Hypera postica
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''Hypera postica'', commonly known as the alfalfa weevil, is a species of beetle in the superfamily
Curculionoidea Weevils are beetles belonging to the superfamily Curculionoidea, known for their elongated snouts. They are usually small, less than in length, and herbivorous. Approximately 97,000 species of weevils are known. They belong to several families, ...
; it can be found in alfalfa fields throughout Europe. Considered a destructive threat to alfalfa production in North America, several accidental introductions have been successfully countered though the use of a variety of biological control species.


Description

The alfalfa weevil grows to a length of about . The rostrum or beak is short and broad. The frons is half as wide as the rostrum while the pronotum is broadest in the centre. The general colour of the insect is brown, with a dark mid-dorsal stripe. The larva has a distinctive black head and no legs; it is yellowish-green, with a white dorsal stripe and faint white lateral stripes. It is about long just before pupation. It
pupa A pupa ( la, pupa, "doll"; plural: ''pupae'') is the life stage of some insects undergoing transformation between immature and mature stages. Insects that go through a pupal stage are holometabolous: they go through four distinct stages in thei ...
tes in a white, pea-sized cocoon made of loosely-woven silk. It resembles the clover leaf weevil (''
Hypera punctata ''Hypera zoilus'' (syn. ''H. punctata''),
Crop Protection Compendium. CABI ...
''), but that species is nearly twice as large, the larvae have tan heads and they seldom cause much damage to alfalfa crops.


Life cycle

In
Illinois Illinois ( ) is a state in the Midwestern United States. Its largest metropolitan areas include the Chicago metropolitan area, and the Metro East section, of Greater St. Louis. Other smaller metropolitan areas include, Peoria and Rock ...
, some eggs are laid in the late fall or the winter, when weather conditions permit. Adults also overwinter and become increasingly active in March and April. Eggs are laid in batches of up to 25 inside alfalfa stems. The larvae feed for three or four weeks, moulting three times, before pupating in the cocoons they make. They emerge as adults in about one or two weeks. After feeding for a week or two, they may experience
aestivation Aestivation ( la, aestas (summer); also spelled estivation in American English) is a state of animal dormancy, similar to hibernation, although taking place in the summer rather than the winter. Aestivation is characterized by inactivity and a ...
during the remainder of the summer, in which they demonstrate a dampening of their metabolic, respiratory and nervous system activities. In fall, the adults hide in the crowns of alfalfa plants or move onto coarse vegetation in ditches or by fences or in nearby woodland.


Damage

Both the larvae and the adults are diurnal and feed on the foliage of alfalfa, the larvae doing the most damage. The adults eat the edges of the leaves, giving them a feathery appearance. At first, the larvae feed on terminal leaves leaving puncture marks, but they later move down the plant to feed on lower leaves. The leaves attacked are skeletonised as the larvae feed between the veins, and plant yields can be reduced by up to 15%. Sometimes the larvae are killed by the pathogenic fungus ''
Zoophthora phytonomi ''Zoophthora'' is a genus of fungi in the family Entomophthoraceae. Like other taxa in this family, ''Zoophthora'' species cause disease in insects and as such are considered entomopathogenic fungi. Like most entomopathogenic fungal taxa, ''Zoo ...
'', especially in warm and humid weather. They may also be parasitized by ichneumon wasps, '' Bathylplectes anurus'' and '' B. curculionis''. The female of these wasps is only about long, and lays an egg inside an early stage of the larva of the weevil. The developing wasp larva is a
parasitoid In evolutionary ecology, a parasitoid is an organism that lives in close association with its host (biology), host at the host's expense, eventually resulting in the death of the host. Parasitoidism is one of six major evolutionarily stable str ...
, living inside the weevil larva and devouring it, eventually pupating soon after its host. The wasp's brownish cocoon has a broad white band in the case of ''B. curculionis'', and a narrower, raised one in the case of ''B. anurus''. When it is disturbed, the cocoon of the latter species can "jump" several centimetres to avoid
predation Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill ...
. Both these species of wasp have been investigated for their potential for biological control of the weevil, and ''B. anurus'' is generally considered superior to ''B. curculionis'' because of its higher reproductive rate.


References

{{DEFAULTSORT:Hypera postica Entiminae Beetles described in 1813