The collared peccary (''Dicotyles tajacu'') is a species of

artiodactyl The even-toed ungulates (Artiodactyla , ) are ungulates—hoofed animals—which bear weight equally on two (an even number) of their five toes: the third and fourth. The other three toes are either present, absent, vestigial, or pointing poster ...

(even-toed)

mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur ...

in the family Tayassuidae found in

North North is one of the four compass points or cardinal directions. It is the opposite of south and is perpendicular to east and west. ''North'' is a noun, adjective, or adverb indicating direction or geography. Etymology The word ''north ...

Central Central is an adjective usually referring to being in the center of some place or (mathematical) object. Central may also refer to: Directions and generalised locations * Central Africa, a region in the centre of Africa continent, also known a ...

, and

South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sou ...

. It is the only member of the genus ''Dicotyles''. They are commonly referred to as ''javelina, saíno'', or ''báquiro'', although these terms are also used to describe other species in the family. The species is also known as the musk hog. In Trinidad, it is colloquially known as ''quenk''.

Taxonomy

Although somewhat related to true Old World pigs, and frequently referred to as a pig, this species and the other peccaries are no longer classified in the pig family,

Suidae Suidae is a family of artiodactyl mammals which are commonly called pigs, hogs or swine. In addition to numerous fossil species, 18 extant species are currently recognized (or 19 counting domestic pigs and wild boars separately), classified in ...

. Although formerly classified in the genus ''Pecari'', studies in 2020 placed them in the genus ''Dicotyles'', based on an unequivocal type-species selection; these studies have been accepted by the

American Society of Mammalogists The American Society of Mammalogists (ASM) was founded in 1919. Its primary purpose is to encourage the study of mammals, and professions studying them. There are over 4,500 members of this society, and they are primarily professional scientists w ...

. Currently, the

IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...

still places them in the genus ''Pecari''.

Description

The collared peccary stands around tall at the shoulder and is about long. It weighs between . The

dental formula Dentition pertains to the development of teeth and their arrangement in the mouth. In particular, it is the characteristic arrangement, kind, and number of teeth in a given species at a given age. That is, the number, type, and morpho-physiolog ...

is: 2/3,1/1,3/3,3/3. The collared peccary has small tusks that point toward the ground when the animal is upright. It has slender legs with a robust or stocky body. The tail is often hidden in the coarse fur of the peccary.

Range and habitat

The collared peccary is widespread throughout much of the tropical and subtropical

Americas The Americas, which are sometimes collectively called America, are a landmass comprising the totality of North America, North and South America. The Americas make up most of the land in Earth's Western Hemisphere and comprise the New World. ...

, ranging from the

Southwestern United States The Southwestern United States, also known as the American Southwest or simply the Southwest, is a geographic and cultural region of the United States that generally includes Arizona, New Mexico, and adjacent portions of California, Colorado, N ...

to northern

Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the List of South American countries by area, second-largest ...

. They were reintroduced to

Uruguay Uruguay (; ), officially the Oriental Republic of Uruguay ( es, República Oriental del Uruguay), is a country in South America. It shares borders with Argentina to its west and southwest and Brazil to its north and northeast; while bordering ...

in 2017, after 100 years of extirpation there. The only

Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean ...

island where it is native, however, is

Trinidad Trinidad is the larger and more populous of the two major islands of Trinidad and Tobago. The island lies off the northeastern coast of Venezuela and sits on the continental shelf of South America. It is often referred to as the southernmos ...

. Until fairly recently, it was also present on the nearby island of

Tobago Tobago () is an island and ward within the Republic of Trinidad and Tobago. It is located northeast of the larger island of Trinidad and about off the northeastern coast of Venezuela. It also lies to the southeast of Grenada. The offic ...

, but is now exceedingly rare (if not extirpated) due to

overhunting Overexploitation, also called overharvesting, refers to harvesting a renewable resource to the point of diminishing returns. Continued overexploitation can lead to the destruction of the resource, as it will be unable to replenish. The term ap ...

by humans. An adaptable species, it inhabits

desert A desert is a barren area of landscape where little precipitation occurs and, consequently, living conditions are hostile for plant and animal life. The lack of vegetation exposes the unprotected surface of the ground to denudation. About on ...

s, xeric shrublands, tropical and subtropical grasslands, savannas, shrublands, flooded grasslands and savannas,

tropical and subtropical dry broadleaf forests The tropical and subtropical dry broadleaf forest is a habitat type defined by the World Wide Fund for Nature and is located at tropical and subtropical latitudes. Though these forests occur in climates that are warm year-round, and may receive ...

, and several other habitats; it is also present in habitats shared by humans, merely requiring sufficient cover. Peccaries can be found in

cities A city is a human settlement of notable size.Goodall, B. (1987) ''The Penguin Dictionary of Human Geography''. London: Penguin.Kuper, A. and Kuper, J., eds (1996) ''The Social Science Encyclopedia''. 2nd edition. London: Routledge. It can be def ...

and agricultural land throughout their range, where they consume garden plants. Notable populations are known to exist in the

suburbs A suburb (more broadly suburban area) is an area within a metropolitan area, which may include commercial and mixed-use, that is primarily a residential area. A suburb can exist either as part of a larger city/urban area or as a separa ...

Phoenix Phoenix most often refers to: * Phoenix (mythology), a legendary bird from ancient Greek folklore * Phoenix, Arizona, a city in the United States Phoenix may also refer to: Mythology Greek mythological figures * Phoenix (son of Amyntor), a ...

and

Tucson, Arizona , "(at the) base of the black ill , nicknames = "The Old Pueblo", "Optics Valley", "America's biggest small town" , image_map = , mapsize = 260px , map_caption = Interactive map ...

. Due to the lack of fossil material or even specimens from archeological sites, it was assumed that javelinas only recently crossed into the US by way of Mexico. However, a fossil jaw of this species was discovered in Florida ("Collared peccary (Mammalia, Artiodactyla, Tayassuidae, Pecari) from the late Pleistocene of Florida", Richard C. Hulbert, Gary S. Morgan & Andreas Kerner), proving that at some point in the late Pleistocene the species had already inhabited part of the Southern US.

Diet

Collared peccaries are often classified as herbivores. They normally feed on

cactus A cactus (, or less commonly, cactus) is a member of the plant family Cactaceae, a family comprising about 127 genera with some 1750 known species of the order Caryophyllales. The word ''cactus'' derives, through Latin, from the Ancient Gree ...

mesquite Mesquite is a common name for several plants in the genus '' Prosopis'', which contains over 40 species of small leguminous trees. They are native to dry areas in the Americas. They have extremely long roots to seek water from very far under gr ...

beans, fruits, berries, seeds, roots,

tuber Tubers are a type of enlarged structure used as storage organs for nutrients in some plants. They are used for the plant's perennation (survival of the winter or dry months), to provide energy and nutrients for regrowth during the next growing ...

bulb In botany, a bulb is structurally a short stem with fleshy leaves or leaf basesBell, A.D. 1997. ''Plant form: an illustrated guide to flowering plant morphology''. Oxford University Press, Oxford, U.K. that function as food storage organs dur ...

palm nut Palm most commonly refers to: * Palm of the hand, the central region of the front of the hand * Palm plants, of family Arecaceae **List of Arecaceae genera * Several other plants known as "palm" Palm or Palms may also refer to: Music * Palm (ba ...

s, grasses, other green vegetation,

fungi A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately fr ...

, and insects. However, they will also eat eggs, snakes, fish, frogs, lizards, dead birds, and rodents if the opportunity presents itself. Despite all this supplementary diet, the main dietary components of this species are

agave ''Agave'' (; ; ) is a genus of monocots native to the hot and arid regions of the Americas and the Caribbean, although some ''Agave'' species are also native to tropical areas of North America, such as Mexico. The genus is primarily known for ...

s and prickly pears. In areas inhabited by humans, they also consume cultivated crops and

ornamental plant Ornamental plants or garden plants are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars that ...

s, such as

tulip Tulips (''Tulipa'') are a genus of spring-blooming perennial herbaceous bulbiferous geophytes (having bulbs as storage organs). The flowers are usually large, showy and brightly coloured, generally red, pink, yellow, or white (usually in warm ...

Predators

The main predators of the collared peccary are

cougar The cougar (''Puma concolor'') is a large cat native to the Americas. Its range spans from the Canadian Yukon to the southern Andes in South America and is the most widespread of any large wild terrestrial mammal in the Western Hemisphere. ...

s (''Puma concolor''), Mexican wolves (''Canis lupus baileyi''),

coyote The coyote (''Canis latrans'') is a species of canine native to North America. It is smaller than its close relative, the wolf, and slightly smaller than the closely related eastern wolf and red wolf. It fills much of the same ecological nich ...

s (''Canis latrans''),

jaguar The jaguar (''Panthera onca'') is a large cat species and the only living member of the genus ''Panthera'' native to the Americas. With a body length of up to and a weight of up to , it is the largest cat species in the Americas and the th ...

s (''Panthera onca''), and

bobcat The bobcat (''Lynx rufus''), also known as the red lynx, is a medium-sized cat native to North America. It ranges from southern Canada through most of the contiguous United States to Oaxaca in Mexico. It is listed as Least Concern on the ...

s (''Lynx rufus'').

Behavior

Collared peccaries are diurnal creatures that live in groups of up to 50 individuals, averaging between six and nine animals. They sleep in burrows (often under bushes or larger systems of tree roots), but sometimes can be found in caves, abandoned mines, old desert tunnels, or among logs, felled trees and abandoned timber. However, collared peccaries are not completely diurnal. In central Arizona, they are often more active at night, and less so in the heat of the daytime. Although they usually ignore humans, they will react if they feel threatened. They defend themselves with their tusks. A collared peccary can release a strong musk or give a sharp bark if it is alarmed. Amazonian peoples (including the Shipibos) sometimes raise and tame juvenile collared peccaries, if they are encountered. Peccary will also rub their scent onto rocks and tree stumps to mark their territory, and rub the scent on each other to help with identification.

The "giant peccary"

The giant peccary (described as ''Pecari maximus'') was a purported fourth species of

peccary A peccary (also javelina or skunk pig) is a medium-sized, pig-like hoofed mammal of the family Tayassuidae (New World pigs). They are found throughout Central and South America, Trinidad in the Caribbean, and in the southwestern area of North ...

, first reported to have been seen in

Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...

in 2000 by Dutch naturalist

Marc van Roosmalen Dr. Marc van Roosmalen (born June 23, 1947) is a Dutch-Brazilian primatologist. He was elected as one of the "Heroes of the Planet" by ''Time'' magazine in 2000. His research has led to the identification of several new monkey species, as well a ...

. In 2003 German natural history filmmaker Lothar Frenz filmed a group and gathered a skull which later served as the type (INPA4272). It had been known locally as '' caitetú-mundè'', which Roosmalen ''et al''. state the locals claimed was

Tupí Tupí, also known as ''formatge de tupí'', is a fermented cheese of a certain area of the Pyrenees and Pre-Pyrenees made from cows' or sheep's milk. It is a cheese traditionally prepared in the mountainous Pallars region, as well as in the Cerd ...

and meant "the collared peccary that is bigger and goes in pairs", as opposed to ''caitetú-de-bando'', "the collared peccary that goes in herds". It was formally described in 2007,Roosmalen, M.G.M.; Frenz, L.; Hooft, W.F. van; Iongh, H.H. de; Leirs, H. 2007
''A New Species of Living Peccary (Mammalia: Tayassuidae) from the Brazilian Amazon.''
Bonner zoologische Beiträge 55(2): 105–112. but the scientific evidence for its species status was quickly questioned,Gongora, J., Taber, A., Keuroghlian, A., Altrichter, M., Bodmer, R.E., Mayor, P., Moran, C., Damayanti, C.S., González S. (2007).
Re-examining the evidence for a ‘new’ peccary species, ''Pecari maximus'', from the Brazilian Amazon
'' Newsletter of the Pigs, Peccaries, and Hippos Specialist Group of the IUCN/SSC. 7(2): 19–26. which also was one of the reasons for its initial evaluation as

data deficient A data deficient (DD) species is one which has been categorized by the International Union for Conservation of Nature (IUCN) as offering insufficient information for a proper assessment of conservation status to be made. This does not necessaril ...

in 2008. A review in 2011 moved the giant peccary into

synonymy A synonym is a word, morpheme, or phrase that means exactly or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are all ...

with the collared peccary (''P. tajacu''),Gongora, J., Biondo, C., Cooper, J.D., Taber, A., Keuroghlian, A., Altrichter, M., Ferreira do Nascimento, F., Chong, A.Y., Miyaki, C.Y., Bodmer, R., Mayor, P. and González, S. (2011).
Revisiting the species status of ''Pecari maximus'' van Roosmalen et al., 2007 (Mammalia) from the Brazilian Amazon
'' Bonn Zoological Bulletin 60(1): 95-101. which was followed by the

the same year. The reported range of the giant peccary encompasses the south-central

Amazon Amazon most often refers to: * Amazons, a tribe of female warriors in Greek mythology * Amazon rainforest, a rainforest covering most of the Amazon basin * Amazon River, in South America * Amazon (company), an American multinational technolog ...

between the

Madeira ) , anthem = ( en, "Anthem of the Autonomous Region of Madeira") , song_type = Regional anthem , image_map=EU-Portugal_with_Madeira_circled.svg , map_alt=Location of Madeira , map_caption=Location of Madeira , subdivision_type=Sovereign st ...

and the Tapajós Rivers and northern

Bolivia , image_flag = Bandera de Bolivia (Estado).svg , flag_alt = Horizontal tricolor (red, yellow, and green from top to bottom) with the coat of arms of Bolivia in the center , flag_alt2 = 7 × 7 square p ...

.Moravec, J., & Böhme, W. (2009).
Second Find of the Recently Discovered Amazonian Giant Peccary, ''Pecari maximus'' (Mammalia: Tayassuidae) van Roosmalen et al., 2007: First Record from Bolivia
.'' Bonner zoologische Beiträge 56(1-2): 49-54. It is restricted to ''terra firme'' forest, which is forest that does not flood annually. Unlike other peccaries in its range, the giant peccary was reported to mainly occur in pairs or small family groups. According to its original description, the giant peccary is larger, longer-legged, and proportionally smaller-headed than the only other member of the genus, the collared peccary. Compared to most individuals of the

sympatric In biology, two related species or populations are considered sympatric when they exist in the same geographic area and thus frequently encounter one another. An initially interbreeding population that splits into two or more distinct species s ...

populations of the collared peccary, the giant peccary also had thinner fur that is grizzled in brown and white, blacker legs, and a relatively faint collar. Five skins of the giant peccary had a total length of , while local hunters have estimated a weight of . Based on a

mtDNA Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial DNA ...

study, the collared and the giant peccaries were estimated to have diverged 1.0–1.2 million years ago, but these results were later questioned due to the small

sample size Sample size determination is the act of choosing the number of observations or replicates to include in a statistical sample. The sample size is an important feature of any empirical study in which the goal is to make inferences about a populati ...

, low bootstrap support, and the absence of

nDNA Nuclear DNA (nDNA), or nuclear deoxyribonucleic acid, is the DNA contained within each cell nucleus of a eukaryotic organism. It encodes for the majority of the genome in eukaryotes, with mitochondrial DNA and plastid DNA coding for the rest. I ...

and

cytogenetic Cytogenetics is essentially a branch of genetics, but is also a part of cell biology/cytology (a subdivision of human anatomy), that is concerned with how the chromosomes relate to cell behaviour, particularly to their behaviour during mitosis an ...

results. In 2011, a review noted that the measurements provided in the initial description were within those generally recognized for the collared peccary, and the behaviors supposedly unique to the giant peccary are also known from the collared peccary. They also provided new genetic evidence showing that collared peccaries from South America form a

monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gr ...

clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English ter ...

that includes the giant peccary (without it the clade is

paraphyletic In taxonomy, a group is paraphyletic if it consists of the group's last common ancestor and most of its descendants, excluding a few monophyletic subgroups. The group is said to be paraphyletic ''with respect to'' the excluded subgroups. In ...

). The major genetic split within the collared peccary is between a clade comprising North and Central American specimens, and a clade comprising South American specimens (the presumed contact zone is in

Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the ...

, which has both clades). Furthermore, extensive infraspecific variations (both individual and locality-based) are known in the morphology of the collared peccary.

Gallery

File:Collared peccaries as seen by a fixed camera in Scottsdale, Arizona.webm, A herd seen by a fixed camera in

Scottsdale, Arizona , settlement_type = City , named_for = Winfield Scott , image_skyline = , image_seal = Seal of Scottsdale (Arizona).svg , image_blank_emblem = City of Scottsdale Script Logo.svg , nic ...

File:Running Javelina.jpg, Running collared peccary File:Mother javelina and baby.jpg, Mother and juvenile File:Ancestral Pueblo, Roosevelt black on white javelina effigy canteen, 1250-1300 CE, Heard Museum.JPG, A Pueblo drinking vessel File:Ogrličasti pekari u Zagrebu.jpg, A herd of collared peccaries in Zagreb Zoo

Notes

References

External links

Smithsonian Institution - North American Mammals: ''Pecari tajacu''

Arizona Game and Fish Department – Living With Javelina
{{Authority control Peccaries Mammals of the United States Fauna of the Southwestern United States Fauna of the Sonoran Desert Ungulates of Central America Mammals of Mexico Mammals of Argentina Mammals of Bolivia Mammals of Brazil Mammals of Colombia Mammals of Ecuador Mammals of French Guiana Mammals of Guyana Mammals of Paraguay Mammals of Peru Mammals of Suriname Mammals of Trinidad and Tobago Mammals of Venezuela Mammals of the Caribbean Fauna of the Amazon Fauna of the Pantanal Mammals described in 1758 Articles containing video clips Taxa named by Carl Linnaeus Fauna of the Sierra Madre Occidental Taxobox binomials not recognized by IUCN