Eyles' harrier
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Eyles's harrier (''Circus teauteensis'') is an
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
bird of prey Birds of prey or predatory birds, also known as raptors, are hypercarnivorous bird species that actively hunt and feed on other vertebrates (mainly mammals, reptiles and other smaller birds). In addition to speed and strength, these predat ...
which lived in
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island coun ...
. Its closest relative is the smaller Swamp harrier (''Circus approximans''), which arrived in New Zealand after its extinction.


Name

This species was named after
Jim Eyles James Roy Eyles (10 January 1926 – 12 November 2004) was a New Zealand archaeologist. Biography Born in Blenheim on 10 January 1926, Eyles was the son of Albert Roy Eyles and Daphne Eyles (née Daken). He spent his early years living with ...
, paleontologist and former director of the
Nelson Provincial Museum The Nelson Provincial Museum, ''Pupuri Taonga O Te Tai Ao'' is a regional museum in the city of Nelson, New Zealand. The museum showcases the Nelson and Tasman regions' history, from geological origins to the stories of individuals and families. N ...
and the West Coast Museum.


Taxonomy

Eyles' harrier was presumably somewhat similar to the living
spotted harrier The spotted harrier (''Circus assimilis'') also known as the smoke hawk, is a large Australasian bird of prey belonging to the family Accipitridae. Taxonomy The spotted harrier belongs to the family Accipitridae. Accipitridae is the largest fami ...
, its closest living relative, from which it diverged around 2.4 million years ago.


Description

It was an example of
island gigantism Island gigantism, or insular gigantism, is a biological phenomenon in which the size of an animal species isolated on an island increases dramatically in comparison to its mainland relatives. Island gigantism is one aspect of the more general Fos ...
, as an adult female weighed around , over twice as much as a swamp harrier. Its shape differed from that of most other harriers, and it was initially mistaken for a huge
hawk Hawks are birds of prey of the family Accipitridae. They are widely distributed and are found on all continents except Antarctica. * The subfamily Accipitrinae includes goshawks, sparrowhawks, sharp-shinned hawks and others. This subfa ...
, possibly a giant ''
Accipiter ''Accipiter'' is a genus of birds of prey in the family Accipitridae. With 51 recognized species it is the most diverse genus in its family. Most species are called goshawks or sparrowhawks, although almost all New World species (excepting th ...
''.


Ecology

It was a generalist predator, taking prey of the same size as small
eagle Eagle is the common name for many large birds of prey of the family Accipitridae. Eagles belong to several groups of genera, some of which are closely related. Most of the 68 species of eagle are from Eurasia and Africa. Outside this area, j ...
species do: land animals weighing one or a few
kilogram The kilogram (also kilogramme) is the unit of mass in the International System of Units (SI), having the unit symbol kg. It is a widely used measure in science, engineering and commerce worldwide, and is often simply called a kilo colloquially. ...
s. In its hunting strategy, however, it was more adapted to
avian Avian may refer to: * Birds or Aves, winged animals *Avian (given name) (russian: Авиа́н, link=no), a male forename Aviation *Avro Avian, a series of light aircraft made by Avro in the 1920s and 1930s *Avian Limited, a hang glider manufactur ...
prey, as mammals were entirely absent from New Zealand. Presumably, it hunted diurnal birds in a manner similar to
goshawk Goshawk may refer to several species of birds of prey, mainly in the genus ''Accipiter'': * Northern goshawk, ''Accipiter gentilis'', often referred to simply as the goshawk, since it is the only goshawk found in much of its range (in Europe and N ...
s.


Taxonomy

The
nomenclature Nomenclature (, ) is a system of names or terms, or the rules for forming these terms in a particular field of arts or sciences. The principles of naming vary from the relatively informal conventions of everyday speech to the internationally ag ...
and
taxonomy Taxonomy is the practice and science of categorization or classification. A taxonomy (or taxonomical classification) is a scheme of classification, especially a hierarchical classification, in which things are organized into groups or types. ...
of ''C. teauteensis'' are quite convoluted. As ''Circus eylesi'', it was only described in the mid-20th century, from remains found in
Pyramid Valley Pyramid Valley is a locality in the Hurunui District of New Zealand. It is well known for its prominent limestone rock formations. It is located near Waikari in the North Canterbury region, 80 km north-west of Christchurch. On the foot of the ...
in the South Island of New Zealand. However, essentially identical bones had been found about 100 years earlier by Augustus Hamilton and discussed subsequently by
Henry Ogg Forbes Henry Ogg Forbes LLD (30 January 1851 – 27 October 1932) was a Scottish explorer, ornithologist, and botanist. He also described a new species of spider, '' Thomisus decipiens''. Biography Forbes was the son of Rev Alexander Forbes M.A. (182 ...
; this material probably comes from the
Te Aute Te Aute College (Māori: Te Kura o Te Aute) is a school in the Hawke's Bay region of New Zealand. It opened in 1854 with twelve pupils under Samuel Williams, an Anglican missionary, and nephew and son-in-law of Bishop William Williams. It has a ...
region in the North Island. Forbes gave it the names ''Circus hamiltoni'' and ''Circus teauteensis'', but since it seems no
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...
was ever formally named nor can be deduced from the minimal description; these names are generally considered ''
nomina nuda In taxonomy, a ''nomen nudum'' ('naked name'; plural ''nomina nuda'') is a designation which looks exactly like a scientific name of an organism, and may have originally been intended to be one, but it has not been published with an adequate descr ...
''. ''C. teauteensis'' is considered valid, based on the argument that as with many New Zealand birds, it is reasonable to assume that North and South Island populations were at least distinct
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. Not all specie ...
, if not
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
. But even though Kálmán Lambrecht mentioned a putative holotype tibiotarsus (which still exists in the BMNH) he explicitly stated that Forbes' names were both invalid. In addition, harrier bones of comparatively recent age in the collection of Walter Mantell, assigned to ''C. gouldi'' (an obsolete name of ''C. approximans'') by
Richard Lydekker Richard Lydekker (; 25 July 1849 – 16 April 1915) was an English naturalist, geologist and writer of numerous books on natural history. Biography Richard Lydekker was born at Tavistock Square in London. His father was Gerard Wolfe Lydekker ...
, seem to be of a more robust bird judging from the published descriptions; this material may now be lost, but all things considered it is not unlikely to be the first remains of Eyles' harrier known to science.Worthy, T. H. and R. N. Holdaway. (2002): ''The lost world of the Moa: Prehistoric Life of New Zealand''.
Indiana University Indiana University (IU) is a system of public universities in the U.S. state of Indiana. Campuses Indiana University has two core campuses, five regional campuses, and two regional centers under the administration of IUPUI. *Indiana Universi ...
Press, Bloomington.


Extinction

The modern swamp harrier occurs all over New Zealand today. Even though the bones of its females can be mistaken for those of Eyles's harrier's males (as they differ little except in stoutness), all swamp harrier remains from the islands that have been studied in detail are some 1000 years old at most. That means that they post-date human settlement, i.e. there is no evidence for widespread coexistence of swamp and Eyles' harriers on New Zealand. Apparently, despite their considerable differences, the two harriers were still ecologically similar enough to competitively exclude one another, and only when the endemic Eyles' harrier became extinct could ''C. approximans'' become established.


Possible sightings

A noted explorer,
Charles Edward Douglas Charles Edward Douglas (1 July 1840 – 23 May 1916) was a New Zealand surveyor and explorer, who came to be known as Mr. Explorer Douglas, owing to his extensive explorations of the West Coast of New Zealand and his work for the New Zealand S ...
, claims in his journals that he had an encounter with two
raptor Raptor or RAPTOR may refer to: Animals The word "raptor" refers to several groups of bird-like dinosaurs which primarily capture and subdue/kill prey with their talons. * Raptor (bird) or bird of prey, a bird that primarily hunts and feeds on v ...
s of immense size in the
Landsborough River The Landsborough River is located in New Zealand's South Island. A major tributary of the Haast River, it flows southwest, parallel with the Southern Alps, for 50 kilometres from its source five kilometres north of Mount Hopkins to meet the Ha ...
valley (probably in the 1870s), and shot and ate them. These birds might have been a last remnant of the mighty '' pouakai'' (mythical represenation of
Haast's eagle Haast's eagle (''Hieraaetus moorei'') is an extinct species of eagle that once lived in the South Island of New Zealand, commonly accepted to be the pouakai of Māori legend.Wingspan Birds of Prey TrustZealand Birds Online
{{Taxonbar, from=Q760860
Eyles's harrier Eyles's harrier (''Circus teauteensis'') is an extinct bird of prey which lived in New Zealand. Its closest relative is the smaller Swamp harrier (''Circus approximans''), which arrived in New Zealand after its extinction. Name This species was ...
Extinct birds of New Zealand Bird extinctions since 1500 Late Quaternary prehistoric birds
Eyles's harrier Eyles's harrier (''Circus teauteensis'') is an extinct bird of prey which lived in New Zealand. Its closest relative is the smaller Swamp harrier (''Circus approximans''), which arrived in New Zealand after its extinction. Name This species was ...