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Elachistinae
Elachistinae
(but see text)

The Elachistidae
Elachistidae
(grass-miner moths) are a family of small moths in the superfamily Gelechioidea. Some authors lump about 3,300 species in eight subfamilies here, but this arrangement almost certainly results in a massively paraphyletic and completely unnatural assemblage, united merely by symplesiomorphies retained from the first gelechioid moths. In fact, most of these moths appear to be either closer to the Oecophorinae
Oecophorinae
and are hence nowadays usually included in the Oecophoridae
Oecophoridae
(Depressariinae, "Deuterogoniinae", Hypertrophinae, Stenomatinae
Stenomatinae
and perhaps the enigmatic Aeolanthes), or constitute quite basal lineages of gelechioids, neither closely related to Elachista
Elachista
nor to Oecophora, and hence best treated as independent families within the Gelechioidea
Gelechioidea
(Agonoxenidae, Ethmiidae). The genus Coelopoeta is sometimes[1] still placed here, but probably belongs in the Oecophorinae. Consequently, the Elachistidae
Elachistidae
are essentially identical to the subfamily Elachistinae
Elachistinae
in the family's wide circumscription. The Agonoxenidae
Agonoxenidae
might perhaps belong here regardless, but even this is doubtful. Nonetheless, a considerable number of genera remain in the present family, and eventually it is likely that subdivisions will again be established (e.g. by raising some or all of the tribes proposed for the former Elachistinae
Elachistinae
to subfamily status). A significant reduction of genera has been proposed, from the current mass of 20 genera to 3. The proposition was made on the premise of several distinguishing classifications, all of which are synapomorphies. Those include genital size and presence of digitate, adult abdomen segments without dorsal spines, absence of maxillary palpi and fronto-clypeal suture, and immobile abdominal segments in pupae and larvae. Various tribes were considered within the proposition, with most differentiation coming from genital structure. In the modern, reduced description, the Elachistidae
Elachistidae
are small to very small moths (wingspans usually around 1 cm). Their wings appear feather-like due to the fine hair covering the wings' fringes, and the hindwings can be significantly reduced in area, essentially consisting of a small strip with a wide hairy fringe. The caterpillars are typically leaf miners or stem miners on Poales.

Contents

1 Genera

1.1 Fossil
Fossil
record

2 Former genera 3 Footnotes 4 References

Genera[edit] The genera of Elachistidae
Elachistidae
are:[2]

Elachistinae

Elachista Eretmograptis Meyrick, 1938 Mylocrita Meyrick, 1922 Myrrhinitis Meyrick, 1913 Perittia Stainton, 1854 Stephensia Stainton, 1858 Urodeta Stainton, 1869

Several small genera colored by some authors are here included in Elachista, as it would otherwise be liable to be non-monophyletic. As noted above, Aeolanthes may also belong here, as the only genus of a subfamily Aeolanthinae.

Parametriotinae Capuse, 1971 Agonoxeninae
Agonoxeninae
Meyrick, 1925 (alternatively treated as a separate family)

Fossil
Fossil
record[edit] Some prehistoric genera of Elachistidae, known only from fossils, have been described:[3]

Elachistites Kozlov, 1987 Microperittia Kozlov, 1987 Palaeoelachista Kozlov, 1987 Praemendesia Kozlov, 1987

Former genera[edit]

Annetennia Traugott-Olsen, 1995 Aristoptila Meyrick, 1932 Atmozostis
Atmozostis
Meyrick, 1932 Atrinia
Atrinia
Sinev, 1992 Austriana Traugott-Olsen, 1995 Atmozostis
Atmozostis
Meyrick, 1932 Bradleyana Traugott-Olsen, 1995 Calamograptis Meyrick, 1937 (Tineidae) Canariana Traugott-Olsen, 1995 Cryphioxena Meyrick, 1921 (Bucculatricidae) Dicasteris
Dicasteris
Meyrick, 1906 Dicranoctetes
Dicranoctetes
A.F.Braun, 1918 Elachistoides
Elachistoides
Sruoga, 1992 Eupneusta
Eupneusta
Bradley, 1974 Gibraltarensis Traugott-Olsen, 1996 Habeleria Traugott-Olsen, 1995 Holstia Traugott-Olsen, 1995 Illantis
Illantis
Meyrick, 1921 Kumia Falkovich, 1986

Kuznetzoviana Traugott-Olsen, 1996 Mendesina de Joannis, 1902 Microplitica Meyrick, 1935 Ogmograptis
Ogmograptis
Meyrick, 1935 (Bucculatricidae) Paraperittia
Paraperittia
Rebel, 1916 Perittoides Sinev, 1992 Petrochroa
Petrochroa
Busck, 1914 Phaneroctena A.J.Turner, 1923 (Cosmopterigidae) Phthinostoma Meyrick, 1914 Polymetis Walsingham, 1908 Proterochyta Meyrick, 1918 (Scythrididae) Sineviana Traugott-Olsen, 1995 Sruogania Traugott-Olsen, 1995 Symphoristis Meyrick, 1918 Whitebreadia Traugott-Olsen, 1995

Footnotes[edit]

^ See e.g. references in Savela (2004) ^ Wikispecies (2009-NOV-09), and see references in Savela (2004) ^ Wikispecies (2009-NOV-09)

References[edit]

Wikimedia Commons has media related to Elachistidae.

Data related to Elachistidae
Elachistidae
at Wikispecies See also Gelechioidea Talk
Talk
page for comparison of some approaches to gelechioid systematics and taxonomy.

Savela, Markku (2004): Markku Savela's Lepidoptera
Lepidoptera
and some other life forms – Elachistidae. Version of 2004-OCT-03. Retrieved 2010-APR-21. Kaila, Lauri (1999): Phylogeny and classification of the Elachistidae s.s. (Lepidoptera: Gelechioidea); Phylogeny and classification. Retrieved 2 November 2010.

v t e

Extant Lepidoptera
Lepidoptera
families

Kingdom: Animalia Phylum: Arthropoda Class: Insecta Subclass: Pterygota Infraclass: Neoptera Superorder: Endopterygota

Suborder Zeugloptera

Micropterigoidea

Micropterigidae
Micropterigidae
(mandibulate archaic moths)

Suborder Aglossata

Agathiphagoidea

Agathiphagidae (kauri moths)

Suborder Heterobathmiina

Heterobathmioidea

Heterobathmiidae

Suborder Glossata

Dacnonypha

Eriocranioidea

Eriocraniidae

Acanthoctesia

Acanthopteroctetoidea

Acanthopteroctetidae (archaic sun moths)

Lophocoronina

Lophocoronoidea

Lophocoronidae

Neopseustina

Neopseustoidea

Neopseustidae (archaic bell moths)

Exoporia

Hepialoidea

Anomosetidae Hepialidae
Hepialidae
(swift moths, ghost moths) Neotheoridae (Amazonian primitive ghost moths) Palaeosetidae (miniature ghost moths) Prototheoridae (African primitive ghost moths)

Mnesarchaeoidea

Mnesarchaeidae (New Zealand primitive moths)

H e t e r o n e u r a

M o n o t r y s i a

Incurvarioidea

Adelidae
Adelidae
(fairy longhorn moths) Cecidosidae Crinopterygidae Heliozelidae Incurvariidae Prodoxidae
Prodoxidae
(yucca moths)

Andesianoidea

Andesianidae (Andean endemic moths)

Nepticuloidea

Nepticulidae
Nepticulidae
(pigmy, or midget moths) Opostegidae
Opostegidae
(white eyecap moths)

Palaephatoidea

Palaephatidae (Gondwanaland moths)

Tischerioidea

Tischeriidae (trumpet leaf miner moths)

D i t r y s i a

Simaethistoidea

Simaethistidae

Tineoidea

Acrolophidae
Acrolophidae
(burrowing webworm moths) Arrhenophanidae Eriocottidae (Old World spiny-winged moths) Lypusidae Psychidae (bagworm moths) Tineidae
Tineidae
(fungus moths)

Gracillarioidea

Bucculatricidae
Bucculatricidae
(ribbed cocoon makers) Douglasiidae (Douglas moths) Gracillariidae Roeslerstammiidae

Yponomeutoidea

Acrolepiidae
Acrolepiidae
(false diamondback moths) Bedelliidae Glyphipterigidae
Glyphipterigidae
(sedge moths) Heliodinidae Lyonetiidae Plutellidae Yponomeutidae (ermine moths) Ypsolophidae

Gelechioidea

Autostichidae Batrachedridae Blastobasidae Coleophoridae
Coleophoridae
(case-bearers, case moths) Cosmopterigidae
Cosmopterigidae
(cosmet moths) Elachistidae
Elachistidae
(grass-miner moths) Gelechiidae
Gelechiidae
(twirler moths) Lecithoceridae
Lecithoceridae
(long-horned moths) Metachandidae Momphidae
Momphidae
(mompha moths) Oecophoridae
Oecophoridae
(concealer moths) Pterolonchidae Scythrididae
Scythrididae
(flower moths) Xyloryctidae
Xyloryctidae
(timber moths)

Galacticoidea

Galacticidae

Zygaenoidea

Heterogynidae Zygaenidae
Zygaenidae
(burnet, forester, or smoky moths) Himantopteridae Lacturidae Somabrachyidae Megalopygidae (flannel moths) Aididae Anomoeotidae Cyclotornidae Epipyropidae
Epipyropidae
(planthopper parasite moths) Dalceridae
Dalceridae
(slug caterpillars) Limacodidae
Limacodidae
(slug, or cup moths)

Cossoidea

Cossidae
Cossidae
(carpenter millers, or goat moths) Dudgeoneidae (dudgeon carpenter moths)

Sesioidea

Brachodidae (little bear moths) Castniidae
Castniidae
(castniid moths: giant butterfly-moths, sun moths) Sesiidae
Sesiidae
(clearwing moths)

Choreutoidea

Choreutidae
Choreutidae
(metalmark moths)

Tortricoidea

Tortricidae
Tortricidae
(tortrix moths)

Urodoidea

Urodidae
Urodidae
(false burnet moths)

Schreckensteinioidea

Schreckensteiniidae
Schreckensteiniidae
(bristle-legged moths)

Epermenioidea

Epermeniidae
Epermeniidae
(fringe-tufted moths)

Alucitoidea

Alucitidae (many-plumed moths) Tineodidae (false plume moths)

Pterophoroidea

Pterophoridae
Pterophoridae
(plume moths)

Whalleyanoidea

Whalleyanidae

Immoidea

Immidae

Copromorphoidea

Copromorphidae (tropical fruitworm moths) Carposinidae
Carposinidae
(fruitworm moths)

Hyblaeoidea

Hyblaeidae
Hyblaeidae
(teak moths)

Pyraloidea

Pyralidae
Pyralidae
(snout moths) Crambidae
Crambidae
(grass moth)

Thyridoidea

Thyrididae
Thyrididae
(picture-winged leaf moths)

Mimallonoidea

Mimallonidae (sack bearer moths)

Lasiocampoidea

Lasiocampidae
Lasiocampidae
(eggars, snout moths, or lappet moths)

Bombycoidea

Anthelidae
Anthelidae
(Australian lappet moth) Bombycidae
Bombycidae
(silk moths) Brahmaeidae
Brahmaeidae
(Brahmin moths) Carthaeidae (Dryandra moth) Endromidae
Endromidae
(Kentish glory and relatives) Eupterotidae Lemoniidae Saturniidae
Saturniidae
(saturniids) Sphingidae
Sphingidae
(hawk moths, sphinx moths and hornworms) Phiditiidae

Noctuoidea

Doidae Erebidae
Erebidae
(underwing, tiger, tussock, litter, snout, owlet moths) Euteliidae Noctuidae
Noctuidae
(daggers, sallows, owlet moths, quakers, cutworms, darts) Nolidae
Nolidae
(tuft moths) Notodontidae
Notodontidae
(prominents, kittens) Oenosandridae

Drepanoidea

Epicopeiidae
Epicopeiidae
(oriental swallowtail moths) Drepanidae
Drepanidae
(hook-tips)

Geometroidea

Sematuridae Uraniidae Geometridae (geometer moths)

Cimelioidea

Cimeliidae (gold moths)

Calliduloidea

Callidulidae
Callidulidae
(Old World butterfly-moths)

Superfamily unassigned

Millieriidae

Rhopalocera (butterflies)

Hedyloidea

Hedylidae
Hedylidae
(American moth-butterflies)

Hesperioidea

Hesperiidae (skippers)

Papilionoidea (true butterflies)

Lycaenidae
Lycaenidae
(gossamer-winged butterflies: blues, coppers and relatives) Nymphalidae
Nymphalidae
(brush-footed, or four-footed butterflies) Papilionidae (swallowtail butterflies) Pieridae
Pieridae
(whites, yellows, orangetips, sulphurs) Riodinidae
Riodinidae
(metalmarks)

Note: division Monotrysia
Monotrysia
is not a clade.

Taxonomy of the Lepidoptera Lists by region

Taxon identifiers

Wd: Q222549 ADW: Elachistidae BAMONA: Elachistidae BugGuide: 43958 EoL: 809 EPPO: 1ELACF Fauna Europaea: 1832 Fossilworks: 134749 GBIF: 7297 ITIS: 118721 NCBI: 26

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