Iteaceae is a flowering plant family of trees and shrubs native to the eastern

USA The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territori ...

, southeastern

Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...

, and south and

Southeastern Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical south-eastern region of Asia, consisting of the regions that are situated south of mainland ...

. Some older taxonomic systems place the genus ''Itea'' in the family

Grossulariaceae ''Ribes'' is a genus of about 200 known species of flowering plants, most of them native to the temperate regions of the Northern Hemisphere. The various species are known as currants or gooseberries, and some are cultivated for their edible ...

. The APG III system of 2009 includes the former Pterostemonaceae in Iteaceae. Consequently, it now has two genera with a total of 18 known species.

Fossil record

The family is known from fossil flowers dating to the

Turonian The Turonian is, in the ICS' geologic timescale, the second age in the Late Cretaceous Epoch, or a stage in the Upper Cretaceous Series. It spans the time between 93.9 ± 0.8 Ma and 89.8 ± 1 Ma (million years ago). The Turonian is preceded b ...

age of the

Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', ...

that have been found in the

Raritan Formation The Raritan Formation is a Cretaceous (Turonian) geologic formation. Dinosaur remains are among the fossils that have been recovered from the formation, although none have yet been referred to a specific genus.Weishampel, et al. (2004). "Dinosaur ...

New Jersey New Jersey is a state in the Mid-Atlantic and Northeastern regions of the United States. It is bordered on the north and east by the state of New York; on the east, southeast, and south by the Atlantic Ocean; on the west by the Delaware ...

and from leaves dating to the

Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', " ...

found in the Klondike Mountain Formation,

Washington Washington commonly refers to: * Washington (state), United States * Washington, D.C., the capital of the United States ** A metonym for the federal government of the United States ** Washington metropolitan area, the metropolitan area centered o ...

. Some seeds and one fruit of †''Itea europea'' were found in two samples of muddy sediment from the Fossil Forest of Dunarobba. In northwestern

Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical ...

, it occurs with a some seeds and fruits in two sites of Early or Middle Pliocene age. These records suggest it was an accessory element of the mid-Pliocene

swamp forest Freshwater swamp forests, or flooded forests, are forests which are inundated with freshwater, either permanently or seasonally. They normally occur along the lower reaches of rivers and around freshwater lakes. Freshwater swamp forests are found ...

. Fossil pollen of ''Itea'' has been detected in a short section of the

Sarzana Sarzana (, ; lij, Sarzann-a) is a town, '' comune'' (municipality) and former short-lived Catholic bishopric in the Province of La Spezia, Liguria, Italy. It is east of Spezia, on the railway to Pisa, at the point where the railway to Parm ...

Basin in north-western part of

central Italy Central Italy ( it, Italia centrale or just ) is one of the five official statistical regions of Italy used by the National Institute of Statistics (ISTAT), a first-level NUTS region, and a European Parliament constituency. Regions Central I ...

tentatively assigned to the

Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...

-Pliocene transition. In this site a high pollen percentages of 12% suggest that ''Itea'' was an important element in the local vegetation. The nearest living relative of †''Itea europaea'' is the American species '' Itea virginica''.The role of central Italy as a centre of refuge for thermophilous plants in the late Cenozoic, Edoardo Martinetto, Acta Palaeobotan. 41(2): 299-319, 2001

References

External links

Iteaceae
in Stevens, P. F. (2001 onwards). Saxifragales Saxifragales families Turonian first appearances Extant Turonian first appearances {{Saxifragales-stub