European plaice
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The European plaice (''Pleuronectes platessa''), commonly referred to as simply plaice, is a species of marine
flatfish A flatfish is a member of the ray-finned demersal fish order Pleuronectiformes, also called the Heterosomata, sometimes classified as a suborder of Perciformes. In many species, both eyes lie on one side of the head, one or the other migrating ...
in the genus Pleuronectes of the family
Pleuronectidae Pleuronectidae, also known as righteye flounders, are a family of flounders. They are called "righteye flounders" because most species lie on the sea bottom on their left sides, with both eyes on their right sides. The Paralichthyidae are the op ...
.


Description

The European plaice is characterized, on their dorsal side, by their dark green to dark brown skin, blotched with conspicuous, but irregularly distributed, orange spots. The ventral side is pearly white. The skin is smooth with small scales. They are able to adapt their colour somewhat to match that of their surroundings, but the orange spots always remain visible. The skin lacks any prickles. The outline of adults is oval. The head is rather small and is less than 25% of the total length. The pointed mouth is terminal and fairly small with its
maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. T ...
reaching just below the right eye. Both eyes are located at the right side of the body. The bony ridge behind the eyes is another characteristic for this species. The
lateral line The lateral line, also called the lateral line organ (LLO), is a system of sensory organs found in fish, used to detect movement, vibration, and pressure gradients in the surrounding water. The sensory ability is achieved via modified epithelial ...
curves slightly above the
pectoral fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
. The dorsal fin reaches the eye. The dorsal and anal fins are distant from the caudal fin. The
anal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as see ...
contains 48 to 59 soft rays and is preceded by a spine. The
dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through c ...
has 65 to 79 soft rays, the pectoral fin 10 to 11, and the ventral fin six. European plaice can live up to 20 years and will reach its maximum size at about 50 to 70 cm but individuals up to 78 cm and 5.7 kg have been observed.


Distribution and habitat

European plaice is a common flatfish which inhabits sandy and muddy bottoms of the European shelf, usually at depths between 10 and 200 m. It is found from the
Barents Sea The Barents Sea ( , also ; no, Barentshavet, ; russian: Баренцево море, Barentsevo More) is a marginal sea of the Arctic Ocean, located off the northern coasts of Norway and Russia and divided between Norwegian and Russian terr ...
down to the
Iberian peninsula The Iberian Peninsula (), ** * Aragonese and Occitan: ''Peninsula Iberica'' ** ** * french: Péninsule Ibérique * mwl, Península Eibérica * eu, Iberiar penintsula also known as Iberia, is a peninsula in southwestern Europe, def ...
and around
Iceland Iceland ( is, Ísland; ) is a Nordic island country in the North Atlantic Ocean and in the Arctic Ocean. Iceland is the most sparsely populated country in Europe. Iceland's capital and largest city is Reykjavík, which (along with its ...
and the
Faroe Islands The Faroe Islands ( ), or simply the Faroes ( fo, Føroyar ; da, Færøerne ), are a North Atlantic island group and an autonomous territory of the Kingdom of Denmark. They are located north-northwest of Scotland, and about halfway bet ...
. It is often reported in ichthyological check lists for the
Mediterranean Sea The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on ...
but this is likely to be misidentified specimens of European flounder. Adult fish are generally found in deeper waters whereas young fish are found in shallow waters e.g. estuaries and sandy coasts. Plaice tend to burrow in the sediment during the day and remain stationary for long periods to avoid predators and ambush prey.


Diet

It is active at night and feeds on polychaetes,
crustaceans Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean g ...
and
bivalves Bivalvia (), in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of marine and freshwater molluscs that have laterally compressed bodies enclosed by a shell consisting of two hinged parts. As a group, bival ...
. Young plaice (between 1 and 2 years old) tend to consume mainly
shrimp Shrimp are crustaceans (a form of shellfish) with elongated bodies and a primarily swimming mode of locomotion – most commonly Caridea and Dendrobranchiata of the decapod order, although some crustaceans outside of this order are ref ...
s.


Lifecycle

The main spawning grounds in the North Sea are located in the Southern Bight and in the eastern
English Channel The English Channel, "The Sleeve"; nrf, la Maunche, "The Sleeve" ( Cotentinais) or ( Jèrriais), ( Guernésiais), "The Channel"; br, Mor Breizh, "Sea of Brittany"; cy, Môr Udd, "Lord's Sea"; kw, Mor Bretannek, "British Sea"; nl, Het Ka ...
. Plaice are determinate spawners in which fecundity is determined before the onset of spawning. Females mature, i.e. are able to spawn, at ages from 3 to 7 years. However, in the
North Sea The North Sea lies between Great Britain, Norway, Denmark, Germany, the Netherlands and Belgium. An epeiric sea on the European continental shelf, it connects to the Atlantic Ocean through the English Channel in the south and the Norwegian ...
, most females mature at 3 years.
Ovary The ovary is an organ in the female reproductive system that produces an ovum. When released, this travels down the fallopian tube into the uterus, where it may become fertilized by a sperm. There is an ovary () found on each side of the body. ...
development begins around late August to September with the spawning being from December to May. Each female releases eggs in batches every 3 to 5 days for around a month. The eggs hatch after about two weeks and drift passively in the
plankton Plankton are the diverse collection of organisms found in water (or air) that are unable to propel themselves against a current (or wind). The individual organisms constituting plankton are called plankters. In the ocean, they provide a cruc ...
. The
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. ...
e drift in the plankton and metamorphose after about 8 to 10 weeks, dependent on temperature, at which time they settle in the intertidal zone of sandy beaches. The larvae exhibit what is sometimes called semiactive tidal transport. As the larvae cannot swim against the prevailing currents, they make use of their ability to alter their vertical position in the water column to ensure they are transported to suitable habitat. On incoming or flood tides (water level is rising), the larvae move up into the water column and are thus transported towards land. On the outgoing or ebb tides (water level is falling), the larvae move down the water column and are not transported away from the intertidal by the tidal currents. When the larvae have reached a suitable site for settlement, the metamorphosis to the asymmetric body shape takes place. This can take up to 10 days. Recently transformed juveniles settle onto shallow intertidal beaches. The very youngest juveniles will, for a period of up to a week, strand themselves in very shallow pools on the intertidal once the tide has receded. The reasons for this behaviour are not clear. During the first year of life (when the fish are called 0+ group), the juveniles will stay in these shallow intertidal habitats for up to 7 months (depending on latitude and/or temperature), before migrating to deeper waters. Some of these fish will return the next year (when they are I+ group) and even fewer when they are II+ group; however, the majority of juveniles do not return after they have migrated during their first year.


Fisheries

European plaice are generally caught as part of a mixed flatfish fishery or in other
groundfish Demersal fish, also known as groundfish, live and feed on or near the bottom of seas or lakes (the demersal zone).Walrond Carl . "Coastal fish - Fish of the open sea floor"Te Ara - the Encyclopedia of New Zealand. Updated 2 March 2009 They oc ...
fisheries. The most common gears used in these fisheries are beam trawls, otter trawls and
Danish seine Seine fishing (or seine-haul fishing; ) is a method of fishing that employs a surrounding net, called a seine, that hangs vertically in the water with its bottom edge held down by weights and its top edge buoyed by floats. Seine nets can be de ...
. It is also caught in other fishing gears such as bottom set gillnets, longlines and hand lines. The majority of landings of European plaice are caught in the
North Sea The North Sea lies between Great Britain, Norway, Denmark, Germany, the Netherlands and Belgium. An epeiric sea on the European continental shelf, it connects to the Atlantic Ocean through the English Channel in the south and the Norwegian ...
. Total landings peaked in 1985-1990 when approximately 200 thousand tonnes were landed, current landings are generally less than 120 thousand tonnes. After a period of time in the 1980s and 1990s where landings exceeded management advice for several stocks, the fishing pressure has been reduced and the status of the stocks has improved and all the stocks are considered to be within safe biological limits as defined by ICES.


As a food

Plaice is sometimes used as the fish in
fish and chips Fish and chips is a popular hot dish consisting of fried fish in crispy batter, served with chips. The dish originated in England, where these two components had been introduced from separate immigrant cultures; it is not known who created ...
, in countries where the dish is popular.Seafish. On Plate. Fish & chips
In North German and
Danish cuisine Danish cuisine ( da, det danske køkken) originated from the peasant population's own local produce and was enhanced by cooking techniques developed in the late 19th century and the wider availability of goods during and after the Industrial Rev ...
, plaice is one of the most commonly eaten fishes. Filleted, battered, and pan-fried plaice is popular hot or cold as an open sandwich topping together with remoulade sauce and lemon slices. Battered plaice can also be served hot with
french fries French fries (North American English), chips (British English), finger chips (Indian English), french-fried potatoes, or simply fries, are '' batonnet'' or ''allumette''-cut deep-fried potatoes of disputed origin from Belgium and France. Th ...
and remoulade sauce as a main dish; this fish and chips variant is commonly available as a children's special in Danish restaurants. Breaded frozen plaice, ready to be baked or fried at home, is readily available in supermarkets. Fresh plaice is also oven-baked.


References

* Cooper, J.A. and F. Chapleau (1998). Monophyly and intrarelationships of the family Pleuronectidae (Pleuronectiformes), with a revised classification. ''Fish. Bull., U.S.'' 96(4):686-726. * * (1991). Changes in fecundity of female North Sea plaice (''Pleuronectes platessa'') between three periods since 1900. ''ICES Journal of Marine Science'' 48:253-280. * (1953). ''The plaice'' being the buckland lectures. Publisher Edward Arnold. * (2004). ''Flatfishes: Biology and Exploitation''. Blackwell Publishing.


External links

* Guide to Responsible Sourcing of Plaice - produced by Seafish https://web.archive.org/web/20071008001434/http://www.seafish.org/upload/file/fisheries_management/Plaice%20Factsheet2%20%20final.pdf
Information about plaice
* * {{Taxonbar, from=Q27098 Pleuronectes Commercial fish Fish of the North Sea Fish described in 1758 Taxa named by Carl Linnaeus