Glareolidae
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Glareolidae is a
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
of
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
s in the
wader 245px, A flock of Dunlins and Red knots">Red_knot.html" ;"title="Dunlins and Red knot">Dunlins and Red knots Waders or shorebirds are birds of the order Charadriiformes commonly found wikt:wade#Etymology 1, wading along shorelines and mudflat ...
suborder Charadrii. It contains two distinct groups, the
pratincole The pratincoles or greywaders are a group of birds which together with the coursers make up the family Glareolidae. They have short legs, very long pointed wings and long forked tails. Description Their most unusual feature for birds classed as ...
s and the
courser The coursers are a group of birds which together with the pratincoles make up the family Glareolidae. They have long legs, short wings and long pointed bills which curve downwards. Their most unusual feature for birds classed as waders is that ...
s. The atypical
Egyptian plover The Egyptian plover (''Pluvianus aegyptius''), also known as the crocodile bird, is a wader, the only member of the genus ''Pluvianus''. Formerly placed in the pratincole and courser family, Glareolidae, it is now regarded as the sole member of ...
(''Pluvianus aegyptius''), traditionally placed in this family, is now known to be only distantly related. The family contains 17 species in 4 genera.


Description

The feature that defines the family from the rest of the order is the
bill Bill(s) may refer to: Common meanings * Banknote, paper cash (especially in the United States) * Bill (law), a proposed law put before a legislature * Invoice, commercial document issued by a seller to a buyer * Bill, a bird or animal's beak Plac ...
, which is arched and has the nostrils at the base. The
pratincole The pratincoles or greywaders are a group of birds which together with the coursers make up the family Glareolidae. They have short legs, very long pointed wings and long forked tails. Description Their most unusual feature for birds classed as ...
s have short legs, long pointed wings and long forked tails. They have a buoyant flight that allows them the unusual (for the order) hunting technique of taking their
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...
prey on the wing like
swallows The swallows, martins, and saw-wings, or Hirundinidae, are a family of passerine songbirds found around the world on all continents, including occasionally in Antarctica. Highly adapted to aerial feeding, they have a distinctive appearance. The ...
. The wings also allow for long migrations in some species. The
courser The coursers are a group of birds which together with the pratincoles make up the family Glareolidae. They have long legs, short wings and long pointed bills which curve downwards. Their most unusual feature for birds classed as waders is that ...
s have long legs, which are used to run (giving the group its name). The wings are shorter and have a more sustained flight than that of the pratincoles.


Distribution and habitat

The pratincoles and coursers have an
Old World The "Old World" is a term for Afro-Eurasia that originated in Europe , after Europeans became aware of the existence of the Americas. It is used to contrast the continents of Africa, Europe, and Asia, which were previously thought of by the ...
distribution, occurring in southern
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
,
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area ...
,
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
(including
Madagascar Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Africa ...
), and
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
. The family is thought to have
evolved Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variation t ...
in Africa, which is where the family achieves its greatest diversity, although fossils of the genus ''Glareola'' belonging to an extinct species ''Glareola neogena'' are known from the
Middle Miocene The Middle Miocene is a sub-epoch of the Miocene Epoch made up of two stages: the Langhian and Serravallian stages. The Middle Miocene is preceded by the Early Miocene. The sub-epoch lasted from 15.97 ± 0.05 Ma to 11.608 ± 0.005 Ma (million y ...
of Europe, while of similar age is of the extinct ''Mioglareola gregaria'' also from European deposits. The older glareolid fossils are of the genus ''Paractiornis'' from the Lower Miocene of North America. The fossil genera and species are: *''Boutersemia belgica'' G. Mayr and R. Smith, 2001
Early Oligocene The Rupelian is, in the geologic timescale, the older of two ages or the lower of two stages of the Oligocene Epoch/Series. It spans the time between . It is preceded by the Priabonian Stage (part of the Eocene) and is followed by the Chatti ...
of Belgium *''Boutersemia parvula'' G. Mayr and R. Smith, 2001 Early Oligocene of Belgium *''Paractiornis perpusillus'' Wetmore, 1930
Early Miocene The Early Miocene (also known as Lower Miocene) is a sub-epoch of the Miocene Epoch made up of two stages: the Aquitanian and Burdigalian stages. The sub-epoch lasted from 23.03 ± 0.05 Ma to 15.97 ± 0.05 Ma (million years ago). It was prece ...
of Nebraska *''Mioglareola dolnicensis'' (Švec, 1980) Early Miocene of the Czech Republic *''Mioglareola gregaria'' Ballmann, 1979 Early Miocene of the Czech Republic and Middle Miocene of Germany *''Glareola neogena'' Ballmann, 1979 Middle Miocene of Germany *''"Gallinago" veterior'' Jánossy, 1979
Early Pliocene Early may refer to: History * The beginning or oldest part of a defined historical period, as opposed to middle or late periods, e.g.: ** Early Christianity ** Early modern Europe Places in the United States * Early, Iowa * Early, Texas * Early ...
of Hungary The coursers are typically found in open and arid environments such as deserts and scrub. The three-banded courser and bronze-winged courser are exceptions, being found in woodland and usually away from open land. The subfamily is usually also associated with lowland areas, although the
Burchell's courser Burchell's courser (''Cursorius rufus'') is a wader in the pratincole and courser family, Glareolidae. The name of this bird commemorates the English naturalist William John Burchell. Native to Africa, Burchell's courser is a small, diurnal, an ...
is found in southern Africa's Afro-alpine areas. The pratincoles are associated with wetlands, rivers, estuaries and other inland waterways. As with the coursers there are exceptions, particularly the
black-winged pratincole The black-winged pratincole (''Glareola nordmanni'') is a wader in the pratincole bird family, Glareolidae. The genus name is a diminutive of Latin ''glarea'', "gravel", referring to a typical nesting habitat for pratincoles. The species nam ...
which breeds and feeds on open steppes. Some species of pratincole are long-distance migrants. Shorter migrations include those of the
Madagascar pratincole The Madagascar pratincole (''Glareola ocularis'') is a species of bird in the family Glareolidae. It is found in Comoros, Ethiopia, Kenya, Madagascar, Mozambique, Somalia, Tanzania, possibly Mauritius, and possibly Réunion. Its natural habitats ...
, which migrates from its breeding grounds in Madagascar to East Africa; in contrast the black-winged pratincole migrates from the steppes of Eastern Europe and Central Asia to West and Southern Africa. The migration, which can measure in distance, is often undertaken as a single non-stop flight and is flown at high altitude. The coursers are not particularly migratory, although the
cream-colored courser The cream-colored courser (''Cursorius cursor'') is a wader in the pratincole and courser family, Glareolidae. Both parts of the scientific name derive from Latin ''cursor'', "runner", from ''currere'', "to run" which describes their usual habi ...
does migrate from the northern extremes of its range in the winter. The coursers are fairly nomadic, but do not undertake long-distance migrations.


Behaviour

The coursers are
crepuscular In zoology, a crepuscular animal is one that is active primarily during the twilight period, being matutinal, vespertine, or both. This is distinguished from diurnal and nocturnal behavior, where an animal is active during the hours of daylig ...
and
nocturnal Nocturnality is an animal behavior characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnal meaning the opposite. Nocturnal creatures generally have highly developed sens ...
in their habits, and are generally inconspicuous, particularly the woodland species. They are not as social as the highly gregarious and noisy pratincoles, some species of which may also active at dawn and dusk.


Diet and feeding

Insects form the majority of the diet of the Glareolidae. The pratincoles forage mainly on the wing, but are able to take prey on the ground as well. They are opportunistic, and have been recorded attending herds of antelope to snatch insects flushed up by their movement, or even insects attracted to street lights. Swarming insects, such as locusts or termites, are particularly targeted. Coursers are exclusively terrestrial, and feed in a plover-like fashion, running, then stopping to scan for prey before moving on. Some species may dig for insects in soft soil with their bills. In addition to insects, coursers may also take molluscs and some seeds.


References

{{Taxonbar, from=Q217272 Bird families Taxa named by Christian Ludwig Brehm