Growth cycle (vine)
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The annual growth cycle of grapevines is the process that takes place in the vineyard each year, beginning with
bud In botany, a bud is an undeveloped or embryonic shoot and normally occurs in the axil of a leaf or at the tip of a stem. Once formed, a bud may remain for some time in a dormant condition, or it may form a shoot immediately. Buds may be spec ...
break in the spring and culminating in leaf fall in autumn followed by winter dormancy. From a
winemaking Winemaking or vinification is the production of wine, starting with the selection of the fruit, its fermentation into alcohol, and the bottling of the finished liquid. The history of wine-making stretches over millennia. The science of wine and ...
perspective, each step in the process plays a vital role in the development of
grape A grape is a fruit, botanically a berry, of the deciduous woody vines of the flowering plant genus '' Vitis''. Grapes are a non- climacteric type of fruit, generally occurring in clusters. The cultivation of grapes began perhaps 8,000 years a ...
s with ideal characteristics for making
wine Wine is an alcoholic drink typically made from fermented grapes. Yeast consumes the sugar in the grapes and converts it to ethanol and carbon dioxide, releasing heat in the process. Different varieties of grapes and strains of yeasts are m ...
.
Viticulturalist Viticulture (from the Latin word for ''vine'') or winegrowing (wine growing) is the cultivation and harvesting of grapes. It is a branch of the science of horticulture. While the native territory of ''Vitis vinifera'', the common grape vine, ran ...
s and vineyard managers monitor the effect of climate,
vine disease This is a list of diseases of grapes (''Vitis'' spp.). Bacterial diseases Fungal diseases Miscellaneous diseases and disorders Nematodes, parasitic Phytoplasma, virus and viruslike diseases See also *'' Ampeloglypter ater'' *'' Am ...
and pests in facilitating or impeding the vine's progression from bud break,
flowering A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechanism ...
,
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...
set,
veraison In viticulture, veraison (french: véraison, ) is the onset of the ripening of the grapes. The official definition of ''veraison'' is "change of color of the grape berries". Veraison represents the transition from berry growth to berry ripening, ...
,
harvesting Harvesting is the process of gathering a ripe crop from the fields. Reaping is the cutting of grain or pulse for harvest, typically using a scythe, sickle, or reaper. On smaller farms with minimal mechanization, harvesting is the most labor- ...
, leaf fall and dormancy-reacting if need be with the use of viticultural practices like
canopy management In viticulture, the canopy of a grapevine includes the parts of the vine visible aboveground - the trunk, cordon, stems, leaves, flowers, and fruit. The canopy plays a key role in light energy capture via photosynthesis, water use as regulated ...
,
irrigation Irrigation (also referred to as watering) is the practice of applying controlled amounts of water to land to help grow crops, landscape plants, and lawns. Irrigation has been a key aspect of agriculture for over 5,000 years and has been devel ...
,
vine training The use of vine training systems in viticulture is aimed primarily to assist in canopy management with finding the balance in enough foliage to facilitate photosynthesis without excessive shading that could impede grape ripening or promote grape ...
and the use of
agrochemicals An agrochemical or agrichemical, a contraction of ''agricultural chemical'', is a chemical product used in industrial agriculture. Agrichemical refers to biocides (pesticides including insecticides, herbicides, fungicides and nematicides) and sy ...
. The stages of the annual growth cycle usually become observable within the first year of a vine's life. The amount of time spent at each stage of the growth cycle depends on a number of factors-most notably the type of climate (warm or cool) and the characteristics of the
grape variety This list of grape varieties includes cultivated grapes, whether used for wine, or eating as a table grape, fresh or dried (raisin, currant, sultana). For a complete list of all grape species including those unimportant to agriculture, see Viti ...
.J. Robinson (ed) ''"The Oxford Companion to Wine"'' Third Edition pg 741–742 Oxford University Press 2006


Bud break

The grape starts its annual growth cycle in the spring with bud break. In the Northern Hemisphere, this stage begins around March while in the Southern Hemisphere it begins around September when daily temperatures begin to surpass 10 °C (50 °F). If the vine had been pruned during the winter, the start of this cycle is signaled by a "bleeding" of the vine. This bleeding occurs when the soil begins to warm and
osmotic Osmosis (, ) is the spontaneous net movement or diffusion of solvent molecules through a selectively-permeable membrane from a region of high water potential (region of lower solute concentration) to a region of low water potential (region o ...
forces pushes water, containing a low concentration of organic acids, hormones,
mineral In geology and mineralogy, a mineral or mineral species is, broadly speaking, a solid chemical compound with a fairly well-defined chemical composition and a specific crystal structure that occurs naturally in pure form.John P. Rafferty, ed. (2 ...
s and sugars, up from the
root system In mathematics, a root system is a configuration of vectors in a Euclidean space satisfying certain geometrical properties. The concept is fundamental in the theory of Lie groups and Lie algebras, especially the classification and representatio ...
of the vine and it is expelled from the cuts (or "wounds") left over from pruning the vine. During this period a single vine can "bleed" up to of water.J. Robinson (ed) ''"The Oxford Companion to Wine"'' Third Edition pg 82 Oxford University Press 2006 Tiny buds on the vine start to swell and eventually
shoot In botany, a plant shoot consists of any plant stem together with its appendages, leaves and lateral buds, flowering stems, and flower buds. The new growth from seed germination that grows upward is a shoot where leaves will develop. In the sp ...
s begin to grow from the buds. Buds are the small part of the vine that rest between the vine's stem and the petiole (leaf stem). Inside the buds contain usually three primordial shoots. These buds appear in the summer of previous growth cycle green and covered in scales. During winter dormancy they turn brown until the spring when the vine begins the process of bud break and the first sign of green in the vineyard emerges in the form of tiny shoots.J. Robinson (ed) ''"The Oxford Companion to Wine"'' Third Edition pg 107 Oxford University Press 2006 The energy to facilitate this growth comes from reserves of
carbohydrate In organic chemistry, a carbohydrate () is a biomolecule consisting of carbon (C), hydrogen (H) and oxygen (O) atoms, usually with a hydrogen–oxygen atom ratio of 2:1 (as in water) and thus with the empirical formula (where ''m'' may or m ...
stored in roots and wood of the vine from the last growth cycle. Eventually the shoots sprout tiny leaves that can begin the process of
photosynthesis Photosynthesis is a process used by plants and other organisms to convert light energy into chemical energy that, through cellular respiration, can later be released to fuel the organism's activities. Some of this chemical energy is stored i ...
, producing the energy to accelerate growth. In warm climates, after about 4 weeks the growth of the shoots starts to rapidly accelerate with the shoots growing in length an average of 3 cm (1 in) a day. In
temperate climate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout ...
s, where temperatures can reach above 10 °C (50 °F) in mid-winter, some early budding varieties (such as Chardonnay) can be at risk of premature bud break. This is a potential viticultural hazard in places like the
Margaret River The Margaret River is a river in southwest Western Australia. In a small catchment, it is the eponym of the town and tourist region of Margaret River. The river arises from a catchment of just 40 square kilometres in the Whicher Range. ...
region of Western Australia where warm currents from the Indian Ocean can coax Chardonnay vines to prematurely bud in the mid-winter month of July. After bud break, the young shoots are very vulnerable to frost damage with vineyard managers going to great lengths protect the fragile shoots should temperature dramatically drop below freezing. This can include setting up heaters or wind circulators in the vineyard to keep cold air from settling on the vines.


Flowering

Depending on temperatures, 40–80 days after bud break the process of flowering begins with small flower clusters resembling buttons appearing on the tips of the young shoots. Flowering occurs when average daily temperatures stay between 15–20 °C (59–68 °F) which in the Northern Hemisphere wine regions is generally around May and for the Southern Hemisphere regions around November. A few weeks after the initial clusters appear, the flowers start to grow in size with individual flowers becoming observable. It is during this stage of flowering that the
pollination Pollination is the transfer of pollen from an anther of a plant to the stigma of a plant, later enabling fertilisation and the production of seeds, most often by an animal or by wind. Pollinating agents can be animals such as insects, birds, a ...
and
fertilization Fertilisation or fertilization (see spelling differences), also known as generative fertilisation, syngamy and impregnation, is the fusion of gametes to give rise to a new individual organism or offspring and initiate its development. Proce ...
of the grapevine takes place with the resulting product being a grape berry, containing 1–4 seeds.J. Robinson (ed) ''"The Oxford Companion to Wine"'' Third Edition pg 276 Oxford University Press 2006 Most cultivated ''
Vitis vinifera ''Vitis vinifera'', the common grape vine, is a species of flowering plant, native to the Mediterranean region, Central Europe, and southwestern Asia, from Morocco and Portugal north to southern Germany and east to northern Iran. There are curr ...
'' grape vines are
hermaphroditic In reproductive biology, a hermaphrodite () is an organism that has both kinds of reproductive organs and can produce both gametes associated with male and female sexes. Many taxonomic groups of animals (mostly invertebrates) do not have s ...
, with both male stamens and female
ovaries The ovary is an organ in the female reproductive system that produces an ovum. When released, this travels down the fallopian tube into the uterus, where it may become fertilized by a sperm. There is an ovary () found on each side of the body. T ...
, while many wild grapes are either male, producing pollen but no fruit, or female, producing fruit only if a pollinator is nearby. Hermaphroditic vines are preferred for cultivation because each vine is more likely to self-pollinate and produce fruit. At the beginning of the flowering process the only part that is visible is the fused cap of petals known as the calyptra. Shortly after the calyptra is shed, liberating the pollen from the anthers of the stamen. Wind and insects generally play only a small role in aiding pollination, with the process being mostly self-contained within the vine. But
cross-pollination Pollination is the transfer of pollen from an anther of a plant to the stigma of a plant, later enabling fertilisation and the production of seeds, most often by an animal or by wind. Pollinating agents can be animals such as insects, bird ...
between vines of different varieties is possible: Cabernet Sauvignon is a cross of Cabernet Franc and Sauvignon blanc;
Petite Sirah Durif is a variety of red wine grape primarily grown in Australia, California, France, and Israel. Since the end of the 20th century, wineries located in Washington's Yakima River Valley, Maryland, Arizona, Texas, West Virginia, Chile, Mexico ...
is a cross of Syrah and
Peloursin Peloursin is red French wine grape variety best known for crossing with Syrah to make the red wine grape Durif (known in the United States as Petite Sirah).J. Robinson ''"The Oxford Companion to Wine"'' Third Edition pg 510 Oxford University P ...
. During the process of fertilization, the pollen fertilizes the ovary which produces seeds as the flower begins the transformation into a grape berry, encapsulating the seed. Detrimental weather (cold, wind & rain) can severely affect the flowering process, causing many flowers not to be fertilized and produce a group. It is during this time when the buds that will become next years crops begin to form.


Fruit set

The stage of fruit set follows flowering almost immediately, when the fertilized flower begins to develop a seed and grape berry to protect the seed. In the Northern Hemisphere, this normally takes place in May and in the Southern Hemisphere in November. This stage is very critical for wine production since it determines the potential crop yield. Not every flower on the vine gets fertilized, with the unfertilized flowers eventually falling off the vine. The percentage of fertilized flowers averages around 30 but can get as high as 60 or be much lower. Climate and the health of the vine play an important role with low
humidity Humidity is the concentration of water vapor present in the air. Water vapor, the gaseous state of water, is generally invisible to the human eye. Humidity indicates the likelihood for precipitation, dew, or fog to be present. Humidity dep ...
, high temperatures and water stress having the potential of severely reducing the amount flowers that get fertilized.
Coulure Coulure (pronounced coo-LYUR) is a viticultural hazard that is the result of metabolic reactions to weather conditions that causes a failure of grapes to develop after flowering. In English the word ''shatter'' is sometimes used. Coulure is trigge ...
occurs when there is an imbalance of carbohydrate levels in the vine tissues and some berries fail to set or simply fall off the bunch. Varieties like Grenache and
Malbec Malbec () is a purple grape variety used in making red wine. The grapes tend to have an inky dark color and robust tannins, and are known as one of the six grapes allowed in the blend of red Bordeaux wine. In France, plantations of Malbec are n ...
are prone to this abnormal fruit set.
Millerandage Millerandage (or shot berries, hens and chicks and pumpkins and peas) is a potential viticultural hazard problem in which grape bunches contain berries that differ greatly in size and, most importantly, maturity. Its most common cause is cold, rain ...
occurs when some fertilized flowers do not form seeds but only small berry clusters. Grape berry size depends on the number of seeds so berries with no seeds will be significantly smaller than berries containing seeds. On one cluster there may be berries of various sizes which can create problems during winemaking due to the varying "skin to pulp" ratio among the grapes.J. Robinson (ed) ''"The Oxford Companion to Wine"'' Third Edition pg 291 Oxford University Press 2006 This can be caused by vine disease, such as fanleaf, or by a boron deficiency in the vine.
Gewürztraminer Gewürztraminer () is an aromatic wine grape variety, used in white wines, and performs best in cooler climates. In English, it is sometimes referred to colloquially as Gewürz (; although this is never the case in German, because "Gewürz" me ...
and the Chardonnay
clones Clone or Clones or Cloning or Cloned or The Clone may refer to: Places * Clones, County Fermanagh * Clones, County Monaghan, a town in Ireland Biology * Clone (B-cell), a lymphocyte clone, the massive presence of which may indicate a pathologi ...
IA and Mendoza are both prone to millerandage.J. Robinson (ed) ''"The Oxford Companion to Wine"'' Third Edition pg 443 Oxford University Press 2006


Veraison

Following fruit set, the grape berries are green and hard to the touch. They have very little sugar and are high in organic acids. They begin to grow to about half their final size when they enter the stage of veraison. This stage signals the beginning of the
ripening Ripening is a process in fruits that causes them to become more palatable. In general, fruit becomes sweeter, less green, and softer as it ripens. Even though the acidity of fruit increases as it ripens, the higher acidity level does not make th ...
process and normally takes place around 40–50 days after fruit set. In the Northern Hemisphere this will be around the end of July and into August and between the end of January into February for the Southern Hemisphere. During this stage the colors of the grape take form—red/black or yellow/green depending on the grape varieties. This color changing is due to the chlorophyll in the berry skin being replaced by
anthocyanins Anthocyanins (), also called anthocyans, are water-soluble vacuolar pigments that, depending on their pH, may appear red, purple, blue, or black. In 1835, the German pharmacist Ludwig Clamor Marquart gave the name Anthokyan to a chemical com ...
(red wine grapes) and
carotenoids Carotenoids (), also called tetraterpenoids, are yellow, orange, and red organic pigments that are produced by plants and algae, as well as several bacteria, and fungi. Carotenoids give the characteristic color to pumpkins, carrots, parsnips, co ...
(white wine grapes). In a process known as engustment, the berries start to soften as they build up
sugars Sugar is the generic name for sweet-tasting, soluble carbohydrates, many of which are used in food. Simple sugars, also called monosaccharides, include glucose, fructose, and galactose. Compound sugars, also called disaccharides or double ...
. Within six days of the start of veraison, the berries begin to grow dramatically as they accumulate
glucose Glucose is a simple sugar with the molecular formula . Glucose is overall the most abundant monosaccharide, a subcategory of carbohydrates. Glucose is mainly made by plants and most algae during photosynthesis from water and carbon dioxide, u ...
and fructose and acids begin to fall.J. Robinson (ed) ''"The Oxford Companion to Wine"'' Third Edition pg 730 Oxford University Press 2006 The onset of veraison does not occur uniformly among all berries. Typically the berries and clusters that are most exposed to warmth, on the outer extents of the
canopy Canopy may refer to: Plants * Canopy (biology), aboveground portion of plant community or crop (including forests) * Canopy (grape), aboveground portion of grapes Religion and ceremonies * Baldachin or canopy of state, typically placed over an ...
, undergo veraison first with the berries and clusters closer to the trunk and under the canopy shade undergoing it last. There are some factors in the vineyards that can control the onset of veraison, limited
water stress Water scarcity (closely related to water stress or water crisis) is the lack of fresh water resources to meet the standard water demand. There are two types of water scarcity: physical or economic water scarcity. Physical water scarcity is where ...
and canopy management that creates a high "fruit to leaf" ratio can encourage veraison. This is because the vine is biologically programmed to channel all its energies and resources into the berries, which houses its seedling offspring, to provide them a better chance of survival. Conversely, very vigorous vines with plentiful leaf shading for photosynthesis and water supply will delay the start of veraison due to the vines energies being directed towards continued shoot growth of new buds. For the production of high-quality wine, it is considered ideal to have an earlier veraison. During this period the cane of the vine starts to ripen as well changing from green and springing to brown and hard. The vines begins to divert some of its energy production into its reserves in preparation for its next growth cycle.


After harvest

In the vineyard, the antepenultimate event is the harvest in which the grapes are removed from the vine and transported to the winery to begin the wine making process. In the Northern Hemisphere this is generally between September and October while in the Southern Hemisphere it is generally between February and April. The time of harvest depends on a variety of factors-most notably the subjective determination of ripeness. As the grape ripens on the vines, sugars and pH increase as acids (such as
malic acid Malic acid is an organic compound with the molecular formula . It is a dicarboxylic acid that is made by all living organisms, contributes to the sour taste of fruits, and is used as a food additive. Malic acid has two stereoisomeric forms ( ...
) decrease.
Tannin Tannins (or tannoids) are a class of astringent, polyphenolic biomolecules that bind to and precipitate proteins and various other organic compounds including amino acids and alkaloids. The term ''tannin'' (from Anglo-Norman ''tanner'' ...
s and other phenolics also develop which can affect the flavors and aromas in the resulting wine. The threat of detrimental weather and vine diseases (such as
grey rot ''Botrytis cinerea'' is a necrotrophic fungus that affects many plant species, although its most notable hosts may be wine grapes. In viticulture, it is commonly known as "botrytis bunch rot"; in horticulture, it is usually called "grey mould" o ...
) can also play a role in the time table. The balance of all these factors contributes to when a winemaker or vineyard manager decides that it is time to harvest. Following the harvest, the vines continue the process of photosynthesis, creating carbohydrate reserves to store in the vine's roots and trunks. It will continue doing this until an appropriate level of reserves have been stored. At that point the chlorophyll in the leaves begins to break down and the leaves change color from green to yellow. Following the first frost the leaves begin to fall as the vine starts to enter its winter dormancy period. The following spring, the cycle begins again.


See also

* BBCH-scale (grape)


References


External links


Annual Cycle of the Grapevine

Stages of Grape Berry Development
information from Cooperative Extension {{Viticulture Viticulture