''Ostrya'' is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...

of eight to 10 small

deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, ...

tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...

s belonging to the birch family

Betulaceae Betulaceae, the birch family, includes six genera of deciduous nut-bearing trees and shrubs, including the birches, alders, hazels, hornbeams, hazel-hornbeam, and hop-hornbeams numbering a total of 167 species. They are mostly natives of the ...

. Common names include hop-hornbeam and hophornbeam. It may also be called ironwood, a name shared with a number of other plants. The genus is native in southern

Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a subcontinent of Eurasia and it is located entirel ...

, southwest and eastern

Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an are ...

, and

North North is one of the four compass points or cardinal directions. It is the opposite of south and is perpendicular to east and west. ''North'' is a noun, adjective, or adverb indicating direction or geography. Etymology The word ''north ...

and

Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...

.Kew World Checklist of Selected Plant Families
/ref> They have a conical or irregular crown and a scaly, rough bark. They have alternate and double-toothed

birch A birch is a thin-leaved deciduous hardwood tree of the genus ''Betula'' (), in the family Betulaceae, which also includes alders, hazels, and hornbeams. It is closely related to the beech- oak family Fagaceae. The genus ''Betula'' contains ...

-like leaves 3–10 cm long. The

flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...

s are produced in spring, with male catkins 5–10 cm long and female

aments A catkin or ament is a slim, cylindrical flower cluster (a spike), with inconspicuous or no petals, usually wind-pollinated ( anemophilous) but sometimes insect-pollinated (as in ''Salix''). They contain many, usually unisexual flowers, arranged ...

2–5 cm long. The

fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...

form in pendulous clusters 3–8 cm long with 6–20

seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...

s; each seed is a small nut 2–4 mm long, fully enclosed in a bladder-like

involucre In botany, a bract is a modified or specialized leaf, especially one associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves. They may be smaller, larger, or o ...

. The

wood Wood is a porous and fibrous structural tissue found in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulose fibers that are strong in tension and embedded in a matrix of lignin ...

is very hard and heavy. The genus name ''Ostrya'' is derived from the Greek word (), which may be related to () "shell (of an animal)". Regarded as a weed tree by some foresters, this hard and stable wood was historically used to fashion plane soles. ''Ostrya'' species are used as food plants by the

larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. ...

e of some Lepidoptera species, including

winter moth :''In North America, "winter moth" usually denotes the invasive species ''Operophtera brumata'', but may also mean refer to a native species, '' Erannis tiliaria'' (linden looper) or '' Operophtera bruceata'' (bruce spanworm).'' The winter moth ( ...

walnut sphinx ''Amorpha juglandis'', the walnut sphinx, is the only species in the monotypic moth genus ''Amorpha'', which is in the family Sphingidae, erected by Jacob Hübner in 1809. The species was first described by James Edward Smith in 1797. Distribu ...

, and ''

Coleophora ostryae ''Coleophora ostryae'' is a moth of the family Coleophoridae found in North America, including Maryland and Ontario. The larvae feed on the leaves of ''Ostrya'', ''Carpinus'', ''Carya'', and ''Quercus An oak is a tree or shrub in the genus ...

''.

Species

''Ostrya'' has the following species: # ''

Ostrya carpinifolia ''Ostrya carpinifolia'', the European hop-hornbeam, is a tree in the family Betulaceae. It is the only species of the genus ''Ostrya'' that is native to Europe. The specific epithet ''carpinifolia'' means "hornbeam-leaved", from , the Latin word ...

'' Scop. – European hop-hornbeam - Mediterranean region of southern Europe, Middle-east,

Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a transcontinental country located mainly on the Anatolian Peninsula in Western Asia, with a small portion on the Balkan Peninsula in ...

Lebanon Lebanon ( , ar, لُبْنَان, translit=lubnān, ), officially the Republic of Lebanon () or the Lebanese Republic, is a country in Western Asia. It is located between Syria to Lebanon–Syria border, the north and east and Israel to Blue ...

, Caucasus # ''

Ostrya chisosensis ''Ostrya chisosensis'', common name Big Bend hop-hornbeam or Chisos hop-hornbeam, is a plant species endemic to Texas. It is known only from the Chisos Mountains inside Big Bend National Park, in Brewster County, although related populations in n ...

'' Correll – Chisos hophornbeam, Big Bend hophornbeam - endemic to Big Bend National Park in Texas # ''

Ostrya japonica ''Ostrya japonica'', known as East Asian hophornbeam, or Japanese hop-hornbeam, is a species of tree in the Betulaceae family growing to 25 m tall. It is native to Japan, Korea and China. In China, it occurs in temperate forests of southern Gansu ...

'' Sarg. – Japanese hophornbeam - Japan, Korea, northern China # ''

Ostrya knowltonii ''Ostrya knowltonii'' is a species of tree known by the common names Knowlton's hophornbeam, western hophornbeam, woolly hophornbeam, and wolf hophornbeam. It is also one of many trees called ironwood. It is native to Utah, Arizona, New Mexico, ...

'' Coville – Knowlton hophornbeam, western hophornbeam, wolf hophornbeam -

Utah Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to it ...

Arizona Arizona ( ; nv, Hoozdo Hahoodzo ; ood, Alĭ ṣonak ) is a state in the Southwestern United States. It is the 6th largest and the 14th most populous of the 50 states. Its capital and largest city is Phoenix. Arizona is part of the Fou ...

New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe , LargestCity = Albuquerque , LargestMetro = Tiguex , OfficialLang = None , Languages = English, Spanish ( New Mexican), Navajo, Ke ...

Texas Texas (, ; Spanish: ''Texas'', ''Tejas'') is a state in the South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2020, it is the second-largest U.S. state by ...

# '' Ostrya multinervis'' Rehd. – Central Chinese hop-hornbeam - central China # '' Ostrya rehderiana'' Chun – Zhejiang hop-hornbeam - Zhejiang Province in China #''

Ostrya trichocarpa ''Ostrya'' is a genus of eight to 10 small deciduous trees belonging to the birch family Betulaceae. Common names include hop-hornbeam and hophornbeam. It may also be called ironwood, a name shared with a number of other plants. The genus is na ...

'' D.Fang & Y.S.Wang –

Guangxi Province Guangxi (; ; alternately romanized as Kwanghsi; ; za, Gvangjsih, italics=yes), officially the Guangxi Zhuang Autonomous Region (GZAR), is an autonomous region of the People's Republic of China, located in South China and bordering Vietnam ( ...

in China # ''

Ostrya virginiana ''Ostrya virginiana'', the American hophornbeam, is a species of '' Ostrya'' native to eastern North America, from Nova Scotia west to southern Manitoba and eastern Wyoming, southeast to northern Florida and southwest to eastern Texas. Population ...

'' (Mill.) K. Koch – eastern hophornbeam, American hophornbeam, ironwood - eastern US, eastern Canada, Mexico, Guatemala, El Salvador, Honduras # '' Ostrya yunnanensis'' W.K.Hu – Yunnan hop-hornbeam -

Yunnan Province Yunnan , () is a landlocked province in the southwest of the People's Republic of China. The province spans approximately and has a population of 48.3 million (as of 2018). The capital of the province is Kunming. The province borders the ...

in China # †'' Ostrya oregoniana'' (fossil) # †'' Ostrya scholzii'' (fossil)

Fossil record

†''Ostrya scholzii'' fossil seeds of the Chattian stage, Oligocene, are known from the Oberleichtersbach Formation in the Rhön Mountains, central

Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...

.The floral change in the tertiary of the Rhön mountains (Germany) by Dieter Hans Mai - Acta Paleobotanica 47(1): 135-143, 2007.

References

* Rushforth, K. (1985). "Ostrya". ''The Plantsman'' 7: 208-212.
Flora of China: ''Ostrya''

Flora Europaea: ''Ostrya''

External links

* {{Taxonbar, from=Q605290 Fagales genera