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''Farancia erytrogramma'' (also known commonly as the rainbow snake, and less frequently as the eel moccasin) is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of large, nonvenomous, highly aquatic,
colubrid Colubridae (, commonly known as colubrids , from la, coluber, 'snake') is a family of snakes. With 249 genera, it is the largest snake family. The earliest species of the family date back to the Oligocene epoch. Colubrid snakes are found on ever ...
snake Snakes are elongated, Limbless vertebrate, limbless, carnivore, carnivorous reptiles of the suborder Serpentes . Like all other Squamata, squamates, snakes are ectothermic, amniote vertebrates covered in overlapping Scale (zoology), scales. Ma ...
, which is
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to coastal plains of the
southeastern United States The Southeastern United States, also referred to as the American Southeast or simply the Southeast, is a geographical region of the United States. It is located broadly on the eastern portion of the southern United States and the southern por ...
. Two
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
are recognized as being valid, one of which has been declared extinct.


Etymology

The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
, ''erytrogramma'', is derived from the
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
words ερυθρóς (''erythros''), meaning "red," and γράμμα (''gramma''), meaning "letter/word" but in this case probably the author means γραμμή (grammi) meaning "line/stripe".


Common names

Other common names for ''F. erytrogramma'' include horn snake, red-lined snake, red-lined horned snake, red-sided snake, sand hog, sand snake, and striped wampum.


Description

Dorsally Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position prov ...
, rainbow snakes have smooth, glossy bluish-black back scales, with three red stripes. They have short tails, with a spiny tip which they sometimes use as a probe. Adults may show yellow coloration along the sides and on the head. They usually grow to a total length (including tail) of 36-48 inches (91–122 cm), although some specimens have been recorded up to 66 inches (168 cm) in total length. Females are larger than males.


Behavior

Rainbow snakes are rarely seen due to their secretive habits. They spend most of their lives in the water, hiding in aquatic vegetation or other forms of cover. They are strong swimmers, and also know how to burrow into mud and sand. Rainbow snakes are not aggressive when captured, and do not bite their captors. In New Kent County,
Virginia Virginia, officially the Commonwealth of Virginia, is a state in the Mid-Atlantic and Southeastern regions of the United States, between the Atlantic Coast and the Appalachian Mountains. The geography and climate of the Commonwealth ar ...
, they are abundant in sandy fields near the
Chickahominy River The Chickahominy is an U.S. Geological Survey. National Hydrography Dataset high-resolution flowline dataThe National Map, accessed April 1, 2011 river in the eastern portion of the U.S. state of Virginia. The river, which serves as the eastern bo ...
, and great numbers are turned up by plows in the spring.


Diet

Rainbow snakes subsist mainly on
eel Eels are ray-finned fish belonging to the order Anguilliformes (), which consists of eight suborders, 19 families, 111 genera, and about 800 species. Eels undergo considerable development from the early larval stage to the eventual adult stage ...
s, but also prey on small
frog A frog is any member of a diverse and largely Carnivore, carnivorous group of short-bodied, tailless amphibians composing the order (biology), order Anura (ανοὐρά, literally ''without tail'' in Ancient Greek). The oldest fossil "proto-f ...
s,
tadpole A tadpole is the larval stage in the biological life cycle of an amphibian. Most tadpoles are fully aquatic, though some species of amphibians have tadpoles that are terrestrial. Tadpoles have some fish-like features that may not be found i ...
s, and
salamander Salamanders are a group of amphibians typically characterized by their lizard-like appearance, with slender bodies, blunt snouts, short limbs projecting at right angles to the body, and the presence of a tail in both larvae and adults. All ten ...
s. They eat their prey alive, usually swallowing them head first.


Reproduction

Adult female rainbow snakes usually lay their eggs in July, leaving them underground in sandy soil. A
clutch A clutch is a mechanical device that engages and disengages power transmission, especially from a drive shaft to a driven shaft. In the simplest application, clutches connect and disconnect two rotating shafts (drive shafts or line shafts). ...
consists of around 20 eggs on average, but large females may lay over 50. The young are hatched in late summer or fall.


Habitat

Rainbow snakes are found in aquatic habitats ranging from
cypress Cypress is a common name for various coniferous trees or shrubs of northern temperate regions that belong to the family Cupressaceae. The word ''cypress'' is derived from Old French ''cipres'', which was imported from Latin ''cypressus'', the ...
swamp A swamp is a forested wetland.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p. Swamps are considered to be transition zones because both land and water play a role in ...
s and
marsh A marsh is a wetland that is dominated by herbaceous rather than woody plant species.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p Marshes can often be found at ...
es to blackwater creeks, slow-moving streams, and sandy coastal plain.


Geographic range

''F. erytrogramma'' is found from southern
Maryland Maryland ( ) is a state in the Mid-Atlantic region of the United States. It shares borders with Virginia, West Virginia, and the District of Columbia to its south and west; Pennsylvania to its north; and Delaware and the Atlantic Ocean to ...
to southeastern
Louisiana Louisiana , group=pronunciation (French: ''La Louisiane'') is a state in the Deep South and South Central regions of the United States. It is the 20th-smallest by area and the 25th most populous of the 50 U.S. states. Louisiana is borde ...
, including eastern
Virginia Virginia, officially the Commonwealth of Virginia, is a state in the Mid-Atlantic and Southeastern regions of the United States, between the Atlantic Coast and the Appalachian Mountains. The geography and climate of the Commonwealth ar ...
, southeastern
North Carolina North Carolina () is a state in the Southeastern region of the United States. The state is the 28th largest and 9th-most populous of the United States. It is bordered by Virginia to the north, the Atlantic Ocean to the east, Georgia and So ...
,
South Carolina )''Animis opibusque parati'' ( for, , Latin, Prepared in mind and resources, links=no) , anthem = " Carolina";" South Carolina On My Mind" , Former = Province of South Carolina , seat = Columbia , LargestCity = Charleston , LargestMetro = ...
,
Georgia Georgia most commonly refers to: * Georgia (country), a country in the Caucasus region of Eurasia * Georgia (U.S. state), a state in the Southeast United States Georgia may also refer to: Places Historical states and entities * Related to the ...
, northern
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and to ...
,
Alabama (We dare defend our rights) , anthem = "Alabama (state song), Alabama" , image_map = Alabama in United States.svg , seat = Montgomery, Alabama, Montgomery , LargestCity = Huntsville, Alabama, Huntsville , LargestCounty = Baldwin County, Al ...
, and
Mississippi Mississippi () is a state in the Southeastern region of the United States, bordered to the north by Tennessee; to the east by Alabama; to the south by the Gulf of Mexico; to the southwest by Louisiana; and to the northwest by Arkansas. Miss ...
. A small population once inhabited the
Lake Okeechobee Lake Okeechobee (), also known as Florida's Inland Sea, is the largest freshwater lake in the U.S. state of Florida. It is the tenth largest natural freshwater lake among the 50 states of the United States and the second-largest natural freshwat ...
region in southern Florida, but was declared extinct on October 5, 2011.Curry, Tierra (2011). Center for Biological Diversity. BiologicalDiversity.com. October 5, 2011. One was seen at the Ocala National Forest, in Marion County, in early 2020 and the sighting was confirmed by the National Museum of Florida as the first in 50 years at this location.


Subspecies

There are two recognized
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
of ''F. erytrogramma'': *''Farancia erytrogramma erytrogramma'' (
Palisot de Beauvois Ambroise Marie François Joseph Palisot, Baron de Beauvois (27 July 1752, in Arras – 21 January 1820, in Paris) was a French naturalist and zoologist. Palisot collected insects in Oware, Benin, Saint Domingue, and the United States, from 1 ...
, 1802)
– common rainbow snake *''Farancia erytrogramma seminola'' (
Neill Neill is an Irish surname, and may refer to * A. S. Neill (1883-1973), British educator and author * Alec Neill (b.1950), New Zealand politician * Ben Neill (b.1957), American composer * Bob Neill (b.1952), British politician * Bud Neill (1911-197 ...
, 1964)
– southern Florida rainbow snake (declared extinct on October 5, 2011))


References


Further reading

* Baird SF, Girard C (1853). ''Catalogue of North American Reptiles in the Museum of the Smithsonian Institution. Part I.—Serpents.'' Washington, District of Columbia: Smithsonian Institution. xvi + 172 pp. (''Abastor erythrogrammus'', pp. 125–126). * Behler JL, King FW (1979). ''The Audubon Society Field Guide to North American Reptiles and Amphibians''. New York: Alfred A. Knopf. 743 pp. . (''Farancia erytrogramma'', pp. 610–611 + Plate 546). * Boulenger GA (1894). ''Catalogue of the Snakes in the British Museum (Natural History). Volume II., Containing the Conclusion of the Colubridæ Aglyphæ.'' London: Trustees of the British Museum (Natural History). (Taylor and Francis, printers). xi + 382 pp. + Plates I-XX. (''Abastor erythrogrammus'', p. 290). * Conant R, Bridges W (1939). ''What Snake is That? A Field Guide to the Snakes of the United States East of the Rocky Mountains''. (With 108 drawings by
Edmond Malnate Edmond is a given name related to Edmund. Persons named Edmond include: * Edmond Canaple (1797–1876), French politician * Edmond Chehade (born 1993), Lebanese footballer * Edmond Conn (1914–1998), American farmer, businessman, and politici ...
). New York and London: D. Appleton-Century. Frontispiece map + 163 pp. + Plates A-C, 1-32. (''Abastor erythrogrammus'', pp. 42–43 + Plate 2, Figure 6). * Gray JE (1849). ''Catalogue of the Specimens of Snakes in the Collection of the British Museum''. London: Trustees of the British Museum. (Edward Newman, printer). xv + 125 pp. (''Abastor erythrogrammus'', p. 78). *
Jan G Jan, JaN or JAN may refer to: Acronyms * Jackson, Mississippi (Amtrak station), US, Amtrak station code JAN * Jackson-Evers International Airport, Mississippi, US, IATA code * Jabhat al-Nusra (JaN), a Syrian militant group * Japanese Article Numb ...
, Sordelli F (1868). ''Iconographie générale des Ophidiens, Vingt-neuvième livraison''. Paris: Baillière. Index + Plates I-VI. (''Calopisma erythrogrammum'', Plate IV, Figure 2; Plate V, Figure 1). (in
French French (french: français(e), link=no) may refer to: * Something of, from, or related to France ** French language, which originated in France, and its various dialects and accents ** French people, a nation and ethnic group identified with Franc ...
). * Morris PA (1948). ''Boy's Book of Snakes: How to Recognize and Understand Them''. A volume of the Humanizing Science Series, edited by
Jacques Cattell Jaques (Jack) Cattell (2 June 1904 in Garrison, New York – 19 December 1961) was an American publisher and founder of a company bearing his name, "Jaques Cattell Press, Inc.," based in Lancaster, Pennsylvania. Jaques Cattell Press, Inc. The Sci ...
. New York: Ronald Press. viii + 185 pp. (''Abastor erythrogrammus'', pp. 88–89, 179). * Neill WT (1964). "Taxonomy, natural history, and zoogeography of the rainbow snake, ''Farancia erytrogramma'' (Palisot de Beauvois)". ''American Midland Naturalist'' 71: 257–295. * Palisot de Beauvois MFJ(1802). ''In'': Sonnini CS, Latreille PA (1802). ''Histoire Naturelle des Reptiles, avec figures dessinées d'après nature. Tome IV. Seconde Partie. Serpens.'' Paris: Deterville. (Crapelet, printer). 410 pp. (''Coluber erytrogrammus'', new species, pp. 141-142). (in French). * Powell R, Conant R, Collins JT (2016). ''Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America, Fourth Edition''. Boston and New York: Houghton Mifflin Harcourt. xiv + 494 pp. . (''Farancia erytrogramma'', p. 406 + Plate 39). * Richmond ND (1954). "Variation and Sexual Dimorphism in Hatchlings of the Rainbow Snake, ''Abastor erythrogrammus'' ". ''Copeia'' 1954 (2): 87–92. * Schmidt KP, Davis DD (1941). ''Field Book of Snakes of the United States and Canada''. New York: G.P. Putnam's Sons. 365 pp. ''Abastor erythrogrammus'', pp. 104–105, Figure 21 + Frontispiece (colored plate). * Smith HM, Brodie ED Jr (1982). ''Reptiles of North America: A Guide to Field Identification''. New York: Golden Press. 240 pp. . (''Farancia erytrogramma'', pp. 162–163). * Steen, David A.; Stevenson, Dirk J.; Beane, Jeffery C.; Willson, John D.; Aresco, Matthew J.; Godwin, James C.; Graham, Sean P.; Smith, Lora J.; Howse, Jennifer M.; Rudolph, J. Craig; Pierce, Josh B.; Lee, James R.; Gregory, Beau B.; Jensen, John; Stile, Sierra H. (2013). "Terrestrial movements of the red-bellied mudsnake (''Farancia abacura'') and rainbow snake (''F. erytrogramma'')". ''Herpetological Review'' 44: 208–213. * Stejneger L, Barbour T (1917). ''A Check List of North American Amphibians and Reptiles''. Cambridge, Massachusetts: Harvard University Press. 125 pp. (''Abastor erythrogrammus'', p. 75). * Zim HS, Smith HM (1956). ''Reptiles and Amphibians: A Guide to Familiar American Species: A Golden Nature Guide''. Revised edition. New York: Simon and Schuster. 160 pp. (''Abastor erythrogrammus'', pp. 74, 156).


External links

*
University of Georgia Savannah River Ecology LaboratoryVirginia Department of Game and Inland Fisheries
{{DEFAULTSORT:Farancia Erytrogramma Colubrids Reptiles described in 1802 Fauna of the Southeastern United States Reptiles of the United States