the power over fire. Jaguar
Jaguar
is widely used as a product name, most prominently for a British luxury car brand. The name has been adopted by sports franchises, including the NFL's Jacksonville Jaguars
Jacksonville Jaguars
and the Mexican soccer club Chiapas
Chiapas
F.C. The crest of Argentina's national federation in rugby union features a jaguar; however, because of a journalist error, the country's national team is nicknamed Los Pumas.[111] In the spirit of the ancient Mayan culture, the 1968 Olympics in Mexico
Mexico
City adopted a red jaguar as the first official Olympic mascot.[112] References

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S.A. p. 231. ISBN 978-958-14-0368-4.  ^ Kruschek, Michael H. (2003). The evolution of the Bogotá chiefdom: A household view (PhD) (PDF). University of Pittsburgh. p. 14. Retrieved 11 January 2017.  ^ "nymy" (in Spanish). Muysccubun Dictionary Online. Retrieved 11 January 2017.  ^ "quyne" (in Spanish). Muysccubun Dictionary Online. Retrieved 11 January 2017.  ^ Christenson, Allen J. (2007). Popol Vuh: The Sacred Book of the Maya. University of Oklahoma Press. p. 196. ISBN 978-0-8061-3839-8. Retrieved 11 December 2011.  ^ "Guyana". RBC Radio. Archived from the original on 6 October 2014. Retrieved 11 November 2011.  ^ Gutterman, D. (2008-07-26). "Amazonas Department (Colombia)". Fotw.net. Archived from the original on 2011-08-05. Retrieved 2010-04-02.  ^ Levi-Strauss, Claude (2004) [1964]. O Cru e o Cozido. São Paulo: Cosac & Naify. Retrieved 2011-11-11.  ^ Davies, Sean (26 July 2007). "Puma power: Argentinian rugby". bbc.co.uk. Retrieved 8 October 2007.  ^ Welch, Paula. Cute Little Creatures: Mascots Lend a Smile to the Games (PDF). la84foundation.org. Retrieved 2011-11-11. 

Bibliography

Baker, W. K. Jr.; et al. Law, Christopher, ed. Guidelines for Captive Management of Jaguars (PDF). Jaguar
Jaguar
Species
Species
Survival Plan. American Zoo and Aquarium Association. Archived from the original on 13 January 2012. Retrieved 2011-11-11. CS1 maint: BOT: original-url status unknown (link)

Further reading

Brown, David, and Carlos A. López González (2001). Borderland Jaguars. University of Utah Press. ISBN 978-0-87480-696-0.

External links

Wikimedia Commons has media related to Panthera
Panthera
onca.

Wikispecies
Wikispecies
has information related to Panthera
Panthera
onca

" Species
Species
portrait Jaguar". International Union for Conservation of Nature Species
Species
Survival Commission Cat
Cat
Specialist Group. Archived from the original on 2015-03-07.  Sound of a jaguar roar at Vivanatura.org " Jaguar
Jaguar
( Panthera
Panthera
onca)". Arkive. Archived from the original on 2014-12-28.  "Jaguar". BBC. 2014. Archived from the original on 2014-09-22.  People and Jaguars a Guide for Coexistence Sky Island Alliance website Felidae
Felidae
Conservation Fund  "Jaguar". Encyclopedia Americana. 1920.  Salman the fat jaguar at Delhi Zoo, India

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v t e

Extant Carnivora
Carnivora
species

Kingdom: Animalia Phylum: Chordata Class: Mammalia Infraclass: Eutheria Superorder: Laurasiatheria

Suborder Feliformia

Nandiniidae

Nandinia

African palm civet
African palm civet
(N. binotata)

Herpestidae (Mongooses)

Atilax

Marsh mongoose
Marsh mongoose
(A. paludinosus)

Bdeogale

Bushy-tailed mongoose
Bushy-tailed mongoose
(B. crassicauda) Jackson's mongoose
Jackson's mongoose
(B. jacksoni) Black-footed mongoose
Black-footed mongoose
(B. nigripes)

Crossarchus

Alexander's kusimanse
Alexander's kusimanse
(C. alexandri) Angolan kusimanse
Angolan kusimanse
(C. ansorgei) Common kusimanse
Common kusimanse
(C. obscurus) Flat-headed kusimanse
Flat-headed kusimanse
(C. platycephalus)

Cynictis

Yellow mongoose
Yellow mongoose
(C. penicillata)

Dologale

Pousargues's mongoose
Pousargues's mongoose
(D. dybowskii)

Galerella

Angolan slender mongoose
Angolan slender mongoose
(G. flavescens) Black mongoose
Black mongoose
(G. nigrata) Somalian slender mongoose
Somalian slender mongoose
(G. ochracea) Cape gray mongoose
Cape gray mongoose
(G. pulverulenta) Slender mongoose
Slender mongoose
(G. sanguinea)

Helogale

Ethiopian dwarf mongoose
Ethiopian dwarf mongoose
(H. hirtula) Common dwarf mongoose
Common dwarf mongoose
(H. parvula)

Herpestes

Short-tailed mongoose
Short-tailed mongoose
(H. brachyurus) Indian gray mongoose
Indian gray mongoose
(H. edwardsii) Indian brown mongoose
Indian brown mongoose
(H. fuscus) Egyptian mongoose
Egyptian mongoose
(H. ichneumon) Small Asian mongoose
Small Asian mongoose
(H. javanicus) Long-nosed mongoose
Long-nosed mongoose
(H. naso) Collared mongoose
Collared mongoose
(H. semitorquatus) Ruddy mongoose
Ruddy mongoose
(H. smithii) Crab-eating mongoose
Crab-eating mongoose
(H. urva) Stripe-necked mongoose
Stripe-necked mongoose
(H. vitticollis)

Ichneumia

White-tailed mongoose
White-tailed mongoose
(I. albicauda)

Liberiictus

Liberian mongoose
Liberian mongoose
(L. kuhni)

Mungos

Gambian mongoose
Gambian mongoose
(M. gambianus) Banded mongoose
Banded mongoose
(M. mungo)

Paracynictis

Selous' mongoose
Selous' mongoose
(P. selousi)

Rhynchogale

Meller's mongoose
Meller's mongoose
(R. melleri)

Suricata

Meerkat
Meerkat
(S. suricatta)

Hyaenidae (Hyenas)

Crocuta

Spotted hyena
Spotted hyena
(C. crocuta)

Hyaena

Brown hyena
Brown hyena
(H. brunnea) Striped hyena
Striped hyena
(H. hyaena)

Proteles

Aardwolf
Aardwolf
(P. cristatus)

Felidae

Large family listed below

Viverridae

Large family listed below

Eupleridae

Small family listed below

Family Felidae

Felinae

Acinonyx

Cheetah
Cheetah
(A. jubatus)

Caracal

Caracal
Caracal
(C. caracal) African golden cat
African golden cat
(C. aurata)

Catopuma

Bay cat
Bay cat
(C. badia) Asian golden cat
Asian golden cat
(C. temminckii)

Felis

European wildcat
European wildcat
(F. silvestris) African wildcat
African wildcat
(F. lybica) Jungle cat
Jungle cat
(F. chaus) Black-footed cat
Black-footed cat
(F. nigripes) Sand cat
Sand cat
(F. margarita) Chinese mountain cat
Chinese mountain cat
(F. bieti) Domestic cat (F. catus)

Leopardus

Ocelot
Ocelot
(L. pardalis) Margay
Margay
(L. wiedii) Pampas
Pampas
cat (L. colocola) Geoffroy's cat
Geoffroy's cat
(L. geoffroyi) Kodkod
Kodkod
(L. guigna) Andean mountain cat
Andean mountain cat
(L. jacobita) Oncilla
Oncilla
(L. tigrinus) Southern tigrina
Southern tigrina
(L. guttulus)

Leptailurus

Serval
Serval
(L. serval)

Lynx

Canadian lynx (L. canadensis) Eurasian lynx
Eurasian lynx
(L. lynx) Iberian lynx
Iberian lynx
(L. pardinus) Bobcat
Bobcat
(L. rufus)

Otocolobus

Pallas's cat
Pallas's cat
(O. manul)

Pardofelis

Marbled cat
Marbled cat
(P. marmorata)

Prionailurus

Fishing cat
Fishing cat
(P. viverrinus) Leopard
Leopard
cat (P. bengalensis) Sundaland leopard cat (P. javanensis) Flat-headed cat
Flat-headed cat
(P. planiceps) Rusty-spotted cat
Rusty-spotted cat
(P. rubiginosus)

Puma

Cougar
Cougar
(P. concolor)

Herpailurus

Jaguarundi
Jaguarundi
(H. yagouaroundi)

Pantherinae

Panthera

Lion
Lion
(P. leo) Jaguar
Jaguar
(P. onca) Leopard
Leopard
(P. pardus) Tiger
Tiger
(P. tigris) Snow leopard
Snow leopard
(P. uncia)

Neofelis

Clouded leopard
Clouded leopard
(N. nebulosa) Sunda clouded leopard
Sunda clouded leopard
(N. diardi)

Family Viverridae
Viverridae
(includes Civets)

Paradoxurinae

Arctictis

Binturong
Binturong
(A. binturong)

Arctogalidia

Small-toothed palm civet
Small-toothed palm civet
(A. trivirgata)

Macrogalidia

Sulawesi palm civet
Sulawesi palm civet
(M. musschenbroekii)

Paguma

Masked palm civet
Masked palm civet
(P. larvata)

Paradoxurus

Golden wet-zone palm civet (P. aureus) Asian palm civet
Asian palm civet
(P. hermaphroditus) Jerdon's palm civet (P. jerdoni) Golden palm civet
Golden palm civet
(P. zeylonensis)

Hemigalinae

Chrotogale

Owston's palm civet
Owston's palm civet
(C. owstoni)

Cynogale

Otter civet
Otter civet
(C. bennettii)

Diplogale

Hose's palm civet
Hose's palm civet
(D. hosei)

Hemigalus

Banded palm civet
Banded palm civet
(H. derbyanus)

Prionodontinae (Asiatic linsangs)

Prionodon

Banded linsang
Banded linsang
(P. linsang) Spotted linsang
Spotted linsang
(P. pardicolor)

Viverrinae

Civettictis

African civet
African civet
(C. civetta)

Genetta (Genets)

Abyssinian genet
Abyssinian genet
(G. abyssinica) Angolan genet
Angolan genet
(G. angolensis) Bourlon's genet
Bourlon's genet
(G. bourloni) Crested servaline genet
Crested servaline genet
(G. cristata) Common genet
Common genet
(G. genetta) Johnston's genet
Johnston's genet
(G. johnstoni) Rusty-spotted genet
Rusty-spotted genet
(G. maculata) Pardine genet
Pardine genet
(G. pardina) Aquatic genet
Aquatic genet
(G. piscivora) King genet
King genet
(G. poensis) Servaline genet
Servaline genet
(G. servalina) Haussa genet
Haussa genet
(G. thierryi) Cape genet
Cape genet
(G. tigrina) Giant forest genet
Giant forest genet
(G. victoriae)

Poiana

African linsang
African linsang
(P. richardsonii) Leighton's linsang
Leighton's linsang
(P. leightoni)

Viverra

Malabar large-spotted civet
Malabar large-spotted civet
(V. civettina) Large-spotted civet
Large-spotted civet
(V. megaspila) Malayan civet
Malayan civet
(V. tangalunga) Large Indian civet
Large Indian civet
(V. zibetha)

Viverricula

Small Indian civet
Small Indian civet
(V. indica)

Family Eupleridae

Euplerinae

Cryptoprocta

Fossa (C. ferox)

Eupleres

Eastern falanouc
Eastern falanouc
(E. goudotii) Western falanouc (E. major)

Fossa

Malagasy civet
Malagasy civet
(F. fossana)

Galidiinae

Galidia

Ring-tailed mongoose
Ring-tailed mongoose
(G. elegans)

Galidictis

Broad-striped Malagasy mongoose
Broad-striped Malagasy mongoose
(G. fasciata) Grandidier's mongoose
Grandidier's mongoose
(G. grandidieri)

Mungotictis

Narrow-striped mongoose
Narrow-striped mongoose
(M. decemlineata)

Salanoia

Brown-tailed mongoose
Brown-tailed mongoose
(S. concolor) Durrell's vontsira (S. durrelli)

Suborder Caniformia
Caniformia
(cont. below)

Ursidae (Bears)

Ailuropoda

Giant panda
Giant panda
(A. melanoleuca)

Helarctos

Sun bear
Sun bear
(H. malayanus)

Melursus

Sloth
Sloth
bear (M. ursinus)

Tremarctos

Spectacled bear
Spectacled bear
(T. ornatus)

Ursus

American black bear
American black bear
(U. americanus) Brown bear
Brown bear
(U. arctos) Polar bear
Polar bear
(U. maritimus) Asian black bear
Asian black bear
(U. thibetanus)

Mephitidae

Conepatus (Hog-nosed skunks)

Molina's hog-nosed skunk
Molina's hog-nosed skunk
(C. chinga) Humboldt's hog-nosed skunk
Humboldt's hog-nosed skunk
(C. humboldtii) American hog-nosed skunk
American hog-nosed skunk
(C. leuconotus) Striped hog-nosed skunk
Striped hog-nosed skunk
(C. semistriatus)

Mephitis

Hooded skunk
Hooded skunk
(M. macroura) Striped skunk
Striped skunk
(M. mephitis)

Mydaus

Sunda stink badger
Sunda stink badger
(M. javanensis) Palawan stink badger
Palawan stink badger
(M. marchei)

Spilogale (Spotted skunks)

Southern spotted skunk
Southern spotted skunk
(S. angustifrons) Western spotted skunk
Western spotted skunk
(S. gracilis) Eastern spotted skunk
Eastern spotted skunk
(S. putorius) Pygmy spotted skunk
Pygmy spotted skunk
(S. pygmaea)

Procyonidae

Bassaricyon (Olingos)

Eastern lowland olingo
Eastern lowland olingo
(B. alleni) Northern olingo
Northern olingo
(B. gabbii) Western lowland olingo
Western lowland olingo
(B. medius) Olinguito
Olinguito
(B. neblina)

Bassariscus

Ring-tailed cat
Ring-tailed cat
(B. astutus) Cacomistle
Cacomistle
(B. sumichrasti)

Nasua (Coatis inclusive)

White-nosed coati
White-nosed coati
(N. narica) South American coati
South American coati
(N. nasua)

Nasuella (Coatis inclusive)

Western mountain coati (N. olivacea) Eastern mountain coati (N. meridensis)

Potos

Kinkajou
Kinkajou
(P. flavus)

Procyon

Crab-eating raccoon
Crab-eating raccoon
(P. cancrivorus) Raccoon
Raccoon
(P. lotor) Cozumel raccoon
Cozumel raccoon
(P. pygmaeus)

Ailuridae

Ailurus

Red panda
Red panda
(A. fulgens)

Suborder Caniformia
Caniformia
(cont. above)

Otariidae (Eared seals) (includes fur seals and sea lions) ( Pinniped
Pinniped
inclusive)

Arctocephalus

South American fur seal
South American fur seal
(A. australis) Australasian fur seal (A. forsteri) Galápagos fur seal
Galápagos fur seal
(A. galapagoensis) Antarctic fur seal
Antarctic fur seal
(A. gazella) Juan Fernández fur seal
Juan Fernández fur seal
(A. philippii) Brown fur seal
Brown fur seal
(A. pusillus) Guadalupe fur seal
Guadalupe fur seal
(A. townsendi) Subantarctic fur seal
Subantarctic fur seal
(A. tropicalis)

Callorhinus

Northern fur seal
Northern fur seal
(C. ursinus)

Eumetopias

Steller sea lion
Steller sea lion
(E. jubatus)

Neophoca

Australian sea lion
Australian sea lion
(N. cinerea)

Otaria

South American sea lion
South American sea lion
(O. flavescens)

Phocarctos

New Zealand sea lion
New Zealand sea lion
(P. hookeri)

Zalophus

California sea lion
California sea lion
(Z. californianus) Galápagos sea lion
Galápagos sea lion
(Z. wollebaeki)

Odobenidae ( Pinniped
Pinniped
inclusive)

Odobenus

Walrus
Walrus
(O. rosmarus)

Phocidae (Earless seals) ( Pinniped
Pinniped
inclusive)

Cystophora

Hooded seal
Hooded seal
(C. cristata)

Erignathus

Bearded seal
Bearded seal
(E. barbatus)

Halichoerus

Gray seal (H. grypus)

Histriophoca

Ribbon seal
Ribbon seal
(H. fasciata)

Hydrurga

Leopard
Leopard
seal (H. leptonyx)

Leptonychotes

Weddell seal
Weddell seal
(L. weddellii)

Lobodon

Crabeater seal
Crabeater seal
(L. carcinophagus)

Mirounga (Elephant seals)

Northern elephant seal
Northern elephant seal
(M. angustirostris) Southern elephant seal
Southern elephant seal
(M. leonina)

Monachus

Mediterranean monk seal
Mediterranean monk seal
(M. monachus) Hawaiian monk seal
Hawaiian monk seal
(M. schauinslandi)

Ommatophoca

Ross seal
Ross seal
(O. rossi)

Pagophilus

Harp seal
Harp seal
(P. groenlandicus)

Phoca

Spotted seal
Spotted seal
(P. largha) Harbor seal
Harbor seal
(P. vitulina)

Pusa

Caspian seal
Caspian seal
(P. caspica) Ringed seal
Ringed seal
(P. hispida) Baikal seal
Baikal seal
(P. sibirica)

Canidae

Large family listed below

Mustelidae

Large family listed below

Family Canidae
Canidae
(includes dogs)

Atelocynus

Short-eared dog
Short-eared dog
(A. microtis)

Canis

Side-striped jackal
Side-striped jackal
(C. adustus) African golden wolf
African golden wolf
(C. anthus) Golden jackal
Golden jackal
(C. aureus) Coyote
Coyote
(C. latrans) Gray wolf
Gray wolf
(C. lupus) Black-backed jackal
Black-backed jackal
(C. mesomelas) Red wolf
Red wolf
(C. rufus) Ethiopian wolf
Ethiopian wolf
(C. simensis)

Cerdocyon

Crab-eating fox
Crab-eating fox
(C. thous)

Chrysocyon

Maned wolf
Maned wolf
(C. brachyurus)

Cuon

Dhole
Dhole
(C. alpinus)

Lycalopex

Culpeo
Culpeo
(L. culpaeus) Darwin's fox
Darwin's fox
(L. fulvipes) South American gray fox
South American gray fox
(L. griseus) Pampas
Pampas
fox (L. gymnocercus) Sechuran fox
Sechuran fox
(L. sechurae) Hoary fox
Hoary fox
(L. vetulus)

Lycaon

African wild dog
African wild dog
(L. pictus)

Nyctereutes

Raccoon
Raccoon
dog (N. procyonoides)

Otocyon

Bat-eared fox
Bat-eared fox
(O. megalotis)

Speothos

Bush dog
Bush dog
(S. venaticus)

Urocyon

Gray fox
Gray fox
(U. cinereoargenteus) Island fox
Island fox
(U. littoralis)

Vulpes (Foxes)

Bengal fox
Bengal fox
(V. bengalensis) Blanford's fox
Blanford's fox
(V. cana) Cape fox
Cape fox
(V. chama) Corsac fox
Corsac fox
(V. corsac) Tibetan sand fox
Tibetan sand fox
(V. ferrilata) Arctic fox
Arctic fox
(V. lagopus) Kit fox
Kit fox
(V. macrotis) Pale fox
Pale fox
(V. pallida) Rüppell's fox
Rüppell's fox
(V. rueppelli) Swift fox
Swift fox
(V. velox) Red fox
Red fox
(V. vulpes) Fennec fox
Fennec fox
(V. zerda)

Family Mustelidae

Lutrinae (Otters)

Aonyx

African clawless otter
African clawless otter
(A. capensis) Oriental small-clawed otter
Oriental small-clawed otter
(A. cinerea)

Enhydra

Sea otter
Sea otter
(E. lutris)

Hydrictis

Spotted-necked otter
Spotted-necked otter
(H. maculicollis)

Lontra

North American river otter
North American river otter
(L. canadensis) Marine otter
Marine otter
(L. felina) Neotropical otter
Neotropical otter
(L. longicaudis) Southern river otter
Southern river otter
(L. provocax)

Lutra

Eurasian otter
Eurasian otter
(L. lutra) Hairy-nosed otter
Hairy-nosed otter
(L. sumatrana)

Lutrogale

Smooth-coated otter
Smooth-coated otter
(L. perspicillata)

Pteronura

Giant otter
Giant otter
(P. brasiliensis)

Mustelinae (including badgers)

Arctonyx

Hog badger
Hog badger
(A. collaris)

Eira

Tayra
Tayra
(E. barbara)

Galictis

Lesser grison
Lesser grison
(G. cuja) Greater grison
Greater grison
(G. vittata)

Gulo

Wolverine
Wolverine
(G. gulo)

Ictonyx

Saharan striped polecat
Saharan striped polecat
(I. libyca) Striped polecat
Striped polecat
(I. striatus)

Lyncodon

Patagonian weasel
Patagonian weasel
(L. patagonicus)

Martes (Martens)

American marten
American marten
(M. americana) Yellow-throated marten
Yellow-throated marten
(M. flavigula) Beech marten
Beech marten
(M. foina) Nilgiri marten
Nilgiri marten
(M. gwatkinsii) European pine marten
European pine marten
(M. martes) Japanese marten
Japanese marten
(M. melampus) Sable
Sable
(M. zibellina)

Pekania

Fisher (P. pennanti)

Meles

Japanese badger
Japanese badger
(M. anakuma) Asian badger
Asian badger
(M. leucurus) European badger
European badger
(M. meles)

Mellivora

Honey badger
Honey badger
(M. capensis)

Melogale (Ferret-badgers)

Bornean ferret-badger
Bornean ferret-badger
(M. everetti) Chinese ferret-badger
Chinese ferret-badger
(M. moschata) Javan ferret-badger
Javan ferret-badger
(M. orientalis) Burmese ferret-badger
Burmese ferret-badger
(M. personata)

Mustela (Weasels and Ferrets)

Amazon weasel
Amazon weasel
(M. africana) Mountain weasel
Mountain weasel
(M. altaica) Stoat
Stoat
(M. erminea) Steppe polecat
Steppe polecat
(M. eversmannii) Colombian weasel
Colombian weasel
(M. felipei) Long-tailed weasel
Long-tailed weasel
(M. frenata) Japanese weasel
Japanese weasel
(M. itatsi) Yellow-bellied weasel
Yellow-bellied weasel
(M. kathiah) European mink
European mink
(M. lutreola) Indonesian mountain weasel
Indonesian mountain weasel
(M. lutreolina) Black-footed ferret
Black-footed ferret
(M. nigripes) Least weasel
Least weasel
(M. nivalis) Malayan weasel
Malayan weasel
(M. nudipes) European polecat
European polecat
(M. putorius) Siberian weasel
Siberian weasel
(M. sibirica) Back-striped weasel
Back-striped weasel
(M. strigidorsa) Egyptian weasel
Egyptian weasel
(M. subpalmata)

Neovison (Minks)

American mink
American mink
(N. vison)

Poecilogale

African striped weasel
African striped weasel
(P. albinucha)

Taxidea

American badger
American badger
(T. taxus)

Vormela

Marbled polecat
Marbled polecat
(V. peregusna)

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Taxon identifiers

Wd: Q35694 ADW: Panthera_onca ARKive: panthera-onca BioLib: 2015 EoL: 328606 EPPO: PNTHON Fossilworks: 49735 GBIF: 5219426 iNaturalist: 41970 ITIS: 180593 IUCN: 15953 MSW: 14000240 NCBI: 9690 Species+: 6385

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LCCN: sh8506

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