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A tide pool or rock pool is a shallow pool of seawater that forms on the rocky intertidal shore. Many of these pools exist as separate bodies of water only at
low tide Tides are the rise and fall of sea levels caused by the combined effects of the gravitational forces exerted by the Moon (and to a much lesser extent, the Sun) and are also caused by the Earth and Moon orbiting one another. Tide tables can ...
. Many tide pool habitats are home to especially adaptable animals that have engaged the attention of naturalists and
marine biologists Marine biology is the scientific study of the biology of marine life, organisms in the sea. Given that in biology many phyla, families and genera have some species that live in the sea and others that live on land, marine biology classifie ...
, as well as philosophical essayists:
John Steinbeck John Ernst Steinbeck Jr. (; February 27, 1902 – December 20, 1968) was an American writer and the 1962 Nobel Prize in Literature winner "for his realistic and imaginative writings, combining as they do sympathetic humor and keen social ...
wrote in ''
The Log from the Sea of Cortez ''The Log from the Sea of Cortez'' is an English-language book written by American author John Steinbeck and published in 1951. It details a six-week (March 11 – April 20) marine specimen-collecting boat expedition he made in 1940 at variou ...
'', "It is advisable to look from the tide pool to the stars and then back to the tide pool."


Zones from shallow to deep

The rocky shoreline exhibits zonation as a feature of the shoreline. Tidal movements of water creates zonation patterns along rocky shores from high to low-tide. The area above the high-tide mark is the supralittoral zone which is virtually a terrestrial environment. The area around the high-tide mark is known as the intertidal fringe. Between the high and low-tide marks is the intertidal or littoral zone. Below the low-tide mark is the sublittoral or subtidal zone. The presence and abundance of animals and algae vary between zones along the rocky shore due to niche adaptations in response to the varying levels of tidal and solar exposure. Tide pools exist in the "intertidal zone" (the area within the tidal range), which is submerged by the sea at high tides and during storms, and may receive spray from wave action. At other times the rocks may undergo other extreme conditions, baking in the sun or exposed to cold winds. Few organisms can survive such harsh conditions.
Lichen A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.barnacle A barnacle is a type of arthropod constituting the subclass Cirripedia in the subphylum Crustacea, and is hence related to crabs and lobsters. Barnacles are exclusively marine, and tend to live in shallow and tidal waters, typically in eros ...
s live in this zone. Different
barnacle A barnacle is a type of arthropod constituting the subclass Cirripedia in the subphylum Crustacea, and is hence related to crabs and lobsters. Barnacles are exclusively marine, and tend to live in shallow and tidal waters, typically in eros ...
species live at very tightly constrained elevations, with tidal conditions precisely determining the exact height of an assemblage relative to sea level. The intertidal zone is periodically exposed to sun and wind, conditions that can cause barnacles to become
desiccated Desiccation () is the state of extreme dryness, or the process of extreme drying. A desiccant is a hygroscopic (attracts and holds water) substance that induces or sustains such a state in its local vicinity in a moderately sealed container. ...
. These animals, therefore, need to be well adapted to water loss. Their
calcite Calcite is a Carbonate minerals, carbonate mineral and the most stable Polymorphism (materials science), polymorph of calcium carbonate (CaCO3). It is a very common mineral, particularly as a component of limestone. Calcite defines hardness 3 on ...
shells are impermeable, and they possess two plates which they slide across their mouth opening when not feeding. These plates also protect against predation.


High tide zone

The high tide zone is flooded during each high tide. Organisms must survive wave action, currents, and exposure to the sun. This zone is predominantly inhabited by
seaweed Seaweed, or macroalgae, refers to thousands of species of macroscopic, multicellular, marine algae. The term includes some types of '' Rhodophyta'' (red), ''Phaeophyta'' (brown) and ''Chlorophyta'' (green) macroalgae. Seaweed species such as ...
and invertebrates, such as
sea anemone Sea anemones are a group of predation, predatory marine invertebrates of the order (biology), order Actiniaria. Because of their colourful appearance, they are named after the ''Anemone'', a terrestrial flowering plant. Sea anemones are classifi ...
s, sea star,
chiton Chitons () are marine molluscs of varying size in the class Polyplacophora (), formerly known as Amphineura. About 940 extant and 430 fossil species are recognized. They are also sometimes known as gumboots or sea cradles or coat-of-mail s ...
s,
crab Crabs are decapod crustaceans of the infraorder Brachyura, which typically have a very short projecting "tail" (abdomen) ( el, βραχύς , translit=brachys = short, / = tail), usually hidden entirely under the thorax. They live in all the ...
s,
green algae The green algae (singular: green alga) are a group consisting of the Prasinodermophyta and its unnamed sister which contains the Chlorophyta and Charophyta/Streptophyta. The land plants (Embryophytes) have emerged deep in the Charophyte alga as ...
, and mussels. Marine
algae Algae (; singular alga ) is an informal term for a large and diverse group of photosynthetic eukaryotic organisms. It is a polyphyletic grouping that includes species from multiple distinct clades. Included organisms range from unicellular mic ...
provide shelter for nudibranchs and hermit crabs. The same waves and currents that make life in the high tide zone difficult bring food to
filter feeder Filter feeders are a sub-group of suspension feeding animals that feed by straining suspended matter and food particles from water, typically by passing the water over a specialized filtering structure. Some animals that use this method of feedin ...
s and other intertidal organisms.


Low tide zone

Also called the "lower littoral zone", this area is mostly submerged and is exposed only during unusually low tide. It often teems with life and has far more marine vegetation, especially seaweeds. There is also greater biodiversity. Organisms in this zone do not have to be as well adapted to drying out and temperature extremes. Low tide zone organisms include
abalone Abalone ( or ; via Spanish , from Rumsen ''aulón'') is a common name for any of a group of small to very large marine gastropod molluscs in the family (biology), family Haliotidae. Other common name In biology, a common name of a taxon o ...
, anemones, brown seaweed, chitons, crabs, green algae, hydroids, isopods,
limpet Limpets are a group of aquatic snails that exhibit a conical shell shape (patelliform) and a strong, muscular foot. Limpets are members of the class Gastropoda, but are polyphyletic, meaning the various groups called "limpets" descended indep ...
s, mussels, and sometimes even small vertebrates such as fish. These creatures can grow to larger sizes because there is more available energy and better water coverage: the water is shallow enough to allow additional sunlight for
photosynthetic Photosynthesis is a process used by plants and other organisms to convert light energy into chemical energy that, through cellular respiration, can later be released to fuel the organism's activities. Some of this chemical energy is stored in c ...
activity, with almost normal levels of
salinity Salinity () is the saltiness or amount of salt dissolved in a body of water, called saline water (see also soil salinity). It is usually measured in g/L or g/kg (grams of salt per liter/kilogram of water; the latter is dimensionless and equal ...
. This area is also relatively protected from large
predators Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill the ...
because of the wave action and shallow water.


Marine life

Tide pools provide a home for hardy organisms such as sea stars,
mussel Mussel () is the common name used for members of several families of bivalve molluscs, from saltwater and Freshwater bivalve, freshwater habitats. These groups have in common a shell whose outline is elongated and asymmetrical compared with other ...
s and clams. Inhabitants must be able to deal with a frequently changing
environment Environment most often refers to: __NOTOC__ * Natural environment, all living and non-living things occurring naturally * Biophysical environment, the physical and biological factors along with their chemical interactions that affect an organism or ...
: fluctuations in water temperature, salinity, and oxygen content. Hazards include waves, strong currents, exposure to midday sun and predators. Waves can dislodge mussels and draw them out to sea. Gulls pick up and drop
sea urchin Sea urchins () are spiny, globular echinoderms in the class Echinoidea. About 950 species of sea urchin live on the seabed of every ocean and inhabit every depth zone from the intertidal seashore down to . The spherical, hard shells (tests) of ...
s to break them open. Sea stars prey on mussels and are eaten by gulls themselves. Black bears are known to sometimes feast on intertidal creatures at low tide. Although tide pool organisms must avoid getting washed away into the ocean, drying up in the sun, or being eaten, they depend on the tide pool's constant changes for food.


Fauna

The
sea anemone Sea anemones are a group of predation, predatory marine invertebrates of the order (biology), order Actiniaria. Because of their colourful appearance, they are named after the ''Anemone'', a terrestrial flowering plant. Sea anemones are classifi ...
''
Anthopleura elegantissima The aggregating anemone (''Anthopleura elegantissima''), or clonal anemone, is the most abundant species of sea anemone found on rocky, tide swept shores along the Pacific coast of North America. This cnidarian hosts endosymbiotic algae called ...
'' reproduces clones of itself through a process of longitudinal
fission Fission, a splitting of something into two or more parts, may refer to: * Fission (biology), the division of a single entity into two or more parts and the regeneration of those parts into separate entities resembling the original * Nuclear fissio ...
, in which the animal splits into two parts along its length. The sea anemone ''
Anthopleura sola The starburst anemone or sunburst anemone (''Anthopleura sola'') is a species of sea anemone in the family Actiniidae. The sunburst anemone was formerly considered the solitary form of the common aggregating anemone, but was identified as a se ...
'' often engages in territorial fights. The white tentacles (acrorhagi), which contain stinging cells, are for fighting. The sea anemones sting each other repeatedly until one of them moves. Some species of sea stars can regenerate lost arms. Most species must retain an intact central part of the body to be able to regenerate, but a few can regrow from a single ray. The regeneration of these stars is possible because the vital organs are in the arms. Sea Urchins (" Echinoidia") move around tide pools with tube like feet. Different species of urchin have different colors, and many are seen in tide pools. With spines, some filled with poison like with " Toxopnesutes pileolus", that protect them from predators they feed almost undisturbed in tide pools.
Algae Algae (; singular alga ) is an informal term for a large and diverse group of photosynthetic eukaryotic organisms. It is a polyphyletic grouping that includes species from multiple distinct clades. Included organisms range from unicellular mic ...
and other microorganism are the food sources that attract them to the tide pools.


Flora

Sea palms ('' Postelsia'') look similar to miniature
palm trees Palm most commonly refers to: * Palm of the hand, the central region of the front of the hand * Palm plants, of family Arecaceae **List of Arecaceae genera * Several other plants known as "palm" Palm or Palms may also refer to: Music * Palm (ba ...
. They live in the middle to upper intertidal zones in areas with greater wave action. High wave action may increase nutrient availability and moves the blades of the thallus, allowing more sunlight to reach the organism so that it can photosynthesize. In addition, the constant wave action removes competitors, such as the mussel species ''
Mytilus californianus The California mussel (''Mytilus californianus'') is a large edible mussel, a marine bivalve mollusk in the family Mytilidae. This species is native to the west coast of North America, occurring from northern Mexico to the Aleutian Islands ...
''. Recent studies have shown that ''Postelsia'' grows in greater numbers when such competition exists; a control group with no competition produced fewer offspring than an experimental group with mussels; from this it is thought that the mussels provide protection for the developing gametophytes. Alternatively, the mussels may prevent the growth of competing
algae Algae (; singular alga ) is an informal term for a large and diverse group of photosynthetic eukaryotic organisms. It is a polyphyletic grouping that includes species from multiple distinct clades. Included organisms range from unicellular mic ...
such as '' Corallina'' or ''
Halosaccion ''Halosaccion'' is a genus of red algae belonging to the family Palmariaceae. The species of this genus are found in Northern Hemisphere. Species Species: *''Halosaccion americanum'' *''Halosaccion glandiforme'' *''Halosaccion hydrophora'' * ...
'', allowing ''Postelsia'' to grow freely after wave action has eliminated the mussels. Coralline algae "Corallinales" are predominant features of mid and low intertidal tide pools. Calcium carbonate (CaCO3) takes the form of calcite in their cell walls providing them with a hard outer shell. This shell protects from herbivores and
desiccation Desiccation () is the state of extreme dryness, or the process of extreme drying. A desiccant is a hygroscopic (attracts and holds water) substance that induces or sustains such a state in its local vicinity in a moderately sealed container. ...
due to lack of water and evaporation. Many forms of the Coralline algae bring herbivores, such as mollusks "Notoacmea", to the tide pools during high tides, increasing the biomass of the area. Once low tides comes, these herbivores are exposed to carnivores in the areas, fueling the food web. File:Anthopleura sola is consuming Velella velella.jpg, A large sea anemone ''(
Anthopleura sola The starburst anemone or sunburst anemone (''Anthopleura sola'') is a species of sea anemone in the family Actiniidae. The sunburst anemone was formerly considered the solitary form of the common aggregating anemone, but was identified as a se ...
)'' consuming a "by-the-wind-sailor" ''( Velella velella),'' a blue hydrozoan File:Postelsia palmaeformis 2.jpg, alt=Photo of speckled rocks, and various irregularly-shaped animals, ''
Postelsia palmaeformis ''Postelsia palmaeformis'', also known as the sea palm (not to be confused with the southern sea palm) or palm seaweed, is a species of kelp and classified within brown algae. It is the only known species in the genus Postelsia. The sea palm is ...
'' at low tide in a tide pool File:Starfishmussel.jpg, alt=Photo of five-legged approximately radially-symmetric animal lying on rock with shelled animal in its mouth, which is in the center of its body, Sea star, ''
Pisaster ochraceus ''Pisaster ochraceus'', generally known as the purple sea star, ochre sea star, or ochre starfish, is a common seastar found among the waters of the Pacific Ocean. Identified as a keystone species, ''P. ochraceus'' is considered an important indi ...
'' consuming a mussel in tide pools File:Close-up of clone war of sea anemones.jpg, Sea anemones, ''
Anthopleura sola The starburst anemone or sunburst anemone (''Anthopleura sola'') is a species of sea anemone in the family Actiniidae. The sunburst anemone was formerly considered the solitary form of the common aggregating anemone, but was identified as a se ...
'' engaged in a battle for territory


See also

*
Intertidal fish Coastal fish, also called inshore fish or neritic fish, inhabit the sea between the shoreline and the edge of the continental shelf. Since the continental shelf is usually less than deep, it follows that pelagic coastal fish are generally epi ...
* List of British Isles rockpool life * Rocky shore * Sydney inter-tidal rock pools


References


External links


Tidal swimming pools in Britain
{{Authority control Bodies of water Tides Marine biology Coastal geography Coastal and oceanic landforms