Eosuchians are an extinct

order Order, ORDER or Orders may refer to: * Categorization, the process in which ideas and objects are recognized, differentiated, and understood * Heterarchy, a system of organization wherein the elements have the potential to be ranked a number of ...

diapsid Diapsids ("two arches") are a clade of sauropsids, distinguished from more primitive eureptiles by the presence of two holes, known as temporal fenestrae, in each side of their skulls. The group first appeared about three hundred million years a ...

reptiles. Depending on which taxa are included the order may have ranged from the late

Carboniferous The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carboniferou ...

to the

Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', ...

but the consensus is that eosuchians are confined to the

Permian The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleo ...

and

Triassic The Triassic ( ) is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.36 Mya. The Triassic is the first and shortest per ...

. Eosuchia was initially defined to include all "thecodontian" reptiles which did not have an antorbital fenestra but did retain tabulars, postparietals and a large pineal foramen (Broom, 1914). Broom coined the term as a new suborder for ''

Youngina ''Youngina'' is an extinct genus of diapsid reptile from the Late Permian Beaufort Group ('' Tropidostoma''-''Dicynodon'' zones) of the Karoo Red Beds of South Africa. This, and a few related forms, make up the family Younginidae, within the Ord ...

''. A definition for inclusion in the order is difficult: it is almost easier to list the primitively-diapsid reptiles that have not been included at one time or another. The order has almost been treated as a dustbin for diapsids that are not obviously lepidosaurian or archosaurian. One consequence has been Romer's suggestion of the alternative order

Younginiformes Younginiformes is a replacement name for the taxon Eosuchia, proposed by Alfred Romer in 1947. The Eosuchia having become a wastebasket taxon for many probably distantly-related primitive diapsid reptiles ranging from the late Carboniferous to ...

to be applied strictly to those forms with the primitive diapsid form, in particular, a complete lowermost arch as the quadratojugal and jugal bones of the skull meet. The one constant eosuchian has been ''

'', a small lizard-shaped reptile from the Upper Permian of

South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring coun ...

. This and a couple of other genera make up the family

Younginidae Younginidae is an extinct family of neodiapsid reptiles from the Late Permian and Early Triassic. In a phylogenetic context, younginids are near the base of the clade Neodiapsida. Younginidae includes the species ''Youngina capensis'' from the La ...

. The tangasaurids, a family that includes forms apparently adapted for swimming in fresh water, is also usually included. In some phylogenies ''Eosuchia'' has been treated (probably erroneously) as a sister lepidosaur taxon to

Squamata Squamata (, Latin ''squamatus'', 'scaly, having scales') is the largest order of reptiles, comprising lizards, snakes, and amphisbaenians (worm lizards), which are collectively known as squamates or scaled reptiles. With over 10,900 species, ...

and

Rhynchocephalia Rhynchocephalia (; ) is an order of lizard-like reptiles that includes only one living species, the tuatara (''Sphenodon punctatus'') of New Zealand. Despite its current lack of diversity, during the Mesozoic rhynchocephalians were a diverse gr ...

.http://tolweb.org/Diapsida At one time or another all of the taxa in black in this phylogeny has been placed within "''Eosuchia''" in one sense or another

Classification

Eosuchia s.s. (Syn:

) :Family ''

incertae sedis ' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertain ...

'' ::'' Sphodrosaurus'' :Family: Galesphyridae ::'' Galesphyrus'' :Family:

::'' Heleosuchus'' ::''

'' :Family:

Tangasauridae Tangasauridae is a family of diapsids. Specimens have been found that are of Late Permian to Early Triassic in age from the Sakamena Group of western Madagascar. They lived alongside other taxa present from the Sakamena Group, including temnos ...

::''

Hovasaurus ''Hovasaurus'' is an extinct genus of diapsid reptile belonging to the order Eosuchia. It lived in what is now Madagascar during the Late Permian and Early Triassic, being a survivor of the Permian–Triassic extinction event and the paleontolog ...

'' ::''

Tangasaurus ''Tangasaurus'' is an extinct genus of aquatic basal tangasaurid neodiapsid known from the Late Permian period (late Changhsingian stage) of Tanga, northeastern Tanzania. It contains a single species, ''Tangasaurus mennelli''. Discovery ''Ta ...

'' ::'' Thadeosaurus'' ::'' Kenyasaurus''

References

Prehistoric reptile taxonomy Prehistoric animal orders Tetrapod orders Permian first appearances Lopingian taxonomic orders Early Triassic taxonomic orders Triassic extinctions {{paleo-reptile-stub