The European wild ass (''Equus hemionus hydruntinus'') or hydruntine is an extinct

equine Equinae is a subfamily of the family Equidae, which have lived worldwide (except Indonesia and Australia) from the Hemingfordian stage of the Early Miocene (16 million years ago) onwards. They are thought to be a monophyletic grouping.B. J. Ma ...

from the

Middle Pleistocene The Chibanian, widely known by its previous designation of Middle Pleistocene, is an age in the international geologic timescale or a stage in chronostratigraphy, being a division of the Pleistocene Epoch within the ongoing Quaternary Period. Th ...

to Late

Holocene The Holocene ( ) is the current geological epoch. It began approximately 11,650 cal years Before Present (), after the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene togeth ...

of Western Eurasia. It appeared first in the fossil record around 350–300,000 years before present. In the late Pleistocene it was widespread throughout much of western Eurasia from the Middle East to Europe, especially along the

Mediterranean The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on ...

, with fossil reports from

Sicily (man) it, Siciliana (woman) , population_note = , population_blank1_title = , population_blank1 = , demographics_type1 = Ethnicity , demographics1_footnotes = , demographi ...

, Turkey, Spain, France and Portugal. In the east the range apparently stretched at least to the Volga and to Iran. In the north it reached almost to the

North Sea The North Sea lies between Great Britain, Norway, Denmark, Germany, the Netherlands and Belgium. An epeiric sea on the European continental shelf, it connects to the Atlantic Ocean through the English Channel in the south and the Norwegian ...

in Germany and the

British Isles The British Isles are a group of islands in the North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner and Outer Hebrides, the Northern Isl ...

. Its range fragmented after the

Last Glacial Maximum The Last Glacial Maximum (LGM), also referred to as the Late Glacial Maximum, was the most recent time during the Last Glacial Period that ice sheets were at their greatest extent. Ice sheets covered much of Northern North America, Northern Eu ...

, surviving into the Holocene, its range gradually contracted further, persisting in small regions of

southern Europe Southern Europe is the southern region of Europe. It is also known as Mediterranean Europe, as its geography is essentially marked by the Mediterranean Sea. Definitions of Southern Europe include some or all of these countries and regions: Alb ...

into the

Bronze Age The Bronze Age is a historic period, lasting approximately from 3300 BC to 1200 BC, characterized by the use of bronze, the presence of writing in some areas, and other early features of urban civilization. The Bronze Age is the second pri ...

, and in Iran and the

Caucasus The Caucasus () or Caucasia (), is a region between the Black Sea and the Caspian Sea, mainly comprising Armenia, Azerbaijan, Georgia, and parts of Southern Russia. The Caucasus Mountains, including the Greater Caucasus range, have historica ...

into the

Iron Age The Iron Age is the final epoch of the three-age division of the prehistory and protohistory of humanity. It was preceded by the Stone Age ( Paleolithic, Mesolithic, Neolithic) and the Bronze Age ( Chalcolithic). The concept has been mostly ...

, around 500 BC. It has been suggested that the Iberian Zebro, extinct in the wild from the 16th century, could correspond to the ''Equus hydruntinus'', although the word " zebro" or "cebro" comes from Latin ''equiferus'' meaning 'wild horse'. Later research judged that it was unlikely that hydruntines persisted in the

Iberian Peninsula The Iberian Peninsula (), ** * Aragonese and Occitan: ''Peninsula Iberica'' ** ** * french: Péninsule Ibérique * mwl, Península Eibérica * eu, Iberiar penintsula also known as Iberia, is a peninsula in southwestern Europe, def ...

beyond the

Chalcolithic The Copper Age, also called the Chalcolithic (; from grc-gre, χαλκός ''khalkós'', "copper" and ''líthos'', "Rock (geology), stone") or (A)eneolithic (from Latin ''wikt:aeneus, aeneus'' "of copper"), is an list of archaeologi ...

. Morphologically the European ass can be distinguished from

asses Ass most commonly refers to: * Buttocks (in informal American English) * Donkey or ass, ''Equus africanus asinus'' **any other member of the subgenus ''Asinus'' Ass or ASS may also refer to: Art and entertainment * Ass (album), ''Ass'' (album ...

and hemiones particularly by its

molars The molars or molar teeth are large, flat teeth at the back of the mouth. They are more developed in mammals. They are used primarily to grind food during chewing. The name ''molar'' derives from Latin, ''molaris dens'', meaning "millstone to ...

and the relatively short

nare A nostril (or naris , plural ''nares'' ) is either of the two orifices of the nose. They enable the entry and exit of air and other gasses through the nasal cavities. In birds and mammals, they contain branched bones or cartilages called tur ...

s. The exact systematic position was formerly unclear but genetic and morphological analysis suggested that it is closely related to the Asiatic wild ass. A 2017 genetic study found that it was a subspecies of Asiatic wild ass, closer to the

Khur The Indian wild ass (''Equus hemionus khur''), also called the Indian onager or, in the local Gujarati language, Ghudkhur and Khur, is a subspecies of the onager native to South Asia. It is currently listed as Near Threatened by IUCN. The ...

than the

Persian onager The Persian onager (''Equus hemionus onager''), also called the Persian wild ass or Persian zebra, is a subspecies of onager (Asiatic wild ass) native to Iran (Persia). It is listed as Endangered, with no more than 600 individuals left in the wil ...

Etymology

The specific epithet, ''hydruntinus'', means ''from

Otranto Otranto (, , ; scn, label=Salentino, Oṭṛàntu; el, label= Griko, Δερεντό, Derentò; grc, Ὑδροῦς, translit=Hudroûs; la, Hydruntum) is a coastal town, port and ''comune'' in the province of Lecce (Apulia, Italy), in a ferti ...

'' ('' Hydruntum'' in

Latin Latin (, or , ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through ...

Ecology

The evidence shows that the European ass favoured semi-arid, steppic conditions and showed a preference for

temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout ...

climates, although it was also found in cool or cold conditions. It may have retreated to warmer locales during the coldest periods although the relatively short muzzle indicates an adaptation for cold conditions. It preferred open

biotopes A biotope is an area of uniform environmental conditions providing a living place for a specific assemblage of plants and animals. ''Biotope'' is almost synonymous with the term "habitat", which is more commonly used in English-speaking countrie ...

, between

shrubland Shrubland, scrubland, scrub, brush, or bush is a plant community characterized by vegetation dominated by shrubs, often also including grasses, herbs, and geophytes. Shrubland may either occur naturally or be the result of human activity. It ...

(favoured by true horses) and

grassland A grassland is an area where the vegetation is dominated by grasses ( Poaceae). However, sedge ( Cyperaceae) and rush ( Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur na ...

(favoured by bovids). It is believed to have shared this habitat with species such as the

woolly rhinoceros The woolly rhinoceros (''Coelodonta antiquitatis'') is an extinct species of rhinoceros that was common throughout Europe and Asia during the Pleistocene epoch and survived until the end of the last glacial period. The woolly rhinoceros was a me ...

. It is considered an ecologically important part of the ecosystem known as

mammoth steppe During the Last Glacial Maximum, the mammoth steppe, also known as steppe-tundra, was the Earth's most extensive biome. It spanned from Spain eastward across Eurasia to Canada and from the arctic islands southward to China. The mammoth step ...

where it filled a niche equivalent to that provided by the

African wild ass The African wild ass (''Equus africanus'') or African wild donkey is a wild member of the horse family, Equidae. This species is thought to be the ancestor of the domestic donkey (''Equus asinus''), which is sometimes placed within the same spec ...

Zebra Zebras (, ) (subgenus ''Hippotigris'') are African equines with distinctive black-and-white striped coats. There are three living species: the Grévy's zebra (''Equus grevyi''), plains zebra (''E. quagga''), and the mountain zebra (''E. zebr ...

in the African

Savanna A savanna or savannah is a mixed woodland- grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground ...

References

Onager Pleistocene mammals of Europe Extinct mammals of Europe Holocene extinctions Middle Pleistocene Late Pleistocene Extinct mammals Mammals described in 1907 {{horse-stub

Etymology

Ecology

See also

References