Eocursor
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''Eocursor'' (meaning "dawn runner") is
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of basal
ornithischia Ornithischia () is an extinct order of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name ''Ornithischia'', or "bird-hipped", reflects this similarity and is derived from the Greek ...
n
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
that lived in what is now
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring coun ...
during the
Early Jurassic The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic-J ...
. Remains of this animal have been found in the Upper Elliot Formation and it is among the most completely known early ornithischians, shedding new light on the origin of the group. The exact age of this taxon has been the subject of uncertainty. It was originally interprereted as living during the
Late Triassic The Late Triassic is the third and final epoch of the Triassic Period in the geologic time scale, spanning the time between Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch and followed by the Early Jurassic Epoch ...
(
Norian The Norian is a division of the Triassic Period. It has the rank of an age ( geochronology) or stage (chronostratigraphy). It lasted from ~227 to million years ago. It was preceded by the Carnian and succeeded by the Rhaetian. Stratigraphic ...
age), around 210
million years ago The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds. Usage Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago ...
; however,
Olsen Olsen or Ölsen may refer to: * Olsen (surname), people with the surname ''Olsen'' * Fred. Olsen & Co., a large shipping company with worldwide headquarters in Oslo, Norway * Ölsen, municipality in Rhineland-Palatinate, Germany. * Olsen House, a h ...
, Kent & Whiteside (2010) stated that there is no independent geochronological support for its assumed age, and the available data makes it impossible to conclusively determine whether ''Eocursor'' is of Triassic or
Early Jurassic The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic-J ...
(potentially as young as
Sinemurian In the geologic timescale, the Sinemurian is an age and stage in the Early or Lower Jurassic Epoch or Series. It spans the time between 199.3 ± 2 Ma and 190.8 ± 1.5 Ma (million years ago). The Sinemurian is preceded by the Hettangian and ...
) age. ''Eocursor'' was subsequently interpreted as a taxon of Early Jurassic age by McPhee ''et al.'' (2017), who identified the specimen as having originated from the Upper (not Lower) Elliot Formation. Fossils of ''Eocursor'' were originally collected in 1993, but were not formally described until fourteen years later. The
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specim ...
, ''Eocursor parvus'', was described in 2007 by Richard J. Butler, Roger M. H. Smith, and
David B. Norman David Bruce Norman (born 20 June 1952 in the United Kingdom) is a British paleontologist, currently the main curator of vertebrate paleontology at the Sedgwick Museum, Cambridge University. From 1991 to 2011, Norman has also been the Sedgwick Mu ...
. ''Eocursor'' was one of the earliest known ornithischians, and sheds some light on early dinosaur relationships because early dinosaurs are known from mostly incomplete skeletons. ''Eocursor'' is known from partial skeletal elements, including skull fragments, spinal elements, pelvis, long leg bones, and unusually large grasping hands.


Description

''Eocursor'' was a lightly built bipedal dinosaur with an estimated length of about 1 m (3 ft). The general aspect of the animal resembles that of the early Jurassic ornithischians such as '' Lesothosaurus'' and '' Scutellosaurus''. Its large hands resembled those of the
Heterodontosauridae Heterodontosauridae is a family of ornithischian dinosaurs that were likely among the most basal (primitive) members of the group. Their phylogenetic placement is uncertain but they are most commonly found to be primitive, outside of the group ...
, a
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English ter ...
of primitive ornithischians. The morphology of the triangular teeth, not unlike those of an iguana, suggests partial herbivory. The
tibia The tibia (; ), also known as the shinbone or shankbone, is the larger, stronger, and anterior (frontal) of the two bones in the leg below the knee in vertebrates (the other being the fibula, behind and to the outside of the tibia); it conn ...
was significantly longer than the
femur The femur (; ), or thigh bone, is the proximal bone of the hindlimb in tetrapod vertebrates. The head of the femur articulates with the acetabulum in the pelvic bone forming the hip joint, while the distal part of the femur articulates ...
, indicating it was a swift runner.


Discovery and naming

In 1993, the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...
specimen of ''Eocursor'' ( SAM-PK-K8025) was collected from the Upper Elliot Formation in Free State, South Africa. The skeleton, although only partially complete, is one of the best-preserved early ornithischian skeletons ever found. The fossils include a partial skeleton: portions of the skull, lower jaw, vertebrae, and limbs. In June 2007, the fossils were formally described by an international team of
paleontologists Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of foss ...
: Richard Butler of the
Natural History Museum A natural history museum or museum of natural history is a scientific institution with natural history collections that include current and historical records of animals, plants, fungi, ecosystems, geology, paleontology, climatology, and more. ...
, London, and the
University of Cambridge , mottoeng = Literal: From here, light and sacred draughts. Non literal: From this place, we gain enlightenment and precious knowledge. , established = , other_name = The Chancellor, Masters and Schola ...
, David Norman of the
University of Cambridge , mottoeng = Literal: From here, light and sacred draughts. Non literal: From this place, we gain enlightenment and precious knowledge. , established = , other_name = The Chancellor, Masters and Schola ...
, and Roger M. H. Smith of the Iziko South African Museum. The fossils were named ''Eocursor parvus'', from the
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
word ''eos'' (meaning "dawn"), and the
Latin Latin (, or , ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through ...
words ''cursor'' (meaning "runner") and ''parvus'' ("little"), "in reference to the early occurrence of this ornithischian, its apparent locomotory abilities and its small size."


Classification

''Eocursor'' was an early ornithischian, one of the first so-called "bird-hipped" dinosaurs, a group which would eventually give rise to animals such as ''
Stegosaurus ''Stegosaurus'' (; ) is a genus of herbivorous, four-legged, armored dinosaur from the Late Jurassic, characterized by the distinctive kite-shaped upright plates along their backs and spikes on their tails. Fossils of the genus have been fou ...
'', ''
Triceratops ''Triceratops'' ( ; ) is a genus of herbivorous chasmosaurine ceratopsid dinosaur that first appeared during the late Maastrichtian stage of the Late Cretaceous period, about 68 million years ago in what is now North America. It is one ...
'', and ''
Iguanodon ''Iguanodon'' ( ; meaning ' iguana-tooth'), named in 1825, is a genus of iguanodontian dinosaur. While many species have been classified in the genus ''Iguanodon'', dating from the late Jurassic Period to the early Cretaceous Period of Asia, ...
''. Butler ''et al.'' consider ''Eocursor'' more primitive than ''Lesothosaurus'' and Heterodontosauridae, but more derived than the ''
Pisanosaurus ''Pisanosaurus'' () is an extinct genus of early dinosauriformes, dinosauriform, likely an Ornithischia, ornithischian or Silesauridae, silesaurid, from the Late Triassic of Argentina. It was a small, lightly built, ground-dwelling herbivore, tha ...
'', as a basal ornithischian forming a sister
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English ter ...
of
Genasauria Genasauria is a clade of extinct beaked, primarily herbivorous dinosaurs. Paleontologist Paul Sereno first named Genasauria in 1986. The name Genasauria is derived from the Latin word ''gena'' meaning ‘cheek’ and the Greek word ''saúra'' (Ï ...
.


References


External links


''Eocursor parvus'' BBC article
{{Taxonbar , from=Q133498 Ornithischian genera Hettangian life Early Jurassic dinosaurs of Africa Jurassic South Africa Fossils of South Africa Fossil taxa described in 2007 Taxa named by David B. Norman