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Grulla or grullo, also called blue dun, gray dun or mouse dun, is a
color Color (American English) or colour (British English) is the visual perceptual property deriving from the spectrum of light interacting with the photoreceptor cells of the eyes. Color categories and physical specifications of color are associ ...
of
horse The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 million yea ...
s in the dun family, characterized by tan-gray or mouse-colored hairs on the body, often with shoulder and dorsal stripes and black barring on the lower legs. In this coloration, each individual hair is mouse-colored, unlike a roan, which is composed of a mixture of dark and light hairs. The several shades of grulla are informally referred to with a variety of terms, including black dun, blue dun, slate grulla, silver grulla or light grulla, silver dun, or lobo dun. Silver grulla may also refer to a grullo horse with
silver dapple The silver or silver dapple (''Z'') gene is a dilution gene that affects the black base coat color and is associated with Multiple Congenital Ocular Abnormalities. It will typically dilute a black mane and tail to a silvery gray or flaxen color ...
, regardless of shade. In the Icelandic horse, the grulla color is called gray dun, in the Highland pony it is called mouse dun, and in the Norwegian
Fjord horse The Fjord Horse or Norwegian Fjord Horse ( no, fjordhest) is a relatively small but very strong horse breed from the mountainous regions of western Norway. It is an agile breed of light draught horse build. All Fjord horses are dun in colour, ...
, ''grå'' or ''gråblakk'' (literally, "gray dun"). The word "grulla" in Spanish means crane. Because of the origin of the name, some people will refer to a mare as a grulla and a stallion or gelding as a grullo, pronounced and . (The original Spanish noun is pronounced in American Spanish and in
Peninsular Spanish Peninsular Spanish ( es, español peninsular) (also known as the Spanish of Spain ( es, español de España, links=no), European Spanish ( es, español europeo, links=no), Iberian Spanish ( es, español ibérico, links=no) or Spanish Spanish ( es ...
.) In terms of equine coat color genetics, all of these shades are based on the
dun gene The dun gene is a dilution gene that affects both red and black pigments in the coat color of a horse. The dun gene lightens most of the body while leaving the mane, tail, legs, and primitive markings the shade of the undiluted base coat color. ...
acting as a
dilution gene A dilution gene is any one of a number of genes that act to create a lighter coat color in living creatures. There are many examples of such genes: General Diluted coat colors have melanocytes, but vary from darker colors due to the concentratio ...
over the
black Black is a color which results from the absence or complete absorption of visible light. It is an achromatic color, without hue, like white and grey. It is often used symbolically or figuratively to represent darkness. Black and white ha ...
gene. Because the grulla color is not due to the
gray Grey (more common in British English) or gray (more common in American English) is an intermediate color between black and white. It is a neutral or achromatic color, meaning literally that it is "without color", because it can be compose ...
gene, a grulla horse remains the same basic color from birth, though some minor shade variation may occur from summer to winter coats. If a grulla also carries the gray gene, it will be born a mouse tan-gray shade, usually with bold primitive markings, but then lighten and eventually develop a white hair coat with age. Because black is less common in general than bay or chestnut, grulla is likewise less common than red duns or bay (classic or zebra) duns. For example, only 0.7% of quarter horses registered each year with the
AQHA The American Quarter Horse Association (AQHA), based in Amarillo, Texas, is an international organization dedicated to the preservation, improvement and record-keeping of the American Quarter Horse. The association sanctions many competitive eve ...
are grulla. The most obvious ways to tell whether a horse is grulla are not only the gray or tan-gray body color, but also its
primitive markings Primitive markings are a group of hair coat markings and qualities seen in several equine species, including horses, donkeys, and asses. In horses, they are associated with primitive breeds, though not limited to such breeds. The markings are ...
, which include some or all of the following: dark face, cobwebbing around the eyes and forehead, dark mottling on the body, leg barring (sometimes called tiger striping), dark ear tips and edging, dark ear barring, dark shadowing of the neck, dark dorsal and transverse striping, and light
guard hairs Fur is a thick growth of hair that covers the skin of mammals. It consists of a combination of oily guard hair on top and thick underfur beneath. The guard hair keeps moisture from reaching the skin; the underfur acts as an insulating blanket t ...
bordering a dark mane and tail.


Primitive roots

The
tarpan The term tarpan (''Equus ferus ferus'') refers to free-ranging horses of the Russian steppe from the 18th to the 20th century. It is generally unknown whether those horses represented genuine wild horses, feral domestic horses or hybrids. The las ...
(''Equus ferus ferus'') was a relative of the domestic horse that became
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
in the nineteenth century, and which appears to have had grulla coloration. The tarpan has been considered a true
wild horse The wild horse (''Equus ferus'') is a species of the genus ''Equus'', which includes as subspecies the modern domesticated horse (''Equus ferus caballus'') as well as the endangered Przewalski's horse (''Equus ferus przewalskii''). The Europea ...
, an undomesticated relative or ancestor of the domestic horse. However, some authorities in the early twentieth century held the opinion that most equines called tarpans were actually domestic or feral horses, not a separate species. Several breeds with the grulla color have been developed in efforts to recreate (" breed back") the tarpan. These breeds include the
Heck horse The Heck horse is a horse breed that is claimed to resemble the tarpan (''Equus ferus ferus''), an extinct equine. The breed was created by the German zoologist brothers Heinz Heck and Lutz Heck in an attempt to breed back the tarpan. Al ...
and
Konik The Konik or Polish Konik, pl, konik polski, is a Polish breed of pony. There are semi- feral populations in some regions. They are usually mouse dun or striped dun in color. The Bilgoray, pl, konik biłgorajski, of south-eastern Poland is ...
. One of the first experiments in this regard was published in 1906 by
James Cossar Ewart James Cossar Ewart FRS FRSE (26 November 1851 – 31 December 1933) was a Scottish zoologist. He performed breeding experiments with horses and zebras which disproved earlier theories of heredity. Life Ewart was born in Penicuik, Midloth ...
, who obtained a "tarpan-like" horse by crossing a Shetland mare and a black Welsh pony.


References


Sources and external links




Grulla Color Genetics and Photos


* http://www.equusite.com/articles/basics/colors/colorsGrullo.shtml
Dun Zygosity test
''from'' Veterinary Genetics Laboratory, School of Veterinary Medicine, University of California, Davis. Web site accessed January 13, 2008.
"Introduction to Coat Color Genetics"
''from'' Veterinary Genetics Laboratory, School of Veterinary Medicine, University of California, Davis. Web Site accessed January 12, 2008 {{Equine coat colors https://www.spanishdict.com/translate/grullo Horse coat colors