European pine marten
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The European pine marten (''Martes martes''), also known as the pine marten, is a
mustelid The Mustelidae (; from Latin ''mustela'', weasel) are a family of carnivorous mammals, including weasels, badgers, otters, ferrets, martens, minks and wolverines, among others. Mustelids () are a diverse group and form the largest family in t ...
native to and widespread in most of
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
,
Asia Minor Anatolia, tr, Anadolu Yarımadası), and the Anatolian plateau, also known as Asia Minor, is a large peninsula in Western Asia and the westernmost protrusion of the Asian continent. It constitutes the major part of modern-day Turkey. The re ...
, the
Caucasus The Caucasus () or Caucasia (), is a region between the Black Sea and the Caspian Sea, mainly comprising Armenia, Azerbaijan, Georgia, and parts of Southern Russia. The Caucasus Mountains, including the Greater Caucasus range, have historica ...
and parts of
Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...
,
Iraq Iraq,; ku, عێراق, translit=Êraq officially the Republic of Iraq, '; ku, کۆماری عێراق, translit=Komarî Êraq is a country in Western Asia. It is bordered by Turkey to Iraq–Turkey border, the north, Iran to Iran–Iraq ...
and
Syria Syria ( ar, سُورِيَا or سُورِيَة, translit=Sūriyā), officially the Syrian Arab Republic ( ar, الجمهورية العربية السورية, al-Jumhūrīyah al-ʻArabīyah as-Sūrīyah), is a Western Asian country loc ...
. It is listed as
Least Concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. ...
on the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biolo ...
. It is less commonly also known as baum marten, or sweet marten.


Description

The European pine marten's fur is usually light to dark brown. It is short and coarse in the summer, growing longer and silkier during the winter. It has a cream- to yellow-coloured "bib" marking on its throat. Its body is up to long, with a bushy tail of about . Males are slightly larger than females; typically, it weighs around . It has excellent senses of sight, smell, and hearing.


Distribution and habitat

The European pine marten inhabits well-wooded areas.


UK and The Republic of Ireland

In
Great Britain Great Britain is an island in the North Atlantic Ocean off the northwest coast of continental Europe. With an area of , it is the largest of the British Isles, the largest European island and the ninth-largest island in the world. It ...
, the species was for many years common only in northwestern
Scotland Scotland (, ) is a country that is part of the United Kingdom. Covering the northern third of the island of Great Britain, mainland Scotland has a border with England to the southeast and is otherwise surrounded by the Atlantic Ocean to ...
. A study in 2012 found that martens have spread from their
Scottish Highlands The Highlands ( sco, the Hielands; gd, a’ Ghàidhealtachd , 'the place of the Gaels') is a historical region of Scotland. Culturally, the Highlands and the Lowlands diverged from the Late Middle Ages into the modern period, when Lowland S ...
stronghold, north into
Sutherland Sutherland ( gd, Cataibh) is a historic county, registration county and lieutenancy area in the Highlands of Scotland. Its county town is Dornoch. Sutherland borders Caithness and Moray Firth to the east, Ross-shire and Cromartyshire (later c ...
and
Caithness Caithness ( gd, Gallaibh ; sco, Caitnes; non, Katanes) is a historic county, registration county and lieutenancy area of Scotland. Caithness has a land boundary with the historic county of Sutherland to the west and is otherwise bounded ...
and southeastwards from the
Great Glen The Great Glen ( gd, An Gleann Mòr ), also known as Glen Albyn (from the Gaelic "Glen of Scotland" ) or Glen More (from the Gaelic ), is a glen in Scotland running for from Inverness on the edge of Moray Firth, in an approximately straight ...
into
Moray Moray () gd, Moireibh or ') is one of the 32 local government council areas of Scotland. It lies in the north-east of the country, with a coastline on the Moray Firth, and borders the council areas of Aberdeenshire and Highland (council area), ...
,
Aberdeenshire Aberdeenshire ( sco, Aiberdeenshire; gd, Siorrachd Obar Dheathain) is one of the 32 council areas of Scotland. It takes its name from the County of Aberdeen which has substantially different boundaries. The Aberdeenshire Council area inclu ...
,
Perthshire Perthshire ( locally: ; gd, Siorrachd Pheairt), officially the County of Perth, is a historic county and registration county in central Scotland. Geographically it extends from Strathmore in the east, to the Pass of Drumochter in the nort ...
,
Tayside Tayside ( gd, Taobh Tatha) was one of the nine regions used for local government in Scotland from 15 May 1975 to 31 March 1996. The region was named for the River Tay. It was created by the Local Government (Scotland) Act 1973, following rec ...
, and
Stirlingshire Stirlingshire or the County of Stirling, gd, Siorrachd Sruighlea) is a historic county and registration countyRegisters of Scotland. Publications, leaflets, Land Register Counties. of Scotland. Its county town is Stirling. It borders Perth ...
, with some in the
Central Belt The Central Belt of Scotland is the area of highest population density within Scotland. Depending on the definition used, it has a population of between 2.4 and 4.2 million (the country's total was around 5.4 million in 2019), including Gre ...
, on the
Kintyre Kintyre ( gd, Cinn Tìre, ) is a peninsula in western Scotland, in the southwest of Argyll and Bute. The peninsula stretches about , from the Mull of Kintyre in the south to East and West Loch Tarbert in the north. The region immediately nor ...
and
Cowal Cowal ( gd, Còmhghall) is a peninsula in Argyll and Bute, in the west of Scotland, that extends into the Firth of Clyde. The northern part of the peninsula is covered by the Argyll Forest Park managed by Forestry and Land Scotland. The Arroch ...
peninsulas and on
Skye The Isle of Skye, or simply Skye (; gd, An t-Eilean Sgitheanach or ; sco, Isle o Skye), is the largest and northernmost of the major islands in the Inner Hebrides of Scotland. The island's peninsulas radiate from a mountainous hub dominated ...
and Mull. The expansion in the Galloway Forest has been limited compared with that in the core marten range. Martens were reintroduced to the
Glen Trool Glen Trool ( gd, Gleann an t-Sruthail) is a glen in the Southern Uplands, Galloway, Scotland. It contains Loch Trool Loch Trool is a narrow, freshwater loch in Galloway, in the Southern Uplands in south-west Scotland. It lies in an elevated p ...
Forest in the early 1980s and only restricted spread has occurred from there. This may be due to ongoing persecution and trapping by local gamekeepers. In
England England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe ...
, pine martens were long considered to be extinct. Analysis of a scat found at
Kidland Kidland is a former civil parish, now in the parish of Alwinton in Northumberland, England, about northwest of Alwinton village. In 1951 the parish had a population of 58. History From 1856 Kidland was a civil parish in its own right until ...
Forest in
Northumberland Northumberland () is a county in Northern England, one of two counties in England which border with Scotland. Notable landmarks in the county include Alnwick Castle, Bamburgh Castle, Hadrian's Wall and Hexham Abbey. It is bordered by land ...
in June 2010 may represent either a recolonisation from Scotland, or a
relict A relict is a surviving remnant of a natural phenomenon. Biology A relict (or relic) is an organism that at an earlier time was abundant in a large area but now occurs at only one or a few small areas. Geology and geomorphology In geology, a r ...
population that has escaped notice previously. There have been numerous reported sightings of pine martens in
Cumbria Cumbria ( ) is a ceremonial and non-metropolitan county in North West England, bordering Scotland. The county and Cumbria County Council, its local government, came into existence in 1974 after the passage of the Local Government Act 1972. ...
, however, it was not until 2011 that concrete proof—some scat that was DNA-tested—was found, followed in October 2022 by images being captured of a pine marten in Grizedale Forest. In July 2015, the first confirmed sighting of a pine marten in England for over a century was recorded by an amateur photographer in woodland in
Shropshire Shropshire (; alternatively Salop; abbreviated in print only as Shrops; demonym Salopian ) is a landlocked historic county in the West Midlands region of England. It is bordered by Wales to the west and the English counties of Cheshire to ...
. Sightings have continued in this area, and juveniles were recorded in 2019, indicating a breeding population. In July 2017, footage of a live pine marten was captured by a camera trap in the
North York Moors The North York Moors is an upland area in north-eastern Yorkshire, England. It contains one of the largest expanses of heather moorland in the United Kingdom. The area was designated as a National Park in 1952, through the National Parks and ...
in
Yorkshire Yorkshire ( ; abbreviated Yorks), formally known as the County of York, is a Historic counties of England, historic county in northern England and by far the largest in the United Kingdom. Because of its large area in comparison with other Eng ...
. In March 2018 the first ever footage of a pine marten in Northumberland was captured by the Back from the Brink pine marten project. In September 2022, the first pine marten to be seen in London in a century was spotted by the
Zoological Society of London The Zoological Society of London (ZSL) is a charity devoted to the worldwide conservation of animals and their habitats. It was founded in 1826. Since 1828, it has maintained the London Zoo, and since 1931 Whipsnade Park. History On 29 ...
's wildlife cameras as part of a hedgehog monitoring program. There is a small population of pine martens in
Wales Wales ( cy, Cymru ) is a country that is part of the United Kingdom. It is bordered by England to the east, the Irish Sea to the north and west, the Celtic Sea to the south west and the Bristol Channel to the south. It had a population in ...
. Scat found in Cwm Rheidol forest in 2007 was confirmed by DNA testing to be from a pine marten. A male was found in 2012 as road kill near
Newtown, Powys Newtown ( cy, Y Drenewydd) is a town in Powys, Wales. It lies on the River Severn in the community of Newtown and Llanllwchaiarn, within the historic boundaries of Montgomeryshire. It was designated a new town in 1967 and saw population growth ...
. This was the first confirmation in Wales of the species, living or dead, since 1971. The
Vincent Wildlife Trust Vincent Wildlife Trust (VWT) was founded in 1975 by the late Honourable Vincent Weir. It is a charity that focuses on mammal conservation in Britain and Ireland. Its Head Office is in Herefordshire, with local offices in south-west England, Wales ...
(VWT) has begun a reinforcement of these mammals in the
mid-Wales Mid Wales ( cy, Canolbarth Cymru or simply ''Y Canolbarth'', meaning "the midlands") or Central Wales refers to a region of Wales, encompassing its midlands, in-between North Wales and South Wales. The Mid Wales Regional Committee of the Se ...
area. During autumn 2015, 20 pine martens were captured in Scotland, in areas where a healthy pine marten population occurs, under licence from
Scottish Natural Heritage NatureScot ( gd, NàdarAlba), which was formerly known as Scottish Natural Heritage, is an executive non-departmental public body of the Scottish Government responsible for the country's natural heritage, especially its natural, genetic and s ...
. These animals were translocated and released in an area of mid-Wales. All of the martens were fitted with radio collars and are being tracked daily to monitor their movements and find out where they have set up territories. During autumn 2016, the VWT planned to capture and release another 20 pine martens in the hope of creating a self-sustaining population. The marten is still quite rare in
Ireland Ireland ( ; ga, Éire ; Ulster-Scots: ) is an island in the North Atlantic Ocean, in north-western Europe. It is separated from Great Britain to its east by the North Channel, the Irish Sea, and St George's Channel. Ireland is the s ...
, but the population is recovering and spreading; its traditional strongholds are in the west and south, especially
the Burren The Burren (; ) is a karst/ glaciokarst landscape centred in County Clare, on the west coast of Ireland.
Burr ...
and
Killarney National Park Killarney National Park ( ga, Páirc Náisiúnta Chill Airne), near the town of Killarney, County Kerry, was the first national park in Ireland, created when the Muckross Estate was donated to the Irish Free State in 1932. The park has sinc ...
, but the population in the Midlands has significantly increased in recent years. A study managed by academics at
Queen's University Belfast , mottoeng = For so much, what shall we give back? , top_free_label = , top_free = , top_free_label1 = , top_free1 = , top_free_label2 = , top_free2 = , established = , closed = , type = Public research university , parent = ...
, using cameras and citizen scientists, published in 2015, showed that pine martens were distributed across all
counties of Northern Ireland Northern Ireland is divided into six counties, namely: Antrim, Armagh, Down, Fermanagh, Londonderry and Tyrone. Six largely rural administrative counties based on these were among the eight primary local government areas of Northern Irela ...
.


Behaviour and ecology

Martens are the only
mustelid The Mustelidae (; from Latin ''mustela'', weasel) are a family of carnivorous mammals, including weasels, badgers, otters, ferrets, martens, minks and wolverines, among others. Mustelids () are a diverse group and form the largest family in t ...
s with semiretractable claws. This enables them to lead more
arboreal Arboreal locomotion is the locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally, but others are exclusively arboreal. The habitats pose nu ...
lifestyles, such as climbing or running on tree branches, although they are also relatively quick runners on the ground. They are mainly active at night and dusk. They have small, rounded, highly sensitive ears and sharp teeth adapted for eating small mammals, birds, insects, frogs, and
carrion Carrion () is the decaying flesh of dead animals, including human flesh. Overview Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures ...
. They have also been known to eat
berries A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit, although many pips or seeds may be present. Common examples are strawberries, ras ...
, birds' eggs, nuts, and honey. The European pine marten is
territorial A territory is an area of land, sea, or space, particularly belonging or connected to a country, person, or animal. In international politics, a territory is usually either the total area from which a state may extract power resources or an ...
and marks its
home range A home range is the area in which an animal lives and moves on a periodic basis. It is related to the concept of an animal's territory which is the area that is actively defended. The concept of a home range was introduced by W. H. Burt in 1943. He ...
by depositing
feces Feces ( or faeces), known colloquially and in slang as poo and poop, are the solid or semi-solid remains of food that was not digested in the small intestine, and has been broken down by bacteria in the large intestine. Feces contain a rela ...
. Its scat is black and twisted. The diet of the pine marten includes small mammals, carrion, birds, insects, and fruits. The recovery of the European pine marten in Ireland has been credited with reducing the population of invasive grey squirrels. Where the range of the expanding European pine marten population meets that of the grey squirrel, the population of the grey squirrels quickly retreats and the
red squirrel The red squirrel (''Sciurus vulgaris'') is a species of tree squirrel in the genus ''Sciurus'' common throughout Europe and Asia. The red squirrel is an arboreal, primarily herbivorous rodent. In Great Britain, Ireland, and in Italy numbe ...
population recovers. Because the grey squirrel spends more time on the ground than the red squirrel, which
co-evolve In biology, coevolution occurs when two or more species reciprocally affect each other's evolution through the process of natural selection. The term sometimes is used for two traits in the same species affecting each other's evolution, as well ...
d with the pine marten, they are thought to be far more likely to come in contact with this predator.


Lifespan

The European pine marten has lived to 18 years in captivity, but the maximum age recorded in the wild is only 11 years, with 3–4 years being more typical. They reach sexual maturity at 2–3 years of age.
Copulation Sexual intercourse (or coitus or copulation) is a sexual activity typically involving the insertion and thrusting of the penis into the vagina for sexual pleasure or reproduction.Sexual intercourse most commonly means penile–vaginal penetra ...
usually occurs on the ground and can last more than 1 hour. Mating occurs in July and August but the fertilized egg does not enter the uterus for about 7 months. The young are usually born in late March or early April after a month-long
gestation Gestation is the period of development during the carrying of an embryo, and later fetus, inside viviparous animals (the embryo develops within the parent). It is typical for mammals, but also occurs for some non-mammals. Mammals during pr ...
period that happens after the implantation of the fertilized egg, in litters of one to five. Young European pine martens weigh around at birth. The young begin to emerge from their dens around 7–8 weeks after birth and are able to disperse from the den around 12–16 weeks after their birth.


Threats

Although they are preyed upon occasionally by
golden eagle The golden eagle (''Aquila chrysaetos'') is a bird of prey living in the Northern Hemisphere. It is the most widely distributed species of eagle. Like all eagles, it belongs to the family Accipitridae. They are one of the best-known bird ...
s,
red fox The red fox (''Vulpes vulpes'') is the largest of the true foxes and one of the most widely distributed members of the Order (biology), order Carnivora, being present across the entire Northern Hemisphere including most of North America, Europe ...
es,
wolves The wolf (''Canis lupus''; : wolves), also known as the gray wolf or grey wolf, is a large canine native to Eurasia and North America. More than thirty subspecies of ''Canis lupus'' have been recognized, and gray wolves, as popularly un ...
, and
wildcat The wildcat is a species complex comprising two small wild cat species: the European wildcat (''Felis silvestris'') and the African wildcat (''F. lybica''). The European wildcat inhabits forests in Europe, Anatolia and the Caucasus, while th ...
s, humans are the largest threat to European pine martens. They are vulnerable from conflict with humans, arising from predator control for other species, or following predation of livestock and the use of inhabited buildings for denning. Martens may also be affected by woodland loss, which results in habitat loss for the animal. Persecution (illegal poisoning and shooting), loss of habitat leading to fragmentation, and other human disturbances have caused a considerable decline in the European pine marten population. They are also prized for their very fine fur in some areas. In the UK, European pine martens and their dens are offered full protection under the
Wildlife and Countryside Act 1981 The Wildlife and Countryside Act 1981 is an Act of Parliament in the United Kingdom implemented to comply with European Council Directive 79/409/EEC on the conservation of wild birds. In short, the act gives protection to native species (especia ...
and the Environmental Protection Act 1990.


References


Further reading

* {{Authority control Martens Mammals of Azerbaijan Mammals of Europe Mammals described in 1758 Taxa named by Carl Linnaeus