Dunlin
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The dunlin (''Calidris alpina'') is a small
wader 245px, A flock of Dunlins and Red knots">Red_knot.html" ;"title="Dunlins and Red knot">Dunlins and Red knots Waders or shorebirds are birds of the order Charadriiformes commonly found wikt:wade#Etymology 1, wading along shorelines and mudflat ...
, formerly sometimes separated with the other "
stint A stint is one of several very small waders in the paraphyletic ''"Calidris"'' assemblage – often separated in ''Erolia'' – which in North America are known as peeps. They are scolopacid waders much similar in ecomorphology to the ...
s" in the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''Erolia''. The English name is a dialect form of "dunling", first recorded in 1531–1532. It derives from ''dun'', "dull brown", with the suffix ''-ling'', meaning a person or thing with the given quality. It is a circumpolar breeder in
Arctic The Arctic ( or ) is a polar regions of Earth, polar region located at the northernmost part of Earth. The Arctic consists of the Arctic Ocean, adjacent seas, and parts of Canada (Yukon, Northwest Territories, Nunavut), Danish Realm (Greenla ...
or subarctic regions. Birds that breed in northern Europe and Asia are long-distance migrants, wintering south to Africa, southeast Asia and the Middle East. Birds that breed in
Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...
and the Canadian Arctic migrate short distances to the Pacific and Atlantic coasts of North America, although those nesting in northern Alaska overwinter in Asia. Many dunlins winter along the Iberian south coast.


Taxonomy

The dunlin was formally described by the Swedish naturalist
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
in 1758 in the tenth edition of his ''
Systema Naturae ' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial nomen ...
'' under the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Tringa alpina''. Linnaeus specified the
location In geography, location or place are used to denote a region (point, line, or area) on Earth's surface or elsewhere. The term ''location'' generally implies a higher degree of certainty than ''place'', the latter often indicating an entity with an ...
as Lapland. This species was formerly placed in the genus ''Erolia'', but is now placed with 23 other sandpipers in the genus ''
Calidris ''Calidris'' is a genus of Arctic-breeding, strongly migratory wading birds in the family Scolopacidae. These birds form huge mixed flocks on coasts and estuaries in winter. They are the typical "sandpipers", small to medium-sized, long-winged ...
'' that was introduced in 1804 by the German naturalist
Blasius Merrem Blasius Merrem (4 February 1761 – 23 February 1824) was a German naturalist, zoologist, ornithologist, mathematician, and herpetologist. In 1804, he became the professor of political economy and botany at the University of Marburg. Early li ...
. The genus name is from
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic peri ...
''kalidris'' or ''skalidris'', a term used by
Aristotle Aristotle (; grc-gre, Ἀριστοτέλης ''Aristotélēs'', ; 384–322 BC) was a Greek philosopher and polymath during the Classical period in Ancient Greece. Taught by Plato, he was the founder of the Peripatetic school of phil ...
for some grey-coloured waterside birds. The specific ''alpina '' is from
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
and means "of high mountains", in this case referring to the
Alps The Alps () ; german: Alpen ; it, Alpi ; rm, Alps ; sl, Alpe . are the highest and most extensive mountain range system that lies entirely in Europe, stretching approximately across seven Alpine countries (from west to east): France, Sw ...
. Ten
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
are recognised: *''C. a. arctica'' (Schiøler, 1922) — breeds in northeast
Greenland Greenland ( kl, Kalaallit Nunaat, ; da, Grønland, ) is an island country in North America that is part of the Kingdom of Denmark. It is located between the Arctic and Atlantic oceans, east of the Canadian Arctic Archipelago. Greenland is t ...
*''C. a. schinzii'' ( Brehm & Schilling, 1822) — breeds in southeast Greenland, Iceland, the
British Isles The British Isles are a group of islands in the North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner and Outer Hebrides, the Northern Isles, ...
,
Scandinavia Scandinavia; Sámi languages: /. ( ) is a subregion#Europe, subregion in Northern Europe, with strong historical, cultural, and linguistic ties between its constituent peoples. In English usage, ''Scandinavia'' most commonly refers to Denmark, ...
&
the Baltic The Baltic Sea is an arm of the Atlantic Ocean that is enclosed by Denmark, Estonia, Finland, Germany, Latvia, Lithuania, Poland, Russia, Sweden and the North and Central European Plain. The sea stretches from 53°N to 66°N latitude and from 10 ...
*''C. a. alpina'' (
Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
, 1758)
— breeds in northern Europe and northwest
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...
*''C. a. centralis'' (
Buturlin Buturlin, feminine: Buturlina (russian: Бутурли́н, Бутурлина) is a Russian surname of a Russian noble . Notable people with this surname include: * Alexander Buturlin (1694–1767), a Russian general * Anna Artemevna Buturlina ( ...
, 1932)
— breeds in north-central and northeast Siberia *''C. a. sakhalina'' (
Vieillot Louis Pierre Vieillot (10 May 1748, Yvetot – 24 August 1830, Sotteville-lès-Rouen) was a French ornithologist. Vieillot is the author of the first scientific descriptions and Linnaean names of a number of birds, including species he collect ...
, 1816)
— breeds in eastern
Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...
to the
Chukchi Peninsula The Chukchi Peninsula (also Chukotka Peninsula or Chukotski Peninsula; russian: Чуко́тский полуо́стров, ''Chukotskiy poluostrov'', short form russian: Чуко́тка, ''Chukotka''), at about 66° N 172° W, is the eastern ...
*''C. a. kistchinski'' Tomkovich, 1986 — breeds around the
Sea of Okhotsk The Sea of Okhotsk ( rus, Охо́тское мо́ре, Ohótskoye móre ; ja, オホーツク海, Ohōtsuku-kai) is a marginal sea of the western Pacific Ocean. It is located between Russia's Kamchatka Peninsula on the east, the Kuril Islands ...
to
Kuril Islands The Kuril Islands or Kurile Islands (; rus, Кури́льские острова́, r=Kuril'skiye ostrova, p=kʊˈrʲilʲskʲɪjə ɐstrɐˈva; Japanese: or ) are a volcanic archipelago currently administered as part of Sakhalin Oblast in the ...
and
Kamchatka The Kamchatka Peninsula (russian: полуостров Камчатка, Poluostrov Kamchatka, ) is a peninsula in the Russian Far East, with an area of about . The Pacific Ocean and the Sea of Okhotsk make up the peninsula's eastern and wes ...
*''C. a. actites'' Nechaev & Tomkovich, 1988 — breeds on
Sakhalin Sakhalin ( rus, Сахали́н, r=Sakhalín, p=səxɐˈlʲin; ja, 樺太 ''Karafuto''; zh, c=, p=Kùyèdǎo, s=库页岛, t=庫頁島; Manchu: ᠰᠠᡥᠠᠯᡳᠶᠠᠨ, ''Sahaliyan''; Orok: Бугата на̄, ''Bugata nā''; Nivkh: ...
*''C. a. arcticola'' (
Todd Todd or Todds may refer to: Places ;Australia: * Todd River, an ephemeral river ;United States: * Todd Valley, California, also known as Todd, an unincorporated community * Todd, Missouri, a ghost town * Todd, North Carolina, an unincorporated ...
, 1953)
— breeds from northwest Alaska to northwest Canada *''C. a. pacifica'' (
Coues Elliott Ladd Coues (; September 9, 1842 – December 25, 1899) was an American army surgeon, historian, ornithologist, and author. He led surveys of the Arizona Territory, and later as secretary of the United States Geological and Geograph ...
, 1861)
— breeds in western and southern Alaska *''C. a. hudsonia'' (
Todd Todd or Todds may refer to: Places ;Australia: * Todd River, an ephemeral river ;United States: * Todd Valley, California, also known as Todd, an unincorporated community * Todd, Missouri, a ghost town * Todd, North Carolina, an unincorporated ...
, 1953)
— breeds in central Canada


Description

Measurements: * Length: * Weight: * Wingspan: An adult dunlin in breeding plumage shows the distinctive black belly which no other similar-sized wader possesses. The winter dunlin is basically grey above and white below. Juveniles are brown above with two whitish "V" shapes on the back. They usually have black marks on the flanks or belly and show a strong white wingbar in flight. The legs and slightly decurved bill are black. There are a number of
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
differing mainly in the extent of rufous colouration in the breeding plumage and the bill length. Bill length varies between sexes, the females having longer bills than the males. On the tip of the Dunlin's bill is a soft covering that fills with blood and with many nerve endings, forming a sensitive probe that is used to locate invertebrate prey in mud and sand. Although the bill can look sharp-pointed in dead specimens, in life it is blunt. The call is a typical sandpiper "peep", and the display song a harsh trill.


Distribution and habitat

Dunlin are small migratory waders, however they show strong philopatry with individuals of the Southern Dunlin (''Calidris alpina schinzii'') in Sweden and Finland returning to, or very close to, their natal patches.
Habitat fragmentation Habitat fragmentation describes the emergence of discontinuities (fragmentation) in an organism's preferred environment (habitat), causing population fragmentation and ecosystem decay. Causes of habitat fragmentation include geological processes ...
has reduced the availability of habitat patches to these birds through reducing patch size and increasing patch isolation. This reduced connectivity between patches has reduced the movements of Dunlin leaving them more susceptible to inbreeding in these locations. Future management for the conservation of Southern Dunlin should include increasing the connectivity between habitat patches.


Behaviour

The dunlin is highly gregarious in winter, sometimes forming large flocks on coastal mudflats or sandy beaches. Large numbers can often be seen swirling in synchronized flight on stop-overs during migration or on their winter habitat. This bird is one of the most common waders throughout its breeding and wintering ranges, and it is the species with which other waders tend to be compared. At length and with a
wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingspan of ...
, it is similar in size to a
common starling The common starling or European starling (''Sturnus vulgaris''), also known simply as the starling in Great Britain and Ireland, is a medium-sized passerine bird in the starling family, Sturnidae. It is about long and has glossy black plumage ...
, but stouter, with a thicker bill. The dunlin moves along the coastal mudflat beaches it prefers with a characteristic "sewing machine" feeding action, methodically picking small food items.
Insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...
s form the main part of the dunlin's diet on the nesting grounds; it eats
molluscs Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000 extant taxon, extant species of molluscs are recognized. The number of fossil sp ...
,
worm Worms are many different distantly related bilateral animals that typically have a long cylindrical tube-like body, no limbs, and no eyes (though not always). Worms vary in size from microscopic to over in length for marine polychaete wor ...
s and
crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...
s in coastal areas.


Breeding

The nest is a shallow scrape on the ground lined with vegetation, into which typically four eggs are laid and incubated by the male and female parents. Chicks are precocial, however are brooded during early development. They start to fly at approximately three weeks of age. The majority of brood care is provided by the male, as the female deserts the brood and often leaves the breeding area. Apparent hybrids between this species and the
white-rumped sandpiper The white-rumped sandpiper (''Calidris fuscicollis'') is a small shorebird that breeds in the northern tundra of Canada and Alaska. This bird can be difficult to distinguish from other similar tiny shorebirds; these are known collectively as "pee ...
as well as with the
purple sandpiper The purple sandpiper (''Calidris maritima'') is a small shorebird in the sandpiper family Scolopacidae. This is a hardy sandpiper that breeds in the arctic and subarctic regions of Eurasia and North America and winters further south on the Atlant ...
have been reported from the Atlantic coasts of North America and Europe, respectively.


Status

The dunlin has an extremely large range and although the population appears to be decreasing, the population is still very large. The
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
(IUCN) has judged that the threat to the species is of "
Least concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. T ...
". The dunlin is one of the species to which the ''Agreement on the Conservation of African-Eurasian Migratory Waterbirds'' (
AEWA The Agreement on the Conservation of African-Eurasian Migratory Waterbirds, or African-Eurasian Waterbird Agreement (AEWA) is an independent international treaty developed under the auspices of the United Nations Environment Programme's Conventio ...
) applies.


Gallery

File:Seasonal-Variations-in-the-Diet-and-Foraging-Behaviour-of-Dunlins-Calidris-alpina-in-a-South-pone.0081174.s001.ogv, Foraging File:Kärrsnäppa - (Calidris alpina).jpg,
Ystad Ystad (; older da, Ysted) is a town and the seat of Ystad Municipality, in Scania County, Sweden. Ystad had 18,350 inhabitants in 2010. The settlement dates from the 11th century and has become a busy ferryport, local administrative centre, and ...
, Sweden File:Alpenstrandläufer (calidris alpina) - Spiekeroog, Nationalpark niedersächsisches Wattenmeer.jpg, Adult in breeding plumage,
Spiekeroog Spiekeroog is one of the East Frisian Islands, off the North Sea coast of Germany. It is situated between Langeoog to its west, and Wangerooge to its east. The island belongs to the district of Wittmund, in Lower Saxony in Germany. The only vi ...
, northern Germany File:Calidris alpina pibydd y mawn.jpg, Adult in winter plumage, Wales File:Calidris alpina alpina, Riga, Latvia 1.jpg, Adult in breeding plumage, Latvia


References


External links

* * *
Dunlin Species Account
- Cornell Lab of Ornithology

- USGS Patuxent Bird Identification InfoCenter
Ageing and sexing (PDF; 1.6 MB) by Javier Blasco-Zumeta & Gerd-Michael Heinze
* * * * * {{Authority control Calidris Birds of the Arctic Holarctic birds Birds of the Dominican Republic Birds of the Caribbean Birds of North Africa Birds described in 1758 Taxa named by Carl Linnaeus Articles containing video clips