Giant petrel
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Giant petrels form a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
, ''Macronectes'', from the family
Procellariidae The family Procellariidae is a group of seabirds that comprises the fulmarine petrels, the gadfly petrels, the diving petrels, the prions, and the shearwaters. This family is part of the bird order Procellariiformes (or tubenoses), which als ...
, which consists of two species. They are the largest birds of this family. Both species are restricted to the Southern Hemisphere, and though their distributions overlap significantly, with both species breeding on the
Prince Edward Islands The Prince Edward Islands are two small uninhabited islands in the sub-Antarctic Indian Ocean that are part of South Africa. The islands are named Marion Island (named after Marc-Joseph Marion du Fresne, 1724–1772) and Prince Edward Island ...
, Crozet Islands, Kerguelen Islands,
Macquarie Island Macquarie Island is an island in the southwestern Pacific Ocean, about halfway between New Zealand and Antarctica. Regionally part of Oceania and politically a part of Tasmania, Australia, since 1900, it became a Tasmanian State Reserve in 197 ...
, and South Georgia, many southern giant petrels nest farther south, with
colonies In modern parlance, a colony is a territory subject to a form of foreign rule. Though dominated by the foreign colonizers, colonies remain separate from the administration of the original country of the colonizers, the '' metropolitan state'' ...
as far south as
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest cont ...
. Giant petrels are extremely aggressive
predator Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill th ...
s and scavengers, inspiring another common name, the stinker. South Sea whalers used to call them gluttons. They are the only member of their family that is capable of walking on land.


Taxonomy

The genus ''Macronectes'' was introduced in 1905 by the American ornithologist
Charles Wallace Richmond Charles Wallace Richmond (December 31, 1868 – May 19, 1932) was an American ornithologist. He is best remembered for a compilation of the Latin names of birds that is called the Richmond Index. Life and work He was born in Kenosha, Wisconsin an ...
to accommodate what is now the southern giant petrel. It replaced the previous genus ''Ossifraga'' which was found to have been earlier applied to a different group of birds. The name ''Macronectes'' combines the
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic p ...
''makros'' meaning "great" and ''nēktēs'' meaning "swimmer". The giant petrels are two large
seabird Seabirds (also known as marine birds) are birds that are adapted to life within the marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent evolution, as the same enviro ...
s from the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
''Macronectes''. Long considered to be
conspecific Biological specificity is the tendency of a characteristic such as a behavior or a biochemical variation to occur in a particular species. Biochemist Linus Pauling stated that "Biological specificity is the set of characteristics of living organis ...
(they were not established as separate species until 1966), the two species, the southern giant petrel, ''M. giganteus'', and
northern giant petrel The northern giant petrel (''Macronectes halli''), also known as Hall's giant petrel, is a large predatory seabird of the southern oceans. Its distribution overlaps broadly, but is slightly north of, the similar southern giant petrel (''Macrone ...
, ''M. halli'', are considered with the two
fulmar The fulmars are tubenosed seabirds of the family Procellariidae. The family consists of two extant species and two extinct fossil species from the Miocene. Fulmars superficially resemble gulls, but are readily distinguished by their flight on ...
s, ''Fulmarus'', to form a distinct subgroup within the Procellariidae, and including the Antarctic petrel, Cape petrel, and
snow petrel The snow petrel (''Pagodroma nivea'') is the only member of the genus ''Pagodroma.'' It is one of only three birds that has been seen at the Geographic South Pole, along with the Antarctic petrel and the south polar skua, which have the most s ...
, they form a separate group from the rest of the family.


Description

The southern giant petrel is slightly larger than the northern giant petrel, at , across the wings, and of body length. The northern giant petrel is , across the wings and of body length. They superficially resemble the albatross, and are the only procellarids that can equal them in size. They can be separated from the albatrosses by their
bill Bill(s) may refer to: Common meanings * Banknote, paper cash (especially in the United States) * Bill (law), a proposed law put before a legislature * Invoice, commercial document issued by a seller to a buyer * Bill, a bird or animal's beak Plac ...
; the two tube nostrils are joined together on the top of the bill, unlike on albatross, where they are separated and on the side of the bill. Giant petrels are also the only members of the family Procellariidae to have strong legs to walk on land. They are also much darker and more mottled brown (except for the white morph southern, which are whiter than any albatross) and have a more hunch-backed look. The bills of Procellariiformes are also unique in that they are split into between seven and nine horny plates. The petrels have a hooked bill called the maxillary unguis which can hold slippery prey. They produce a stomach oil made up of
wax ester A wax ester (WE) is an ester of a fatty acid and a fatty alcohol. Wax esters comprise the main components of three commercially important waxes: carnauba wax, candelilla wax, and beeswax.. Wax esters are formed by combining one fatty acid with on ...
s and
triglycerides A triglyceride (TG, triacylglycerol, TAG, or triacylglyceride) is an ester derived from glycerol and three fatty acids (from ''tri-'' and ''glyceride''). Triglycerides are the main constituents of body fat in humans and other vertebrates, as we ...
which is stored in the
proventriculus The proventriculus is part of the digestive system of birds.Encarta World English Dictionary orth American Edition(2007). ''Proventriculus''. Source: (accessed: December 18, 2007) An analogous organ exists in invertebrates and insects. Birds Th ...
. This can be sprayed out of their mouths as a defence against predators and as a protein-rich food source for chicks and for the adults during their long flights. Petrels have a
salt gland The salt gland is an organ for excreting excess salts. It is found in the cartilaginous fishes subclass elasmobranchii (sharks, rays, and skates), seabirds, and some reptiles. Salt glands can be found in the rectum of sharks. Birds and reptiles ...
situated above the nasal passage that helps to desalinate their bodies by excreting a high saline solution from their noses. The two species are difficult to tell from each other, possessing similar long, pale, orange bills and uniform, mottled grey plumage (except for around 15% of southern petrels, which are almost completely white). The billtip of ''M. halli'' is reddish-pink and that of ''M. giganteus'' is pale green, appearing slightly darker and lighter than the rest of the bill, respectively. The underside of older ''M. halli'' birds is paler and more uniform than ''M. giganteus'', the latter showing a contrast between paler head and neck and darker belly. Additionally, adults of ''M. halli'' typically appear pale-eyed, while adults of ''M. giganteus'' of the normal morph typically appear dark-eyed (occasionally flecked paler). Classic examples of northern giant are identifiable at some range. Young birds of both species are all dark and very hard to distinguish unless bill tip colour can be seen. Some relatively young northern giant petrels can appear to be paler on the head, suggesting southern giant, thus this species is harder to confirm.


Etymology

''Macronectes'' comes from the
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
words ''makros'' meaning "long" and ''nēktēs'' meaning "swimmer". Also, petrel is derived from St. Peter and the story of his walking on water, as they appear to run on the water when they take off.


Behaviour


Feeding

Petrels are highly opportunistic feeders. Unique among procellarids, they will feed both on land and at sea; in fact, they find most of their food near coastlines. On land, they feed on carrion, and regularly scavenge the breeding colonies of
penguins Penguins (order Sphenisciformes , family Spheniscidae ) are a group of aquatic flightless birds. They live almost exclusively in the Southern Hemisphere: only one species, the Galápagos penguin, is found north of the Equator. Highly adapt ...
and seals. They will display their dominance over carcasses with a "sealmaster posture": the head and the wings are held outstretched, the head pointing at the opponent and the wingtips pointing slightly back; the tail is raised to a vertical position. They are extremely aggressive and will kill other seabirds (usually penguin chicks, sick or injured adult penguins and the chicks of other seabirds), even those as large as an albatross, which they kill either by battering them to death or drowning. At sea, they feed on
krill Krill are small crustaceans of the order Euphausiacea, and are found in all the world's oceans. The name "krill" comes from the Norwegian word ', meaning "small fry of fish", which is also often attributed to species of fish. Krill are consi ...
, squid, and fish. They often follow
fishing boats A fishing vessel is a boat or ship used to catch fish in the sea, or on a lake or river. Many different kinds of vessels are used in commercial, artisanal and recreational fishing. The total number of fishing vessels in the world in 2016 was es ...
and other ships, in the hope of picking up
offal Offal (), also called variety meats, pluck or organ meats, is the organs of a butchered animal. The word does not refer to a particular list of edible organs, which varies by culture and region, but usually excludes muscle. Offal may also refe ...
and other waste.


Reproduction

The southern giant petrel is more likely to form loose colonies than the northern, both species laying a single egg in a rough
nest A nest is a structure built for certain animals to hold eggs or young. Although nests are most closely associated with birds, members of all classes of vertebrates and some invertebrates construct nests. They may be composed of organic materi ...
built about off the ground. The
egg An egg is an organic vessel grown by an animal to carry a possibly fertilized egg cell (a zygote) and to incubate from it an embryo within the egg until the embryo has become an animal fetus that can survive on its own, at which point the a ...
is incubated for about 60 days; once hatched the chick is brooded for three weeks. Chicks fledge after about four months, but do not achieve sexual maturity for six or seven years after fledging.


Conservation

While both species were listed as
near threatened A near-threatened species is a species which has been categorized as "Near Threatened" (NT) by the International Union for Conservation of Nature as that may be vulnerable to endangerment in the near future, but it does not currently qualify f ...
in the 2008
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biol ...
, subsequent evidence suggested they were less threatened than previously believed, and the populations of both actually appeared to have increased, at least locally. Consequently, they were listed as
least concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. T ...
on the 2009 Red List and afterwards (as of IUCN's last assessment in 2018, they continue to be listed as least concern). The southern giant petrel is listed as endangered on the Australian
Environment Protection and Biodiversity Conservation Act 1999 The ''Environment Protection and Biodiversity Conservation Act 1999'' (Cth) is an Act of the Parliament of Australia that provides a framework for protection of the Australian environment, including its biodiversity and its natural and cult ...
, while the northern giant petrel is listed on the same act as vulnerable. Their conservation status also varies from state to state within Australia.


Gallery

File:Giant petrel and king penguins.jpg, Giant petrel which has just killed a
king penguin The king penguin (''Aptenodytes patagonicus'') is the second largest species of penguin, smaller, but somewhat similar in appearance to the emperor penguin. There are two subspecies: ''A. p. patagonicus'' and ''A. p. halli''; ''patagonicus'' ...
chick File:Giant petrel flies at soutg georgia.jpg, Giant petrel flying above South Georgia Island


References


Sources

* * * del Hoyo, Josep, Elliott, Andrew & Sargatal, Jordi (1992). ''Handbook of Birds of the World'' Vol 1. Barcelona:Lynx Edicions,


External links

*Southern and Northern Giant Petrels
Species text in The Atlas of Southern African Birds

Video of George the Giant Petrel being released by New Zealand Department of Conservation
{{Taxonbar, from=Q598979 Macronectes Taxa named by Charles Wallace Richmond