Hipposiderinae
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The Hipposideridae are a
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
of
bat Bats are mammals of the order Chiroptera.''cheir'', "hand" and πτερόν''pteron'', "wing". With their forelimbs adapted as wings, they are the only mammals capable of true and sustained flight. Bats are more agile in flight than most ...
s commonly known as the Old World leaf-nosed bats. While it has often been seen as a subfamily, Hipposiderinae, of the family Rhinolophidae, it is now more generally classified as its own family.Simmons, 2005, p. 365 Nevertheless, it is most closely related to Rhinolophidae within the suborder
Yinpterochiroptera The Yinpterochiroptera (or Pteropodiformes) is a suborder of the Chiroptera, which includes taxa formerly known as megabats and five of the microbat families: Rhinopomatidae, Rhinolophidae, Hipposideridae, Craseonycteridae, and Megadermatid ...
.


Taxonomy

The Hipposideridae contain 10 living genera and more than 70 species, mostly in the widespread genus ''
Hipposideros ''Hipposideros'' is one of the most diverse genera of bats, with more than 70 species. They are collectively called roundleaf bats after the shape of their nasal ornament. It is the type genus of the family Hipposideridae. It is divided into s ...
''. In addition, several fossil genera are known; the oldest fossils attributed to the family are from the
middle Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', "da ...
of Europe. In their 1997 ''Classification of Mammals'', Malcolm C. McKenna and Susan K. Bell proposed a division of Hipposideridae (called Rhinonycterinae in their work) into three
tribes The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide usage of the term in English is in the discipline of anthropology. This definition is contested, in part due to confli ...
, one with two subtribes, but these tribes turned out to be non- monophyletic and have been abandoned. A different classification was proposed by Hand and Kirsch in 2003. In 2009, Petr Benda and Peter Vallo proposed a separate tribe, Triaenopini, for the genera ''
Triaenops ''Triaenops'' is a genus of bat in the family Hipposideridae. It is classified in the tribe Triaenopini, along with the closely related genus '' Paratriaenops'' and perhaps the poorly known '' Cloeotis''. The species of ''Paratriaenops'', which ...
'', '' Paratriaenops'', and possibly '' Cloeotis'', synonymised in a 2014 revision (Foley, ''et al.'') that elevated the family
Rhinonycteridae Rhinonycteridae is a family of bats, allied to the suborder Microchiroptera. The type species, the orange nose-leafed species group ''Rhinonicteris aurantia'', is found across the north of Australia. Description The family accords with the de ...
.


Genera

The genera included in Hipposideridae are:


Living

*'' Anthops'' (one species;
Solomon Islands Solomon Islands is an island country consisting of six major islands and over 900 smaller islands in Oceania, to the east of Papua New Guinea and north-west of Vanuatu. It has a land area of , and a population of approx. 700,000. Its capit ...
and Bougainville Island) *'' Asellia'' (four species; Africa and southwestern Asia; Miocene fossils from Europe) *'' Aselliscus'' (three species; southeastern Asia and Melanesia) *''
Coelops ''Coelops'' is a genus of bat in the family Hipposideridae. It contains the following species: * East Asian tailless leaf-nosed bat (''Coelops frithii'') * Malayan tailless leaf-nosed bat The Malayan tailless leaf-nosed bat (''Coelops robinson ...
'' (at least two species; southeastern Asia; Miocene fossils from Africa) *''
Hipposideros ''Hipposideros'' is one of the most diverse genera of bats, with more than 70 species. They are collectively called roundleaf bats after the shape of their nasal ornament. It is the type genus of the family Hipposideridae. It is divided into s ...
'' (more than sixty species; Africa, southern Eurasia, and Australasia; oldest fossils from the Eocene of Europe; includes ''Pseudorhinolophus'', sometimes considered a separate genus) *'' Macronycteris'' (five species, Africa and Madagascar) (Note that genus Paracoelops was previously listed for Vietnam is now a synonym of Hipposideros pomona)


Extinct

*'' Archerops'' (
Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
of Australia) *'' Miophyllorhina'' (Miocene of Australia)Archer et al., 2006, p. 7 *'' Palaeophyllophora'' (Eocene to Miocene of Europe) *'' Paraphyllophora'' (Eocene or Oligocene to Miocene of Europe) *''
Riversleigha ''Riversleigha williamsi'' is a species of hipposiderid bat discovered in fossil deposits located the Riversleigh World Heritage Area in the north of Australia. Taxonomy The description of ''Riversleigha williamsi'' was published in 1998 by Suza ...
'' (Miocene of Australia) *'' Vaylatsia'' (Oligocene of Europe)Ziegler, 2000, p. 652; Hand and Kirsch, 2003, table 3; cf. McKenna and Bell, 1997, p. 305 (excluded from Rhinonycterinae) *''
Xenorhinos ''Xenorhinos halli'' is a species of bat that existed in the early Miocene. It was discovered at a fossil deposit of the Riversleigh World Heritage Area in the north of Australia. Taxonomy The description of ''Xenorhinos halli'' was publish ...
'' (Miocene of Australia)


List of species

* Family Hipposideridae — Old World leaf-nosed bats ** Genus '' Anthops'' *** Flower-faced bat, ''Anthops ornatus'' ** Genus '' Asellia'' — trident leaf-nosed bats *** Arabian trident bat, ''Asellia arabica'' *** Somalian trident bat, ''Asellia italosomalica'' *** Patrizi's trident leaf-nosed bat, ''Asellia patrizii'' ***
Trident bat The trident bat or trident leaf-nosed bat (''Asellia tridens'') is a species of bat in the family Hipposideridae. It is widely distributed in the Middle East, South and Central Asia, and North, East, and Central Africa. Its natural habitats a ...
, ''Asellia tridens'' ** Genus '' Aselliscus'' — Tate's trident-nosed bats ***
Stoliczka's trident bat Stoliczka's trident bat (''Aselliscus stoliczkanus'') is a species of bat in the family Hipposideridae. It is found in China, Laos, Malaysia, Myanmar, Thailand, and Vietnam. In northern Thailand Northern Thailand, or more specifically Lanna, ...
, ''Aselliscus stoliczkanus'' *** Temminck's trident bat, ''Aselliscus tricuspidatus'' ** Genus ''
Coelops ''Coelops'' is a genus of bat in the family Hipposideridae. It contains the following species: * East Asian tailless leaf-nosed bat (''Coelops frithii'') * Malayan tailless leaf-nosed bat The Malayan tailless leaf-nosed bat (''Coelops robinson ...
'' — tailless leaf-nosed bats ***
East Asian tailless leaf-nosed bat The East Asian tailless leaf-nosed bat or tail-less leaf-nosed bat (''Coelops frithii'') is a species of bat in the family Hipposideridae. It is found in Bangladesh, China, India, Indonesia, Laos, Malaysia, Myanmar, Taiwan, and Vietnam ...
, ''Coelops frithii'' *** Philippine tailless leaf-nosed bat, ''Coelops hirsutus'' ***
Malayan tailless leaf-nosed bat The Malayan tailless leaf-nosed bat (''Coelops robinsoni'') is a species of bat in the family Hipposideridae. It is a very small bat which has long and soft fur. The fur coloration is brown to blackish on the dorsal surface and ashy on the ventra ...
, ''Coelops robinsoni'' ** Genus ''
Hipposideros ''Hipposideros'' is one of the most diverse genera of bats, with more than 70 species. They are collectively called roundleaf bats after the shape of their nasal ornament. It is the type genus of the family Hipposideridae. It is divided into s ...
'' — roundleaf bats *** Aba roundleaf bat, ''Hipposideros abae'' *** Great roundleaf bat, ''Hipposideros armiger'' *** Dusky roundleaf bat, ''Hipposideros ater'' *** Benito roundleaf bat, ''Hipposideros beatus'' *** Boeadi’s roundleaf bat, ''Hipposideros boeadii'' *** Bicolored roundleaf bat, ''Hipposideros bicolor'' *** Short-headed roundleaf bat, ''Hipposideros breviceps'' *** Sundevall's roundleaf bat, ''Hipposideros caffer'' *** Spurred roundleaf bat, ''Hipposideros calcaratus'' *** Greater roundleaf bat, ''Hipposideros camerunensis'' ***
Fawn leaf-nosed bat The fawn leaf-nosed bat (''Hipposideros cervinus'') is a species of bat in the family Hipposideridae found in Australia, Indonesia, Malaysia, the Philippines and Vanuatu. Taxonomy The species was first described by John Gould in his Mammals of ...
or fawn roundleaf bat, ''Hipposideros cervinus'' *** Ashy roundleaf bat, ''Hipposideros cineraceus'' *** Large Mindanao roundleaf bat, ''Hipposideros coronatus'' *** Telefomin roundleaf bat, ''Hipposideros corynophyllus'' *** Cox's roundleaf bat, ''Hipposideros coxi'' *** Timor roundleaf bat, ''Hipposideros crumeniferus'' *** Short-tailed roundleaf bat, ''Hipposideros curtus'' *** Cyclops roundleaf bat, ''Hipposideros cyclops'' *** Makira roundleaf bat, ''Hipposideros demissus'' *** Diadem roundleaf bat, ''Hipposideros diadema'' *** Fierce roundleaf bat, ''Hipposideros dinops'' ***
Borneo roundleaf bat The Borneo roundleaf bat or Bornean leaf-nosed bat (''Hipposideros doriae'') is a species of bat in the family Hipposideridae. It is found in Borneo, Sumatra and Peninsular Malaysia. ''Hipposideros sabanus'' is a synonym of this species. Taxon ...
, ''Hipposideros doriae'' *** Khajuria's leaf-nosed bat, ''Hipposideros durgadasi'' *** Dayak roundleaf bat, ''Hipposideros dyacorum'' *** Hill's roundleaf bat, ''Hipposideros edwardshilli'' ***'' Hipposideros fasensis'' ***
Sooty roundleaf bat The sooty roundleaf bat (''Hipposideros fuliginosus'') is a species of bat in the family Hipposideridae. It is found in Cameroon, Democratic Republic of the Congo, Ivory Coast, Gabon, Ghana, Guinea, Liberia, Nigeria, Sierra Leone, and Uganda. I ...
, ''Hipposideros fuliginosus'' *** Fulvus roundleaf bat, ''Hipposideros fulvus'' ***
Cantor's roundleaf bat Cantor's roundleaf bat (''Hipposideros galeritus'') is a species of bat in the family Hipposideridae. It is found in Bangladesh, Brunei, Cambodia, India, Indonesia, Laos, Malaysia, Sri Lanka, Thailand and Vietnam. It is named after Theodore Ed ...
, ''Hipposideros galeritus'' *** Grand roundleaf bat, ''Hipposideros grandis'' ***
Thailand roundleaf bat The Thailand roundleaf bat (''Hipposideros halophyllus'') is a species of bat in the family Hipposideridae. It is endemic to Thailand. Distribution ''Hipposideros halophyllus'' is found in: * Khao Samor Khon, Lopburi Province (1,000-1,400 indi ...
, ''Hipposideros halophyllus'' *** Kolar leaf-nosed bat, ''Hipposideros hypophyllus'' *** Crested roundleaf bat, ''Hipposideros inexpectatus'' *** Arnhem leaf-nosed bat, ''Hipposideros inornatus'' *** Jones's roundleaf bat, ''Hipposideros jonesi'' *** Phou Khao Khouay leaf-nosed bat, ''Hipposideros khaokhouayensis'' *** Khasian leaf-nosed bat, ''Hipposideros khasiana'' ***
Lamotte's roundleaf bat Lamotte's roundleaf bat (''Hipposideros lamottei'') is a species of bat found only at Mount Nimba on the border of Côte d'Ivoire, Guinea and Liberia. It is critically endangered. Taxonomy It was described as a new species in 1984 by French ...
, ''Hipposideros lamottei'' *** Indian roundleaf bat, ''Hipposideros lankadiva'' *** Intermediate roundleaf bat, ''Hipposideros larvatus'' *** Large Asian roundleaf bat, ''Hipposideros lekaguli'' *** Shield-faced roundleaf bat, ''Hipposideros lylei'' *** Big-eared roundleaf bat, ''Hipposideros macrobullatus'' ***
Maduran leaf-nosed bat The Maduran leaf-nosed bat (''Hipposideros madurae'') is a species of bat in the family Hipposideridae endemic to Indonesia. This species is known from the eastern half of Java and on Madura Island. It is found below . It is an IUCN Red List Near ...
, ''Hipposideros madurae'' *** Maggie Taylor's roundleaf bat, ''Hipposideros maggietaylorae'' *** Aellen's roundleaf bat, ''Hipposideros marisae'' *** Ethiopian large-eared roundleaf bat, ''Hipposideros megalotis'' *** Fly River roundleaf bat, ''Hipposideros muscinus'' *** Malayan roundleaf bat, ''Hipposideros nequam'' *** Philippine forest roundleaf bat, ''Hipposideros obscurus'' *** Orbiculus leaf-nosed bat, ''Hipposideros orbiculus'' *** Biak roundleaf bat, ''Hipposideros papua'' ***'' Hipposideros parnabyi'' *** Peleng leaf-nosed bat, ''Hipposideros pelingensis'' *** Pomona roundleaf bat, ''Hipposideros pomona'' *** Pratt's roundleaf bat, ''Hipposideros pratti'' *** Philippine pygmy roundleaf bat, ''Hipposideros pygmaeus'' *** Ridley's leaf-nosed bat, ''Hipposideros ridleyi'' *** Laotian leaf-nosed bat, ''Hipposideros rotalis'' *** Noack's roundleaf bat, ''Hipposideros ruber'' ***'' Hipposideros scutinares'' *** Semon's roundleaf bat, ''Hipposideros semoni'' *** Sorensen's leaf-nosed bat, ''Hipposideros sorenseni'' *** Schneider's leaf-nosed bat, ''Hipposideros speoris'' *** Northern leaf-nosed bat or narrow-eared roundleaf bat, ''Hipposideros stenotis'' *** Sumba roundleaf bat or Sumban leaf-nosed bat, ''Hipposideros sumbae'' ***'' Hipposideros tephrus'' *** Lesser great leaf-nosed bat, ''Hipposideros turpis'' *** Wollaston's roundleaf bat, ''Hipposideros wollastoni'' ** Genus '' Macronycteris'' *** Commerson's leaf-nosed bat or Commerson's roundleaf bat, ''Macronycteris commersoni'' ***'' Macronycteris cryptovalorona'' *** Giant roundleaf bat, ''Macronycteris gigas'' *** Saõ Tomé leaf-nosed bat, ''Macronycteris thomensis'' *** Striped leaf-nosed bat, ''Macronycteris vittatus''


Notes


References

;Bibliography *Archer, M., Arena, D.A., Bassarova, M., Beck, R.M.D., Black, K., Boles, W.E., Brewer, P., Cooke, B.N., Crosby, K., Gillespie, A., Godthelp, H., Hand, S.J., Kear, B.P., Louys, J., Morrell, A., Muirhead, J., Roberts, K.K., Scanlon, J.D., Travouillon, K.J. and Wroe, S. 2006. Current status of species-level representation in faunas from selected fossil localities in the Riversleigh World Heritage Area, northwestern Queensland. Alcheringa Special Issue 1:1-17. *Benda, P. and Vallo, P. 2009
Taxonomic revision of the genus ''Triaenops'' (Chiroptera: Hipposideridae) with description of a new species from southern Arabia and definitions of a new genus and tribe
Folia Zoologica 58(Monograph 1):1–45. *Hand, S.J. and Archer, M. 2005. A new hipposiderid genus (Microchiroptera) from an early Miocene bat community in Australia. Palaeontology 48(2):371–383. *Hand, S.J. and Kirsch, J.A.W. 2003. ''Archerops'', a new annectent hipposiderid genus (Mammalia: Microchiroptera) from the Australian Miocene. Journal of Paleontology 77(6):1139–1151. *Hutcheon, J.M. and Kirsch, J.A.W. 2006. A moveable face: deconstructing the Microchiroptera and a new classification of extant bats. Acta Chiropterologica 8(1):1–10. *McKenna, M.C. and Bell, S.K. 1997. Classification of Mammals: Above the species level. New York: Columbia University Press, 631 pp. *Simmons, N.B. 2005. Order Chiroptera. Pp. 312–529 in Wilson, D.E. and Reeder, D.M. (eds.)
Mammal Species of the World: a taxonomic and geographic reference
3rd ed. Baltimore: The Johns Hopkins University Press, 2 vols., 2142 pp. *Ziegler, R. 2000. The bats (Chiroptera, Mammalia) from the Late Oligocene fissure fillings Herrlingen 8 and Herrlingen 9 near Ulm (Baden-Württemberg).
Senckenbergiana Lethaea This is a list of academic journals, monographic series, and other serials published by E. Schweizerbart. A B C D E F G H I J K L M N P Q R S T U V W Z {{columns-list, colwidth=30em, *''Zeitschrift ...
80(2):647–683. {{Taxonbar, from=Q604995 Bat families Taxa named by Richard Lydekker