Eastern Afromontane
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The Afromontane regions are subregions of the Afrotropical realm, one of the Earth's eight
biogeographic realm A biogeographic realm or ecozone is the broadest biogeographic division of Earth's land surface, based on distributional patterns of terrestrial organisms. They are subdivided into bioregions, which are further subdivided into ecoregions. De ...
s, covering the plant and animal species found in the mountains of
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
and the southern Arabian Peninsula. The Afromontane regions of Africa are discontinuous, separated from each other by lower-lying areas, and are sometimes referred to as the Afromontane archipelago, as their distribution is analogous to a series of
sky island Sky islands are isolated mountains surrounded by radically different lowland environments. The term originally referred to those found on the Mexican Plateau, and has extended to similarly isolated high-elevation forests. The isolation has s ...
s.


Geography

Afromontane communities occur above elevation near the equator, and as low as elevation in the Knysna-Amatole montane forests of
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. It is bounded to the south by of coastline that stretch along the Atlantic Ocean, South Atlantic and Indian Oceans; to the ...
. Afromontane forests are generally cooler and more humid than the surrounding lowlands. The Afromontane archipelago mostly follows the East African Rift from the
Red Sea The Red Sea ( ar, البحر الأحمر - بحر القلزم, translit=Modern: al-Baḥr al-ʾAḥmar, Medieval: Baḥr al-Qulzum; or ; Coptic: ⲫⲓⲟⲙ ⲛ̀ϩⲁϩ ''Phiom Enhah'' or ⲫⲓⲟⲙ ⲛ̀ϣⲁⲣⲓ ''Phiom ǹšari''; ...
to
Zimbabwe Zimbabwe (), officially the Republic of Zimbabwe, is a landlocked country located in Southeast Africa, between the Zambezi and Limpopo Rivers, bordered by South Africa to the south, Botswana to the south-west, Zambia to the north, and ...
, with the largest areas in the
Ethiopian Highlands The Ethiopian Highlands is a rugged mass of mountains in Ethiopia in Northeast Africa. It forms the largest continuous area of its elevation in the continent, with little of its surface falling below , while the summits reach heights of up to . ...
, the
Albertine Rift Mountains The Albertine Rift is the western branch of the East African Rift, covering parts of Uganda, the Democratic Republic of the Congo (DRC), Rwanda, Burundi and Tanzania. It extends from the northern end of Lake Albert to the southern end of Lake ...
of
Uganda }), is a landlocked country in East Africa. The country is bordered to the east by Kenya, to the north by South Sudan, to the west by the Democratic Republic of the Congo, to the south-west by Rwanda, and to the south by Tanzania. The sou ...
, Rwanda, Burundi,
Democratic Republic of the Congo The Democratic Republic of the Congo (french: République démocratique du Congo (RDC), colloquially "La RDC" ), informally Congo-Kinshasa, DR Congo, the DRC, the DROC, or the Congo, and formerly and also colloquially Zaire, is a country in ...
, and
Tanzania Tanzania (; ), officially the United Republic of Tanzania ( sw, Jamhuri ya Muungano wa Tanzania), is a country in East Africa within the African Great Lakes region. It borders Uganda to the north; Kenya to the northeast; Comoro Islands ...
, and the
Eastern Arc The Eastern Academic Research Consortium, or "Eastern Arc", is a regional research collaboration between the University of East Anglia, the University of Essex, and the University of Kent. The three partner institutions are all part of the plate ...
highlands of
Kenya ) , national_anthem = " Ee Mungu Nguvu Yetu"() , image_map = , map_caption = , image_map2 = , capital = Nairobi , coordinates = , largest_city = Nairobi ...
and Tanzania. Other Afromontane regions include the
Drakensberg The Drakensberg (Afrikaans: Drakensberge, Zulu: uKhahlambha, Sotho: Maluti) is the eastern portion of the Great Escarpment, which encloses the central Southern African plateau. The Great Escarpment reaches its greatest elevation – within t ...
range of southern Africa, the
Cameroon Highlands The Western High Plateau, Western Highlands or Bamenda Grassfields is a region of Cameroon characterised by high relief, cool temperatures, heavy rainfall and savanna vegetation. The region lies along the Cameroon line and consists of mounta ...
, and the Cameroon Line volcanoes, including
Mount Cameroon Mount Cameroon is an active volcano in the South West region of Cameroon next to the city of Buea near the Gulf of Guinea. Mount Cameroon is also known as Cameroon Mountain or Fako (the name of the higher of its two peaks) or by its indigenous n ...
,
Bioko Bioko (; historically Fernando Po; bvb, Ëtulá Ëria) is an island off the west coast of Africa and the northernmost part of Equatorial Guinea. Its population was 335,048 at the 2015 census and it covers an area of . The island is located of ...
, and
São Tomé São Tomé is the capital and largest city of the Central African island country of São Tomé and Príncipe. Its name is Portuguese for " Saint Thomas". Founded in the 15th century, it is one of Africa's oldest colonial cities. History Álvar ...
.


Flora

Although some Afromontane enclaves are widely separated, they share a similar mix of plant species which are often distinct from the surrounding lowland regions.
Podocarp Podocarpaceae is a large family of mainly Southern Hemisphere conifers, known in English as podocarps, comprising about 156 species of evergreen trees and shrubs.James E. Eckenwalder. 2009. ''Conifers of the World''. Portland, Oregon: Timber Pr ...
s, of genera ''
Podocarpus ''Podocarpus'' () is a genus of conifers, the most numerous and widely distributed of the podocarp family, the Podocarpaceae. The name comes from Greek πούς (poús, “foot”) + καρπός (karpós, “fruit”). ''Podocarpus'' species ...
'' and '' Afrocarpus'', are a characteristic tree, along with ''
Prunus africana ''Prunus africana'', the African cherry, has a wide distribution in Africa, occurring in montane regions of central and southern Africa and on the islands of Bioko, São-Tomé, Grande Comore, and Madagascar. It can be found at above sea level. I ...
'', ''
Hagenia abyssinica ''Hagenia'' is a monotypic genus of flowering plant with the sole species ''Hagenia abyssinica'', native to the high-elevation Afromontane regions of central and eastern Africa. It also has a disjunct distribution in the high mountains of East ...
'', ''
Juniperus procera ''Juniperus procera'' (known by the common English names African juniper, African pencil-cedar, East African juniper, East African-cedar, and Kenya-cedar) is a coniferous tree native to mountainous areas in Africa and the Arabian Peninsula. It is ...
'', and ''
Olea ''Olea'' ( ) is a genus of about 40 species in the family Oleaceae, native to warm temperate and tropical regions of the Middle East, southern Europe, Africa, southern Asia, and Australasia. They are evergreen trees and shrubs, with small ...
spp.''. In the higher mountains, the Afromontane forest or woodland zone transitions to a higher Afroalpine zone of grasslands, shrublands, or moorlands. The plant families
Curtisiaceae ''Curtisia dentata'' (commonly known as the Assegai tree or Cape lancewood, af, Assegaai, xh, Umgxina, zu, Umagunda) is a flowering tree from Southern Africa. It is the sole species in genus ''Curtisia'', which was originally classed as a typ ...
and Oliniaceae are Afromontane endemics and family Barbeyaceae is a near-endemic. The tree genera '' Afrocrania, Balthasaria,
Curtisia ''Curtisia dentata'' (commonly known as the Assegai tree or Cape lancewood, af, Assegaai, xh, Umgxina, zu, Umagunda) is a flowering tree from Southern Africa. It is the sole species in genus ''Curtisia'', which was originally classed as a typ ...
, Ficalhoa,
Hagenia ''Hagenia'' is a monotypic genus of flowering plant with the sole species ''Hagenia abyssinica'', native to the high-elevation Afromontane regions of central and eastern Africa. It also has a disjunct distribution in the high mountains of East ...
, Kiggelaria, Leucosidea, Platypterocarpus,
Trichocladus ''Trichocladus'' is a genus of plant in family Hamamelidaceae, consisting of shrubs or small trees. The distinguishing features of the genus ''Trichocladus'' are as follows: *Branches and leaves are often covered in dense, velvet-like, stellate h ...
,'' ''
Widdringtonia ''Widdringtonia'' is a genus of coniferous trees in the Cupressaceae (cypress family). The name was Austrian botanist Stephan Endlicher's way of honouring an early expert on the coniferous forests of Spain, Capt. Samuel Edward Widdrington (1787 ...
'', and '' Xymalos'' are Afromontane endemics or near-endemics, as are the plant genera '' Ardisiandra, Cincinnobotrys,'' and '' Stapfiella''.


Plant communities

Afromontane areas have a wide range of plant communities, including intermediate types. These include: * Afromontane rain forest. Afromontane rain forest is found on wetter slopes from southern Ethiopia to Malawi, mostly between 1200 and 2500 meters elevation. It occurs on wetter slopes where average annual rainfall is from 1250 to 2500 mm, or higher. Elevation and location varies somewhat based on distance from the equator or from the sea, and the size and configuration of the highland where it occurs. Mature rain forests generally have an upper stratum of trees 25 to 45 meters high, a middle stratum 14 to 30 meters high, a lower stratum 6 to 15 meters high, a shrub layer of 3 to 6 meters high, and a sparse herbaceous ground layer. The tree crowns of the upper strata are typically open, and the middle strata may be continuous but rarely forms a dense canopy, while the lower tree stratum is typically dense. Most trees are evergreen. Afromontane rain forests can be similar in structure to lowland Guineo-Congolian rain forests, but the species mostly differ from lowland forests. Typical trees include '' Pouteria adolfi-friedericii,
Cola greenwayi ''Cola greenwayi'', commonly known as hairy cola or Zulu coshwood, is a species of flowering plant in the family Malvaceae. It was first described in 1956 by the British botanist John Patrick Micklethwait Brenan. It is native to southeastern Afri ...
, Cylicomorpha parviflora,
Diospyros abyssinica ''Diospyros abyssinica'' (also known as ''giant diospyros'', or ''Kôforonto'' and ''Baforonto'' in local languages spoken in parts of Mali, ''blidzo'', ''blitcho'', ''gblit∫o'', ''blonyat∫o'' or ''gblεt∫o'' in some Ghanaian languages) is ...
, Drypetes gerrardii, Entandrophragma excelsum, Ficalhoa laurifolia,
Gambeya gorungosana ''Gambeya gorungosana'' is species of evergreen tree native to eastern and central Africa. Range and habitat ''Gambeya gorungosana'' ranges across eastern and central Africa, from Mozambique and Zimbabwe through Zambia, Malawi, Tanzania, Burundi ...
, Mitragyna rubrostipulata, Myrianthus holstii,
Ochna holstii ''Ochna''Linnaeus C (1753) ''Sp. Pl.'' 1: 513. is a genus comprising 86 species of evergreen trees, shrubs and shrublets belonging to the flowering plant family ''Ochnaceae''. These species are native to tropical woodlands of Africa, the Mascare ...
, Ocotea usambarensis,
Olea capensis ''Olea capensis'', the black ironwood, is an African tree species in the olive family Oleaceae. It is widespread in sub-Saharan Africa: from the east in Somalia, Ethiopia and Sudan, south to the tip of South Africa, and west to Cameroon, Sierra ...
, Parinari excelsa,
Podocarpus milanjianus ''Podocarpus milanjianus'' is a species of conifer in the family Podocarpaceae. It is native to the highlands and mountains of tropical Africa. Description ''Podocarpus milanjianus'' is an evergreen tree which can grow slowly up to 35 meters in ...
,
Prunus africana ''Prunus africana'', the African cherry, has a wide distribution in Africa, occurring in montane regions of central and southern Africa and on the islands of Bioko, São-Tomé, Grande Comore, and Madagascar. It can be found at above sea level. I ...
, Strombosia scheffleri'', ''Syzygium guineense'' subsp. ''afromontanum'', '' Tabernaemontana johnstonii'', and '' Xymalos monospora''. Tree ferns (''
Cyathea ''Cyathea'' is a genus of tree ferns, the type genus of the fern order Cyatheales. The genus name ''Cyathea'' is derived from the Greek ''kyatheion'', meaning "little cup", and refers to the cup-shaped sori on the underside of the fronds. De ...
'' spp.), lianas, and epiphytes, including ferns, mosses, and species of ''
Begonia ''Begonia'' is a genus of perennial flowering plants in the family Begoniaceae. The genus contains more than 2,000 different plant species. The Begonias are native to moist subtropical and tropical climates. Some species are commonly grown ind ...
, Impatiens, Streptocarpus'', and ''
Peperomia ''Peperomia'' is one of the two large genera of the family Piperaceae. It is estimated that there are at least over 1,000 species, occurring in all tropical and subtropical regions of the world. They are concentrated in northern South America an ...
'', are abundant.


Distribution

In
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. It is bounded to the south by of coastline that stretch along the Atlantic Ocean, South Atlantic and Indian Oceans; to the ...
, Afromontane forests cover only 0.5% of the country's land area. The Afromontane forests occur along the mountainous arc of the
Drakensberg The Drakensberg (Afrikaans: Drakensberge, Zulu: uKhahlambha, Sotho: Maluti) is the eastern portion of the Great Escarpment, which encloses the central Southern African plateau. The Great Escarpment reaches its greatest elevation – within t ...
Range, from
Limpopo Province Limpopo is the northernmost province of South Africa. It is named after the Limpopo River, which forms the province's western and northern borders. The capital and largest city in the province is Polokwane, while the provincial legislature is ...
in the northeast to the
Western Cape Province The Western Cape is a province of South Africa, situated on the south-western coast of the country. It is the fourth largest of the nine provinces with an area of , and the third most populous, with an estimated 7 million inhabitants in 2020 ...
in the southwest. The Afromontane forests generally occur in well-watered areas, including ravines and south-facing slopes. The Afromontane forests are intolerant of fire, and the frequent fires of the surrounding fynbos,
savanna A savanna or savannah is a mixed woodland- grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground to ...
, and
grassland A grassland is an area where the vegetation is dominated by grasses ( Poaceae). However, sedge ( Cyperaceae) and rush ( Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur na ...
limit the expansion of the forests. Despite their small area, the Afromontane forests of South Africa produce valuable timber, particularly the
real yellowwood ''Podocarpus latifolius'' (broad-leaved yellowwood or real yellowwood, af, Opregte-geelhout, nso, Mogôbagôba, xh, Umcheya, zu, Umkhoba) is a large evergreen tree up to 35 m high and 3 m trunk diameter, in the conifer family Podoca ...
''(Podocarpus latifolius)'', Outeniqua yellowwood ''(Afrocarpus falcatus)'', and stinkwood ''(Ocotea bullata)''.


Afromontane ecoregions

;
Tropical and subtropical moist broadleaf forests Tropical and subtropical moist broadleaf forests (TSMF), also known as tropical moist forest, is a subtropical and tropical forest habitat type defined by the World Wide Fund for Nature. Description TSMF is generally found in large, discon ...
* Albertine Rift montane forests (Democratic Republic of the Congo, Burundi, Rwanda, Tanzania, Uganda) *
Cameroonian Highlands forests The Cameroonian Highlands forests, also known as the Cameroon Highlands forests, are a montane tropical moist broadleaf forest ecoregion located on the range of mountains that runs inland from the Gulf of Guinea and forms the border between Camer ...
(Cameroon, Nigeria) *
East African montane forests The East African montane forests is a montane tropical moist forest ecoregion of eastern Africa. The ecoregion comprises several separate areas above 2000 meters in the mountains of South Sudan, Uganda, Kenya, and Tanzania. Geography The East Afr ...
(Kenya, South Sudan, Tanzania, Uganda) *
Eastern Arc forests The Eastern Arc forests is a montane tropical moist forest ecoregion of eastern Africa. The ecoregion comprises several separate highland areas above 800 meters in Kenya, and (mostly) Tanzania. Setting The Eastern Arc forests extend across a tot ...
(Tanzania, Kenya) *
Ethiopian montane forests The Ethiopian montane forests is a tropical moist broadleaf forest ecoregion in eastern Africa. It covers the middle elevations of the Ethiopian Highlands in Ethiopia and extends into neighboring Eritrea, Sudan, Djibouti, and Somaliland. The ecore ...
(Djibouti, Eritrea, Ethiopia, Somalia, Sudan) *
Guinean montane forests The Guinean montane forests are a tropical moist broadleaf forest ecoregion of West Africa. The ecoregion occupies the portions of the Guinea Highlands lying above 600 meters elevation, extending across portions of Guinea, Sierra Leone, Lib ...
(Guinea, Côte d'Ivoire, Liberia, Sierra Leone) * Knysna-Amatole montane forests (South Africa) * Southern Afrotemperate Forest (Western Cape Province of South Africa) *
Mount Cameroon and Bioko montane forests The Mount Cameroon and Bioko montane forests is a tropical moist broadleaf forest ecoregion in central Africa. It occupies the upper slopes of coastal Mount Cameroon in Cameroon, and the mountains of nearby Bioko island in Equatorial Guinea. Ge ...
(Cameroon, Equatorial Guinea) ; Montane grasslands, shrublands, and woodlands * Angolan montane forest-grassland mosaic (Angola) *
Angolan scarp savanna and woodlands The Angolan Scarp savanna and woodlands is an ecoregion located on the coast of Angola, an area with a variety of habitats and rich in wildlife including many endemic birds and animals. Geography This ecoregion consists of the strip of land that ...
(Angola) *
Drakensberg alti-montane grasslands and woodlands The Drakensberg (Afrikaans: Drakensberge, Zulu: uKhahlambha, Sotho: Maluti) is the eastern portion of the Great Escarpment, which encloses the central Southern African plateau. The Great Escarpment reaches its greatest elevation – within t ...
(Lesotho, South Africa) *
Drakensberg montane grasslands, woodlands and forests The Drakensberg (Afrikaans: Drakensberge, Zulu language, Zulu: uKhahlambha, Sotho language, Sotho: Maluti) is the eastern portion of the Great Escarpment, Southern Africa, Great Escarpment, which encloses the central South Africa#Geography, Sou ...
(Lesotho, South Africa, Eswatini) *
East African montane moorlands The East African montane moorlands is a montane grasslands and shrublands ecoregion which occupies several high mountain peaks in Kenya, South Sudan, Tanzania, and Uganda. Geography The ecoregion occupies an area of , covering several small mount ...
(Kenya, South Sudan, Tanzania, Uganda) *
Eastern Zimbabwe montane forest-grassland mosaic :''"Eastern Highlands" also refers to Eastern Highlands Province in Papua New Guinea, and part of the Great Dividing Range, Australia.'' The Eastern Highlands, also known as the Manica Highlands, is a mountain range on the border of Zimbabw ...
(Mozambique, Zimbabwe) *
Ethiopian montane grasslands and woodlands The Ethiopian montane grasslands and woodlands is a montane grasslands and shrublands ecoregion in Ethiopia. It occupies the middle elevations of the Ethiopian Highlands, between the high-altitude Ethiopian montane moorlands and the lower-elevatio ...
(Eritrea, Ethiopia) *
Ethiopian montane moorlands The Ethiopian montane moorlands is a montane grasslands and shrublands ecoregion in Ethiopia. It lies above 3,000 meters elevation in the Ethiopian Highlands, the largest Afroalpine region in Africa. The montane moorlands lie above the tree line, ...
(Ethiopia) * Highveld grasslands (Lesotho, South Africa) * Jos Plateau forest-grassland mosaic (Nigeria) * Maputaland-Pondoland bushland and thickets (Mozambique, South Africa, Eswatini) * Southern Afrotemperate Forest (Western Cape Province of South Africa) * Rwenzori-Virunga montane moorlands (Democratic Republic of the Congo, Rwanda, Uganda) * South Malawi montane forest-grassland mosaic (Malawi, Mozambique) *
Southern Rift montane forest-grassland mosaic Southern may refer to: Businesses * China Southern Airlines, airline based in Guangzhou, China * Southern Airways, defunct US airline * Southern Air, air cargo transportation company based in Norwalk, Connecticut, US * Southern Airways Express, M ...
(Malawi, Tanzania) ;
Deserts and xeric shrublands Deserts and xeric shrublands are a biome defined by the World Wide Fund for Nature. Deserts and xeric (ancient Greek xērós, “dry") shrublands form the largest terrestrial biome, covering 19% of Earth's land surface area. Ecoregions in this h ...
* Southwestern Arabian montane woodlands (Saudi Arabia, Yemen)


External links


Eastern Afromontane Biodiversity Hotspot (Conservation International)



References

{{Reflist * Galley, C. & Linder, H. P. (2006) Geographical affinities of the Cape flora, South Africa. ''Journal of Biogeography'' 33 (2), 236–250. Afrotropical ecoregions Tropical and subtropical moist broadleaf forests Montane grasslands and shrublands Deserts and xeric shrublands Ecoregions of Africa Ecoregions of Saudi Arabia Ecoregions of Yemen * *