Great Egret (Casmerodius albus)- Non-breeding plumage in Kolkata W2 IMG 4341.jpg
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The great egret (''Ardea alba''), also known as the common egret, large egret, or (in the
Old World The "Old World" is a term for Afro-Eurasia that originated in Europe , after Europeans became aware of the existence of the Americas. It is used to contrast the continents of Africa, Europe, and Asia, which were previously thought of by the ...
) great white egret or great white heron is a large, widely distributed egret. The four subspecies are found in Asia, Africa, the Americas, and southern Europe. Recently it is also spreading to more northern areas of Europe. Distributed across most of the tropical and warmer temperate regions of the world, it builds tree nests in colonies close to water.


Taxonomy and systematics

Like all egrets, it is a member of the heron family, Ardeidae. Traditionally classified with the
stork Storks are large, long-legged, long-necked wading birds with long, stout bills. They belong to the family called Ciconiidae, and make up the order Ciconiiformes . Ciconiiformes previously included a number of other families, such as herons an ...
s in the Ciconiiformes, the Ardeidae are closer relatives of
pelican Pelicans (genus ''Pelecanus'') are a genus of large water birds that make up the family Pelecanidae. They are characterized by a long beak and a large throat pouch used for catching prey and draining water from the scooped-up contents before s ...
s and belong in the Pelecaniformes, instead. The great egret—unlike the typical egrets—does not belong to the genus ''
Egretta ''Egretta'' is a genus of medium-sized herons, mostly breeding in warmer climates. Representatives of this genus are found in most of the world, and the little egret, as well as being widespread throughout much of the Old World, has now started ...
'', but together with the great herons is today placed in '' Ardea''. In the past, however, it was sometimes placed in ''Egretta'' or separated in a monotypic genus ''Casmerodius''. The Old World population is often referred to as the "great white egret". This species is sometimes confused with the great white heron of the
Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean Se ...
, which is a white
morph Morph may refer to: Biology * Morph (zoology), a visual or behavioral difference between organisms of distinct populations in a species * Muller's morphs, a classification scheme for genetic mutations * "-morph", a suffix commonly used in tax ...
of the closely related
great blue heron The great blue heron (''Ardea herodias'') is a large wading bird in the heron family Ardeidae, common near the shores of open water and in wetlands over most of North America and Central America, as well as the Caribbean and the Galápagos ...
. The scientific name comes from Latin ''ardea'', "heron", and ''alba'', "white".


Subspecies

Four
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
are found in various parts of the world, which differ but little. Differences among them include bare-part coloration in the breeding season and size. The smallest subspecies, ''A. a. modesta'', is from Asia and Australasia and some taxonomists consider it to be a full species, the
eastern great egret The eastern great egret (''Ardea alba modesta''), a white heron in the genus '' Ardea'', is usually considered a subspecies of the great egret (''A. alba''). In New Zealand it is known as the white heron or by its Māori name ''kōtuku''. The sub ...
(''Ardea modesta''), but most scientists treat it as a subspecies. * ''A. a. alba'' Linnaeus, 1758nominate, found in Europe and across the
Palearctic The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa. The realm consists of several bioregions: the Euro-Sibe ...
. * ''A. a. egretta'' Gmelin, JF, 1789 – found in the
Americas The Americas, which are sometimes collectively called America, are a landmass comprising the totality of North and South America. The Americas make up most of the land in Earth's Western Hemisphere and comprise the New World. Along with th ...
* ''A. a. melanorhynchos'' Wagler, 1827 – found in Africa * '' A. a. modesta'' Gray, JE, 1831 – eastern great egret, found in India, Southeast Asia, East Asia and Oceania


Description

The great egret is a large heron with all-white
plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
. Standing up to tall, this species can measure in length with a wingspan of . Body mass can range from , with an average around . It is thus only slightly smaller than the great blue or grey heron (''A. cinerea''). Apart from size, the great egret can be distinguished from other white egrets by its yellow
bill Bill(s) may refer to: Common meanings * Banknote, paper cash (especially in the United States) * Bill (law), a proposed law put before a legislature * Invoice, commercial document issued by a seller to a buyer * Bill, a bird or animal's beak Plac ...
and black legs and feet, though the bill may become darker and the lower legs lighter in the breeding season. In breeding plumage, delicate ornamental feathers are borne on the back. Males and females are identical in appearance; juveniles look like nonbreeding adults. Differentiated from the intermediate egret (''Ardea intermedia'') by the gape, which extends well beyond the back of the eye in case of the great egret, but ends just behind the eye in case of the intermediate egret. It's flight is slow with its neck retracted. This is characteristic of herons and bitterns, and distinguishes them from
stork Storks are large, long-legged, long-necked wading birds with long, stout bills. They belong to the family called Ciconiidae, and make up the order Ciconiiformes . Ciconiiformes previously included a number of other families, such as herons an ...
s, cranes, ibises, and
spoonbill Spoonbills are a genus, ''Platalea'', of large, long-legged wading birds. The spoonbills have a global distribution, being found on every continent except Antarctica. The genus name ''Platalea'' derives from Ancient Greek and means "broad", refe ...
s, which extend their necks in flight. The great egret walks with its neck extended and wings held close. The great egret is not normally a vocal bird; it gives a low, hoarse croak when disturbed, and at breeding colonies, it often gives a loud croaking ''cuk cuk cuk'' and higher-pitched squawks. Owing to its wide distribution across so much of the Americas, as well as Africa, Europe and Asia, the great egret shares its habitat with many other similar species. For example, the little egret (''Egretta garzetta''), intermediate egret (''Ardea intermedia''), Chinese egret (''Egretta eulophotes''), and the
western reef heron The western reef heron (''Egretta gularis''), also called the western reef egret, is a medium-sized heron found in southern Europe, Africa and parts of Asia. It has a mainly coastal distribution and occurs in several plumage forms: a slaty-grey p ...
(''Egretta gularis''). In the Americas, the
snowy egret The snowy egret (''Egretta thula'') is a small white heron. The genus name comes from Provençal French for the little egret, , which is a diminutive of , 'heron'. The species name ''thula'' is the Araucano term for the black-necked swan, app ...
(''Egretta thula'')—a medium-sized heron that shares the same habitat as the great egret—is one such species. The snowy egret is readily distinguished from the great egret because it is noticeably smaller, and it has a more slender bill which is black in color and yellow feet, whereas the great egret has a yellow bill and black feet. Another species that—in North America—is easily confused with the great egret is the white
morph Morph may refer to: Biology * Morph (zoology), a visual or behavioral difference between organisms of distinct populations in a species * Muller's morphs, a classification scheme for genetic mutations * "-morph", a suffix commonly used in tax ...
of the
great blue heron The great blue heron (''Ardea herodias'') is a large wading bird in the heron family Ardeidae, common near the shores of open water and in wetlands over most of North America and Central America, as well as the Caribbean and the Galápagos ...
(''Ardea herodias''). The great blue heron is a bit larger, and has a thicker bill than that of the great egret.


Distribution and habitat

The great egret is generally a very successful species with a large and expanding range, occurring worldwide in temperate and tropical habitats. It is ubiquitous across the
Sun Belt The Sun Belt is a region of the United States generally considered to stretch across the Southeast and Southwest. Another rough definition of the region is the area south of the 36th parallel. Several climates can be found in the region — des ...
of the United States and in the Neotropics.


Conservation

In
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
, large numbers of great egrets were killed around the end of the 19th century so that their plumes, known as "
aigrette The term aigrette (; from the French for egret, or ''lesser white heron'') refers to the tufted crest or head-plumes of the egret, used for adorning a headdress. The word may also identify any similar ornament, in gems. History and description ...
s", could be used to decorate hats. Numbers have since recovered as a result of conservation measures. Its range has expanded as far north as southern Canada. However, in some parts of the southern United States, its numbers have declined due to habitat loss, particularly wetland degradation through drainage, grazing, clearing, burning, increased salinity, groundwater extraction and invasion by exotic plants. Nevertheless, the species adapts well to human habitation and can be readily seen near wetlands and bodies of water in urban and suburban areas. The great egret is partially migratory, with
northern hemisphere The Northern Hemisphere is the half of Earth that is north of the Equator. For other planets in the Solar System, north is defined as being in the same celestial hemisphere relative to the invariable plane of the solar system as Earth's Nort ...
birds moving south from areas with colder winters. It is one of the species to which the Agreement on the Conservation of African-Eurasian Migratory Waterbirds (AEWA) applies. In 1953, the great egret in flight was chosen as the symbol of the
National Audubon Society The National Audubon Society (Audubon; ) is an American non-profit environmental organization dedicated to conservation of birds and their habitats. Located in the United States and incorporated in 1905, Audubon is one of the oldest of such organ ...
, which was formed in part to prevent the killing of birds for their feathers. On 22 May 2012, a pair of great egrets was observed nesting in the UK for the first time at the Shapwick Heath nature reserve in Somerset. The species was a rare visitor to the UK and Ben Aviss of the BBC stated that the news could mean the UK's first great egret colony had become established. The following week, Kevin Anderson of Natural England confirmed a great egret chick had hatched, making it a new breeding bird record for the UK. In 2017, seven nests in Somerset fledged 17 young, and a second breeding site was announced at Holkham National Nature Reserve in Norfolk where a pair fledged three young. In January 2021, ''Bird Guides'', a UK website and magazine which reports sightings of rare birds, dropped the species from its list of nationally rare birds because sightings had become so numerous. A similar move northwards has been observed in the
Nordic countries The Nordic countries (also known as the Nordics or ''Norden''; literal translation, lit. 'the North') are a geographical and cultural region in Northern Europe and the Atlantic Ocean, North Atlantic. It includes the sovereign states of Denmar ...
where historically it was only a rare visitor. The first confirmed breeding in Sweden was 2012 and in Denmark was 2014. Both countries now have small colonies. In 2018, a pair of great egrets nested in Finland for the first time, raising four young in a grey heron colony in Porvoo.


Ecology

The species breeds in colonies in trees close to large lakes with
reed bed A reedbed or reed bed is a natural habitat found in floodplains, waterlogged depressions and estuaries. Reedbeds are part of a succession from young reeds colonising open water or wet ground through a gradation of increasingly dry ground. As ...
s or other extensive wetlands, preferably at height of . It begins to breed at 2–3 years of age by forming monogamous pairs each season. Whether the pairing carries over to the next season is not known. The male selects the nest area, starts a nest, and then attracts a female. The nest, made of sticks and lined with plant material, could be up to 3 feet across. Up to six bluish green eggs are laid at one time. Both sexes incubate the eggs, and the incubation period is 23–26 days. The young are fed by regurgitation by both parents and are able to fly within 6–7 weeks.


Diet

The great egret forages in shallow water or in drier habitats, feeding mainly on fish, frogs, other amphibians, small mammals (such as mice), and occasionally small reptiles (such as snakes), crustaceans (such as crayfish) and insects (such as crickets and grasshoppers). This species normally impales its prey with its long, sharp bill by standing still and allowing the prey to come within the striking distance of its bill, which it uses as a spear. It often waits motionless for prey or slowly stalks its victim.


Parasites

A long-running field study (1962–2013) suggested that the great egrets of central Europe host 17 different
helminth Parasitic worms, also known as helminths, are large macroparasites; adults can generally be seen with the naked eye. Many are intestinal worms that are soil-transmitted and infect the gastrointestinal tract. Other parasitic worms such as schi ...
species. Juvenile great egrets were shown to host fewer species, but the intensity of infection was higher in the juveniles than in the adults. Of the
digenea Digenea (Gr. ''Dis'' – double, ''Genos'' – race) is a class of trematodes in the Platyhelminthes phylum, consisting of parasitic flatworms (known as ''flukes'') with a syncytial tegument and, usually, two suckers, one ventral and one oral. ...
ns found in central European great egrets, numerous species likely infected their definitive hosts outside of central Europe itself.


In culture

The great egret is depicted on the
reverse Reverse or reversing may refer to: Arts and media * ''Reverse'' (Eldritch album), 2001 * ''Reverse'' (2009 film), a Polish comedy-drama film * ''Reverse'' (2019 film), an Iranian crime-drama film * ''Reverse'' (Morandi album), 2005 * ''Reverse'' ...
side of a 5- Brazilian reais banknote. The great egret is the symbol of the
National Audubon Society The National Audubon Society (Audubon; ) is an American non-profit environmental organization dedicated to conservation of birds and their habitats. Located in the United States and incorporated in 1905, Audubon is one of the oldest of such organ ...
. An airbrushed photograph of a great egret in breeding plumage by Werner Krutein is featured in the cover art of the 1992
Faith No More Faith No More is an American rock band from San Francisco, California, formed in 1979. Before settling on the current name in July 1983, the band performed under the names Sharp Young Men and later Faith No Man. Bassist Billy Gould, keyboardist/r ...
album '' Angel Dust''. In Belarus, a commemorative coin has the image of a great egret. The great egret also features on the New Zealand $2 coin and on the Hungarian 5-forint coin.


Gallery

File:Great Egret during mating season at Smith Oaks Sanctuary, High Island.jpg, Great egret during mating season at High Island File:Great Egret Fish.jpg, Adult in nonbreeding plumage File:Ardea alba-ibaraki-2016624.webm, (video) An individual of '' A. a. modesta'' flying off in Ibaraki,
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...
File:Ardea alba-head.jpg, Bright green facial skin during breeding season File:Ardea alba -chicks and nest -Morro Bay Heron Rookery -8.jpg, Parent on nest with chicks File:Great Egret - Centennial Lakes Park, Edina, Minnesota.jpg, Great egret at
Centennial Lakes Park Centennial Lakes Park is a park and man-made pond located in the southeast portion of Edina, Minnesota, United States. The park features more than of paved pathways meandering around a lake and interspersed with landscaped grounds, formal an ...
in Edina, Minnesota File:Guide leaflet (1901) (14579611617).jpg, Stuffed specimens at the
American Museum of Natural History The American Museum of Natural History (abbreviated as AMNH) is a natural history museum on the Upper West Side of Manhattan in New York City. In Theodore Roosevelt Park, across the street from Central Park, the museum complex comprises 26 inter ...
File:Great egret in GWC (43539).jpg, Plunging its head into the water to hunt inside Green-Wood Cemetery in Brooklyn File:Silberreiher 01 CH.jpg, Adult at
Piraqueaçu River The Piraquê Açu River is a river of Espírito Santo state in eastern Brazil. Part of the river basin is contained in the Augusto Ruschi Biological Reserve, a fully protected area. See also *List of rivers of Espírito Santo List of rivers i ...
in Santa Cruz in Brazil File:Great Egret with fish lake Coatepeque.jpg, Adult in flight over Lake Coatepeque, El Salvador File:Male Great Egret (ardea alba) 2.jpg, Adult male ''A. a. modesta in the
Tamar River Conservation Area The Tamar Conservation Area covers approximately 4,633 ha on the Tamar River estuary in Tasmania. It includes the Tamar Island Wetlands Reserve and a stretches through the upper part of the Tamar Estuary from St Leonards to the Batman Bridge ...
File:Большая белая цапля на Верхнем Знаменском пруду.jpg, Great egret in
Bitsa Park Bitsevski Park (russian: Битцевский парк), or Bitsa Park, is one of the largest natural parks (forests) in Moscow, Russia. The park, traversed by the Chertanovka River and the Bitsa River, sprawls for some 10 km from north to sou ...
, Moscow, Russia File:Great egret preening in a tree (84368).jpg, Preening in New York City File:Egrets Cormorants Trees Kabini Apr22 D72 23589.jpg, Sharing a perch with great cormorants (Phalacrocorax carbo),
Kabini The Kabini River is one of the major tributaries of the river Cauvery in southern India. It originates near Kavilumpara in Kozhikode district of Kerala state by the confluence of the Panamaram River and the Mananthavady River. It flows eastw ...
, India


See also

* Little egret * Intermediate egret


References


External links

*
Ageing and sexing (PDF) by Javier Blasco-Zumeta & Gerd-Michael Heinze

Great White Heron
– '' The Atlas of Southern African Birds''
Great White Egret
– National Park Neusiedlersee Seewinkel in Austria
Great Egret
– Cornell Lab of Ornithology

– USGS Patuxent Bird Identification InfoCenter * * * * {{Authority control great egret great egret Birds of Africa Birds of the Americas Birds of the Dominican Republic Birds of Eurasia Birds of Japan Fauna of the San Francisco Bay Area great egret Taxa named by Carl Linnaeus Cosmopolitan birds