Neofelis nebulosa.jpg
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''Neofelis'' is a genus comprising two extant cat species in Southeast Asia: the
clouded leopard The clouded leopard (''Neofelis nebulosa''), also called the mainland clouded leopard, is a wild cat inhabiting dense forests from the foothills of the Himalayas through mainland Southeast Asia into South China. In the early 19th century, a cl ...
(''Neofelis nebulosa'') of mainland Asia, and the Sunda clouded leopard (''Neofelis diardi'') of
Sumatra Sumatra is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the sixth-largest island in the world at 473,481 km2 (182,812 mi.2), not including adjacent i ...
and Borneo The
scientific name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Neofelis'' is a composite of the Greek word ''neo-'' (νέος-) meaning 'young' and 'new', and the Latin word ''fēles'' meaning 'cat'.


Taxonomic history

The generic name ''Neofelis'' was first proposed by John Edward Gray in 1867 as comprising two species; ''Neofelis macrocelis'' occurring in the Himalaya,
Malacca Malacca ( ms, Melaka) is a state in Malaysia located in the southern region of the Malay Peninsula, next to the Strait of Malacca. Its capital is Malacca City, dubbed the Historic City, which has been listed as a UNESCO World Heritage Site si ...
, and Thailand, and ''Neofelis brachyurus'' occurring in the former
Formosa Taiwan, officially the Republic of China (ROC), is an island country located in East Asia. The main island of Taiwan, formerly known in the Western political circles, press and literature as Formosa, makes up 99% of the land area of the territorie ...
.Gray, J.E. (1867)
''Notes on the skulls of the Cats. 5. Neofelis''
Page 265–266 in: Proceedings of the Scientific Meetings of the Zoological Society of London for the year 1867.
Reginald Innes Pocock Reginald Innes Pocock F.R.S. (4 March 1863 – 9 August 1947) was a British zoologist. Pocock was born in Clifton, Bristol, the fourth son of Rev. Nicholas Pocock and Edith Prichard. He began showing interest in natural history at St. Edward ...
recognized the taxonomic classification of ''Neofelis'' in 1917, but admitted only the single species ''Neofelis nebulosa'' with several
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
and ''macrocelis'' as the type specimen.Pocock, R. I. (1917)
"The classification of existing Felidae"
''The Annals and Magazine of Natural History: Including Zoology, Botany, and Geology'', 8th ser. vol. 20 no. 119: 329–350.
For almost 90 years, the classification of ''Neofelis'' as a monotypic genus was widely accepted. In 2006, ''Neofelis diardi'' was found to be distinct from its continental relative ''Neofelis nebulosa'' and classified as a separate species.


Characteristics

Gray described the genus ''Neofelis'' as having an elongate skull, a broad and rather produced face on the same plane as the forehead, a large and elongate
nasal Nasal is an adjective referring to the nose, part of human or animal anatomy. It may also be shorthand for the following uses in combination: * With reference to the human nose: ** Nasal administration, a method of pharmaceutical drug delivery ** ...
, a moderate orbit, a truncated lower jaw and very long conical upper and lower canine teeth with a sharp cutting hinder edge. This skull has resemblances to that of the fossil '' Smilodon'', with very much elongated upper canines. As a result of this unusual skull anatomy, Neofelids have a maximum gape of approximately 90 degrees, the biggest of any extant Carnivoran, a trait shared by the extinct Machirodont subfamily. Pocock described the skull of ''Neofelis'' as recalling in general features that of '' Panthera pardus'', especially in the shortness and wide separation of the frontal and malar postorbital processes, relative proportion of mandibular teeth; but differing in the greater posterior width of the nasals, the thicker, more salient inferior edge of the orbit, and the mandible being greatly elevated anteriorly. The Sunda clouded leopard has longer upper canines and a narrower palate between them.


Distribution and habitat

''Neofelis'' species range from Nepal and Sikkim eastward to south
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
and Hainan, southeastward to
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John C. Wells, Joh ...
, Annam, the
Malay Peninsula The Malay Peninsula (Malay: ''Semenanjung Tanah Melayu'') is a peninsula in Mainland Southeast Asia. The landmass runs approximately north–south, and at its terminus, it is the southernmost point of the Asian continental mainland. The area ...
,
Sumatra Sumatra is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the sixth-largest island in the world at 473,481 km2 (182,812 mi.2), not including adjacent i ...
, Java and Borneo. They are most closely associated with primary evergreen tropical rainforest, but make use of other types of habitat. Sightings have also been made in
secondary Secondary may refer to: Science and nature * Secondary emission, of particles ** Secondary electrons, electrons generated as ionization products * The secondary winding, or the electrical or electronic circuit connected to the secondary winding i ...
and logged forest, as well as grassland and
scrub Scrub(s) may refer to: * Scrub, low shrub and grass characteristic of scrubland * Scrubs (clothing), worn by medical staff * ''Scrubs'' (TV series), an American television program * Scrubs (occupation), also called "scrub tech," "scrub nurse," ...
. In the Himalayan foothills they have been recorded up to .


Distribution of species

Between 1821 and 1862, several felids have been described from Southeast Asia that are subordinated under ''Neofelis'' today: *''Felis nebulosa'' was first described in 1821 by Edward Griffith based on a specimen brought from
Canton Canton may refer to: Administrative division terminology * Canton (administrative division), territorial/administrative division in some countries, notably Switzerland * Township (Canada), known as ''canton'' in Canadian French Arts and ent ...
in southern
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
. Populations range from the Himalayan foothills in Nepal through mainland Southeast Asia into China. *''Felis diardi'' was first described in 1823 by
Georges Cuvier Jean Léopold Nicolas Frédéric, Baron Cuvier (; 23 August 1769 – 13 May 1832), known as Georges Cuvier, was a French natural history, naturalist and zoology, zoologist, sometimes referred to as the "founding father of paleontology". Cuvier ...
based on a skin and a drawing received from Java. The Sunda clouded leopard is probably restricted to the islands of Sumatra and Borneo. In Java only clouded leopard fossils were found. *''Leopardus brachyurus'' was first described in 1862 by
Robert Swinhoe Robert Swinhoe FRS (1 September 1836 – 28 October 1877) was an English diplomat and naturalist who worked as a Consul in Formosa. He catalogued many Southeast Asian birds, and several, such as Swinhoe's pheasant, are named after him. Bio ...
based on two to three skins from Taiwan. Today the
Formosan clouded leopard The Formosan clouded leopard is an extinct clouded leopard (''Neofelis nebulosa'') population that was endemic to Taiwan. Camera trapping studies carried out in several protected areas in Taiwan between 1997 and 2012 did not record any clouded le ...
is considered a clouded leopard subspecies ''Neofelis nebulosa brachyurus''. It is now considered to be
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
.


Threats

Deforestation is the foremost threat for both ''Neofelis'' species. They are also threatened by commercial poaching for the wildlife trade. Skins, claws and teeth are offered for decoration and clothing, bones and meat as substitute for tiger in traditional Asian medicines and tonics, and live animals for the pet trade. Few poaching incidents have been documented, but all range states are believed to have some degree of commercial poaching. In recent years, substantial domestic markets existed in Indonesia,
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John C. Wells, Joh ...
and Vietnam.Nowell, K. (2007)
''Asian big cat conservation and trade control in selected range States: evaluating implementation and effectiveness of CITES Recommendations''
A TRAFFIC Report, June 2007


Conservation

Both ''Neofelis'' species are listed in CITES Appendix I and are protected over most of their range. Hunting is banned in Bangladesh, Brunei, Cambodia, China, India, Indonesia, Malaysia,
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John C. Wells, Joh ...
, Nepal, Taiwan, Thailand, and Vietnam. Hunting regulations apply in
Laos Laos (, ''Lāo'' )), officially the Lao People's Democratic Republic ( Lao: ສາທາລະນະລັດ ປະຊາທິປະໄຕ ປະຊາຊົນລາວ, French: République démocratique populaire lao), is a socialist ...
.


References

{{Taxonbar, from=Q269871 Mammals of Southeast Asia Pantherinae Mammal genera Taxa named by John Edward Gray