Graphium (butterfly)
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''Graphium'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of mostly tropical swallowtail butterflies commonly known as swordtails, kite swallowtails, or ladies. Native to
Eurasia Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelago ...
, Africa, and
Oceania Oceania (, , ) is a geographical region that includes Australasia, Melanesia, Micronesia, and Polynesia. Spanning the Eastern and Western hemispheres, Oceania is estimated to have a land area of and a population of around 44.5 million ...
, the genus is represented by over 100
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
. Their colouration is as variable as the habitats they frequent; from rainforest to
savanna A savanna or savannah is a mixed woodland- grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground to ...
h. Some possess tails which may be long and swordlike, while others lack any hindwing extensions. ''Graphium'' species are often sighted at mud puddles. The more colourful species are popular with collectors and are commonly seen mounted in frames for sale. Well-known species include the tailed jay (''Graphium agamemnon''), common bluebottle (''G. sarpedon''), and the purple-spotted swallowtail (''G. weiskei''). One species, '' G. idaeoides'', is notable for being a perfect mimic of the danainid '' Idea leuconoe''. Larvae feed variously on
Annonaceae The Annonaceae are a family of flowering plants consisting of trees, shrubs, or rarely lianas commonly known as the custard apple family or soursop family. With 108 accepted genera and about 2400 known species, it is the largest family in the Ma ...
(most commonly),
Magnoliaceae The Magnoliaceae () are a flowering plant family, the magnolia family, in the order Magnoliales. It consists of two genera: ''Magnolia'' and '' Liriodendron'' (tulip trees). Unlike most angiosperms, whose flower parts are in whorls (rings), t ...
(commonly), Lauraceae (commonly),
Rutaceae The Rutaceae is a family, commonly known as the rueRUTACEAE
in BoDD – Botanical Der ...
,
Dioscoreaceae Dioscoreaceae () is a family of monocotyledonous flowering plants, with about 715 known species in nine genera. The best-known member of the family is the yam (some species of ''Dioscorea''). The APG system (1998) and APG II system (2003) both ...
, Bombacaceae,
Piperaceae The Piperaceae (), also known as the pepper family, are a large family of flowering plants. The group contains roughly 3,600 currently accepted species in 5 genera. The vast majority of species can be found within the two main genera: ''Piper'' ...
,
Anacardiaceae The Anacardiaceae, commonly known as the cashew family or sumac family, are a family of flowering plants, including about 83 genera with about 860 known species. Members of the Anacardiaceae bear fruits that are drupes and in some cases produce ...
,
Apocynaceae Apocynaceae (from ''Apocynum'', Greek for "dog-away") is a family of flowering plants that includes trees, shrubs, herbs, stem succulents, and vines, commonly known as the dogbane family, because some taxa were used as dog poison Members of ...
, Malpighiaceae,
Hernandiaceae The Hernandiaceae are a family of flowering plants (angiosperms) in the order Laurales. Consisting of five genera with about 58 known species, they are distributed over the world's tropical areas, some of them widely distributed in coastal areas ...
, Guttiferae,
Monimiaceae The Monimiaceae is a family of flowering plants in the magnoliid order Laurales.Peter F. Stevens (2001 onwards). "Monimiaceae" At: Angiosperm Phylogeny Website. At: Botanical Databases At: Missouri Botanical Garden Website. (see ''External links' ...
, Pandanaceae, Winteraceae, and
Euphorbiaceae Euphorbiaceae, the spurge family, is a large family of flowering plants. In English, they are also commonly called euphorbias, which is also the name of a genus in the family. Most spurges, such as '' Euphorbia paralias'', are herbs, but some, ...
.


Species

Listed alphabetically: subgenus: ''Idaides'' :
species group In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...
: ''codrus'' ::* '' Graphium anthedon'' (C. Felder & R. Felder, 1864) – Wallacea bluebottle ::* ''
Graphium cloanthus ''Graphium cloanthus'', the glassy bluebottle, is a common, non-threatened tropical butterfly of the family Papilionidae. Description Upperside, forewing: costal margin up to a line through the anterior half of the cell to the apex of the wing a ...
'' (Westwood, 1841) – glassy bluebottle ::* '' Graphium codrus'' (Cramer, 777 – eastern olive triangle ::* '' Graphium empedocles'' (Fabricius, 1787), originally ''Papilio empedocles'' Fabricius, 1787; ''Mantissa Insectorum'' 2: 10, no. 94 = homonym of ''Papilio empedocles'' Cramer, 1779; replaced by ''empedovana'' Corbet, 1941. ::* '' Graphium empedovana'' (Corbet, 1941) ::* '' Graphium gelon'' (Boisduval, 1859) ::* '' Graphium macleayanus'' (Leach, 1814) – Macleay's swallowtail ::* '' Graphium mendana'' ( Godman & Salvin, 1888) ::* ''
Graphium monticolus ''Graphium monticolus'', the Sulawesi blue triangle, is a butterfly of the family Papilionidae that is found in Sulawesi in Indonesia. Taxonomy Originally described as a subspecies of '' Papilio'' (''Papilio sarpedon monticolus'' Fruhstorfer, 18 ...
'' (Fruhstorfer, 1896) – Sulawesi blue triangle ::* ?'' Graphium protensor'' (Gistel,1857) ''Papilio protensor'' Gistel, J.,1857 Achthundert und zwanzig neue oder unbeschriebene wirbellose Theire charakterisirt von Doctor Johannes Gistel ''Vacuna'' 2(2):513-606. synonym of '' Graphium sarpedon'' ::* '' Graphium sarpedon'' (Linnaeus, 1758) – common bluebottle or blue triangle ::* '' Graphium stresemanni'' ( Rothschild, 1916) ::* '' Graphium weiskei'' (Ribbe, 1900) – purple spotted swallowtail :species group: ''eurypylus'' ::* '' Graphium agamemnon'' (Linnaeus, 1758) – tailed jay or green-spotted triangle ::* '' Graphium arycles'' (Boisduval, 1836) – spotted jay ::* '' Graphium bathycles'' (Zinken, 1831) ::* '' Graphium chironides'' (Honrath, 1884) – veined jay ::* '' Graphium doson'' (C. Felder & R. Felder, 1864) – common jay ::* '' Graphium eurypylus'' (Linnaeus, 1758) – great jay or pale green triangle ::* '' Graphium evemon'' (Boisduval, 1836) – lesser jay ::* '' Graphium leechi'' (Rothschild, 1895) ::* '' Graphium macfarlanei'' (Butler, 1877) – green triangle butterfly or green triangle ::* '' Graphium meeki'' (Rothschild & Jordan, 1901) – Meek's graphium ::* '' Graphium meyeri'' (Hopffer, 1874) ::* '' Graphium procles'' (Grose-Smith, 1887) :species group: ''wallacei'' ::* ''
Graphium browni ''Graphium browni'' is a butterfly found in Oceania - New Britain, Duke of York Islands, New Hanover Island and St Matthias Islands - that belongs to the swallowtail family. Description The discal band of the hindwing narrow and linear before ...
'' (Godman & Salvin, 1879) ::* ''
Graphium hicetaon ''Graphium hicetaon'' is a butterfly found in the Solomon Islands - Bougainville Island, Choiseul Island, Shortland Island, Florida Island, Guadalcanal, New Georgia Group and Ugi Island - that belongs to the swallowtail family. Taxonomy ''Gra ...
'' (Mathew, 1886) ::* ''
Graphium sandawanum ''Graphium sandawanum'', the Apo swallowtail, is a species of butterfly in the family Papilionidae. It is endemic to the Philippines. Taxonomy ''Graphium hicetaon'' belongs to the ''wallacei'' species group. This clade has four members: *'' Gra ...
'' Yamamoto, 1977 – Apo swallowtail ::* ''
Graphium wallacei ''Graphium wallacei'' is a butterfly found in New Guinea and the Moluccas that belongs to the swallowtail family. Subspecies *''G. w. wallacei'' (Aru Islands, western Irian, New Guinea) *''G. w. rubrosignatus'' (Rothschild, 1895) (Halmahera, B ...
'' (Hewitson, 1858) subgenus: ''Arisbe'' Hübner, 819/small> :species group: ''antheus'' ::* '' Graphium antheus'' (Cramer, 779 – large striped swordtail ::* '' Graphium evombar'' (Boisduval, 1836) :species group: ''policenes'' ::* '' Graphium biokoensis'' Gauthier, 1984 – Gauthier's striped swordtail ::* '' Graphium colonna'' (Ward, 1873) – black swordtail or mamba swordtail ::* '' Graphium gudenusi'' (Rebel, 1911) ::* '' Graphium illyris'' (Hewitson, 1872) - cream-banded swordtail ::* '' Graphium junodi'' (Trimen, 1893) – Junod's swordtail ::* '' Graphium kirbyi'' (Hewitson, 1872) – Kirby's swordtail ::* '' Graphium liponesco'' (Suffert, 1904) ::* '' Graphium policenes'' (Cramer, 775 – small striped swordtail or common swordtail ::* '' Graphium policenoides'' (Holland, 1892) ::* '' Graphium polistratus'' (Grose-Smith, 1889) – dancing swordtail ::* '' Graphium porthaon'' (Hewitson, 1865) – coastal swordtail, cream-striped swordtail, or dark swordtail :species group: ''angolanus'' ::* '' Graphium angolanus'' (Goeze, 1779) – Angola white lady (swordtail) ::* '' Graphium endochus'' (Boisduval, 1836) ::* '' Graphium morania'' (Angas, 1849) – white lady (swallowtail) or small white-lady swordtail ::* '' Graphium ridleyanus'' (White, 1843) - acraea swordtail ::* '' Graphium schaffgotschi'' (Niepelt, 1927) – Schaffgotsch's swordtail ::* '' Graphium taboranus'' (Oberthür, 1886) :species group: ''leonidas'' ::* '' Graphium cyrnus'' (Boisduval, 1836) ::* '' Graphium leonidas'' (Fabricius, 1793) – veined swordtail, veined swallowtail, or common graphium ::* '' Graphium levassori'' Oberthür, 1890 :species group: ''tynderaeus'' ::* '' Graphium tynderaeus'' (Fabricius, 1793) – electric green swordtail or green-spotted swallowtail ::* '' Graphium latreillianus'' (Godart, 1819) – coppery swordtail :species group: ''philonoe'' ::* '' Graphium philonoe'' (Ward, 1873) – eastern white-lady swordtail, eastern graphium, white-dappled swallowtail :species group: ''adamastor'' ::* '' Graphium abri'' Smith & Vane-Wright, 2001 ::* '' Graphium adamastor'' (Boisduval, 1836) – Boisduval's white-lady ::* ''
Graphium agamedes ''Graphium agamedes'', the Westwood's white lady, is a butterfly in the family Papilionidae (swallowtails). It is found in Ivory Coast, Ghana, Togo, Benin, Nigeria, Cameroon, the Central African Republic and the Democratic Republic of the Congo. ...
'' (Westwood, 1842) – Westwood's white-lady or glassy graphium ::* ''
Graphium almansor ''Graphium almansor'', the Almansor white-lady swordtail or Honrath's white lady, is a butterfly in the family Papilionidae (swallowtails). It is found in Guinea, Ghana, Togo, Nigeria, Cameroon, the Republic of the Congo, Angola, the Central Afr ...
'' (Honrath, 1884) – Almansor white-lady swordtail or Honrath's white-lady ::* '' Graphium auriger'' (Butler, 1876) ::* '' Graphium aurivilliusi'' (Seeldrayers, 1896) ::* '' Graphium fulleri'' (Grose-Smith, 1883) ::* '' Graphium hachei'' (Dewitz, 1881) ::* '' Graphium kigoma'' Carcasson, 1964 ::* '' Graphium olbrechtsi'' Berger, 1950 ::* '' Graphium poggianus'' (Honrath, 1884) ::* '' Graphium rileyi'' Berger, 1950 - Riley's graphium ::* '' Graphium schubotzi'' (Schultze, 1913) ::* '' Graphium simoni'' (Aurivillius, 1899) ::* '' Graphium ucalegon'' (Hewitson, 1865) – creamy graphium ::* '' Graphium ucalegonides'' (Staudinger, 1884) subgenus: ''Pathysa'' :species group: ''antiphates'' ::* '' Graphium agetes'' ( Westwood, 1841) – fourbar swordtail ::* '' Graphium androcles'' (Boisduval, 1836) – lion swordtail, giant swordtail ::* '' Graphium antiphates'' (Cramer, 775 – fivebar swordtail ::* '' Graphium aristeus'' (Stoll, 780 – chain swordtail ::* '' Graphium decolor'' (Staudinger, 1888) ::* '' Graphium dorcus'' (de Haan, 1840) – Tabitha's swordtail ::* '' Graphium epaminondas'' (Oberthür, 1879) ::* '' Graphium euphrates'' (C. Felder & R. Felder, 1862) – Euphrates swordtail ::* '' Graphium euphratoides'' (Eimer, 1889) ::* '' Graphium nomius'' (Esper, 1799) – spot swordtail ::* '' Graphium rhesus'' (Boisduval, 1836) – monkey swordtail ::* '' Graphium stratiotes'' (Grose-Smith, 1887) subgenus: ''Paranticopsis'' Wood-Mason & de Nicéville, 1887 ::* '' Graphium delessertii'' (Guérin-Méneville, 1839) – Malayan zebra ::* '' Graphium deucalion'' (Boisduval, 1836) – yellow zebra ::* '' Graphium encelades'' (Boisduval, 1836) ::* '' Graphium idaeoides'' (Hewitson, 1853) ::* '' Graphium macareus'' (Godart, 1819) – lesser zebra ::* '' Graphium megaera'' ( Staudinger, 1888) ::* '' Graphium megarus'' (Westwood, 1844) – spotted zebra ::* '' Graphium ramaceus'' (Westwood, 1872) ::* '' Graphium phidias'' (Oberthür, 1906) ::* '' Graphium stratocles'' (C. Felder & R. Felder, 1861) ::* '' Graphium thule'' (Wallace, 1865) ::* '' Graphium xenocles'' (Doubleday, 1842) – great zebra subgenus: ''Pazala'' Moore, 1888 ::* '' Graphium alebion'' (Gray, 853 ::* '' Graphium eurous'' (Leech, 893 – sixbar swordtail ::* '' Graphium glycerion'' (Gray, 1831) – spectacle swordtail ::* '' Graphium incerta'' Bang-Haas, 1927 ::* '' Graphium mandarinus'' (Oberthür, 1879) – spectacle swordtail ::* '' Graphium mullah'' (Alphéraky 1897)::* '' Graphium tamerlana'' (Oberthür, 1876) ::* '' Graphium timur'' (Ney, 1911) synonymised with '' Graphium mullah'' by Cotton and Racheli, T., 2007 ::* '' Graphium wenlingae'' Hu, Condamine, Monastyrskii & Cotton, 2019


See also

* '' Protographium''


References

*Smith, Campbell R. and Vane-Wright R.I., 2001 A review of the Afrotropical species of the genus Graphium (Lepidoptera: Rhopalocera: Papilionidae) ''Bulletin of the Natural History Museum Entomology Series'' Volume 70:503-71
online
*Miller, L. D., and Miller, J. Y. (2004). ''The Butterfly Handbook'', pp. 20–25, 52. Barron's Educational Series, Inc.; Hauppauge, New York. *Smart, P. (1976). ''The Illustrated Encyclopedia of the Butterfly World''.


External links

* *
"Distribution of Afrotropical Kite Swallowtails"
(broadly subgenera ''Pazala'' and ''Arisbe'') Natural History Museum.
"Genus ''Graphium''"
''Insecta.pro''. {{Taxonbar, from=Q1768585 Butterfly genera Papilionidae Taxa named by Giovanni Antonio Scopoli