The heart and dart (''Agrotis exclamationis'') is a

moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...

of the family

Noctuidae The Noctuidae, commonly known as owlet moths, cutworms or armyworms, are a family of moths. They are considered the most controversial family in the superfamily Noctuoidea because many of the clades are constantly changing, along with the other ...

. The species was first described by

Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, ...

in his 1758 10th edition of ''Systema Naturae''. A familiar moth to many, it is considered one of the most common of the European region. It occurs throughout the

Palearctic realm The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa. The realm consists of several bioregions: the Euro-Si ...

from Ireland to Japan. This is a quite variable species with forewings ranging from pale to dark brown but always recognizable by the distinctively shaped dark

stigmata Stigmata ( grc, στίγματα, plural of , 'mark, spot, brand'), in Roman Catholicism, are bodily wounds, scars and pain which appear in locations corresponding to the crucifixion wounds of Jesus Christ: the hands, wrists, and feet. Sti ...

which give it its common name. The

wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingspan o ...

is 35–44 mm. The hindwings are whitish (compared with other common ''Agrotis'' species, the hindwings of this species are usually paler than in heart and club but darker than in turnip moth). This species usually has a dark area at the front of the

thorax The thorax or chest is a part of the anatomy of humans, mammals, and other tetrapod animals located between the neck and the abdomen. In insects, crustaceans, and the extinct trilobites, the thorax is one of the three main divisions of the c ...

, visible as a horizontal bar when viewing the moth head on. The differences are not consistent however; they are highly variable in both colour and markings, and identification of atypical or worn examples may prove impossible without examination of genitalia. See Townsend et al. This moth flies at night from May to July and is attracted to light, sometimes in large numbers. It also frequently visits

nectar Nectar is a sugar-rich liquid produced by plants in glands called nectaries or nectarines, either within the flowers with which it attracts pollinating animals, or by extrafloral nectaries, which provide a nutrient source to animal mutualist ...

-rich flowers such as '' Buddleia'', ragwort and red valerian. The

larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. ...

The caterpillars are coloured grey-brown, paler ventrally and have a pale dorsal line and clearly recognizable black point warts. The pupa is bright red. On the cremaster sit two short curved thorns.Larvae feed on a variety of plants, both wild and cultivated (see list below). This is one of the notorious cutworms and often severs or fatally damages plants at the base. The species overwinters as a full-grown larva in a chamber in the soil before

pupa A pupa ( la, pupa, "doll"; plural: ''pupae'') is the life stage of some insects undergoing transformation between immature and mature stages. Insects that go through a pupal stage are holometabolous: they go through four distinct stages in thei ...

ting in the spring. #''The flight season refers to the

British Isles The British Isles are a group of islands in the North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner and Outer Hebrides, the Northern Isl ...

. This may vary in other parts of the range.''

Recorded host plants

*''

Anemone ''Anemone'' () is a genus of flowering plants in the buttercup family Ranunculaceae. Plants of the genus are commonly called windflowers. They are native to the temperate and subtropical regions of all continents except Australia, New Zealand an ...

'' *'' Artemisia'' *''Beta'' – beet *''

Brassica rapa ''Brassica rapa'' is a plant species growing in various widely cultivated forms including the turnip (a root vegetable); napa cabbage, bomdong, bok choy, and rapini. ''Brassica rapa'' subsp. ''oleifera'' is an oilseed which has many common n ...

'' – turnip *''Fragaria'' –

strawberry The garden strawberry (or simply strawberry; ''Fragaria × ananassa'') is a widely grown hybrid species of the genus '' Fragaria'', collectively known as the strawberries, which are cultivated worldwide for their fruit. The fruit is widely ap ...

*''Lactuca'' –

lettuce Lettuce (''Lactuca sativa'') is an annual plant of the family Asteraceae. It is most often grown as a leaf vegetable, but sometimes for its stem and seeds. Lettuce is most often used for salads, although it is also seen in other kinds of food, ...

*''

Plantago ''Plantago'' is a genus of about 200 species of flowering plants in the family Plantaginaceae, commonly called plantains or fleaworts. The common name plantain is shared with the unrelated cooking plantain. Most are herbaceous plants, though ...

'' - plantain *''

Polygonum ''Polygonum'' is a genus of about 130 species of flowering plant in the buckwheat and knotweed family Polygonaceae. Common names include knotweed and knotgrass (though the common names may refer more broadly to plants from Polygonaceae). In ...

'' *''Quercus'' – oak *''

Rubus ''Rubus'' is a large and diverse genus of flowering plants in the rose family, Rosaceae, subfamily Rosoideae, with over 1,350 species. Raspberries, blackberries, and dewberries are common, widely distributed members of the genus. Most of the ...

'' – bramble *''

Rumex The docks and sorrels, genus ''Rumex'', are a genus of about 200 species of annual, biennial, and perennial herbs in the buckwheat family, Polygonaceae. Members of this genus are very common perennial herbs with a native almost worldwide distri ...

'' – docks, sorrels *''Solanum'' –

potato The potato is a starchy food, a tuber of the plant ''Solanum tuberosum'' and is a root vegetable native to the Americas. The plant is a perennial in the nightshade family Solanaceae. Wild potato species can be found from the southern Uni ...

*''Spinacia'' –

spinach Spinach (''Spinacia oleracea'') is a leafy green flowering plant native to central and western Asia. It is of the order Caryophyllales, family Amaranthaceae, subfamily Chenopodioideae. Its leaves are a common edible vegetable consumed either f ...

*''

Stellaria ''Stellaria'' is a genus of about 190 species of flowering plants in the family Caryophyllaceae, with a cosmopolitan distribution. Common names include starwort, stitchwort and chickweed. Description ''Stellaria'' species are relatively small ...

'' *''Zea'' –

maize Maize ( ; ''Zea mays'' subsp. ''mays'', from es, maíz after tnq, mahiz), also known as corn (North American English, North American and Australian English), is a cereal grain first domesticated by indigenous peoples of Mexico, indigenous ...

Subspecies

* ''A. e. corsica'' -

Corsica Corsica ( , Upper , Southern ; it, Corsica; ; french: Corse ; lij, Còrsega; sc, Còssiga) is an island in the Mediterranean Sea and one of the 18 regions of France. It is the fourth-largest island in the Mediterranean and lies southeast of ...

* ''A. e. exclamationis'' - Europe * ''A. e. informis'' Leech,

889 __NOTOC__ Year 889 ( DCCCLXXXIX) was a common year starting on Wednesday (link will display the full calendar) of the Julian calendar. Events By place Europe * Guy III, duke of Spoleto, defeats the Lombard king Berengar I at the ...

/small> - Japan Agrotis exclamationis-s.jpg, Profile Agrotis exclamationis-f.jpg, Front mark Agrotis exclamationis-o.jpg, Color variation

References

Chinery, Michael Michael Chinery (born 1938, in London) is an English naturalist. He studied in Cambridge where he graduated in natural sciences and anthropology. He edits '' Cecidology'', the journal of the British Plant Gall Society The British Plant Gal ...

(1986, reprinted 1991). ''Collins Guide to the Insects of Britain and Western Europe''. * Skinner, Bernard (1984). ''

The Colour Identification Guide to Moths of the British Isles ''The Colour Identification Guide to Moths of the British Isles (Macrolepidoptera)'' by Bernard Skinner is a single volume identification guide to the macromoths of Britain and Ireland published by Viking Books, often referred by moth recorder ...

''.

External links

* *
''Lepiforum e.V.''
{{Taxonbar , from=Q771090 Agrotis Moths of Asia Moths of Europe Moths of Japan Moths of the Middle East Taxa named by Carl Linnaeus