Donax (bivalve)
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''Donax'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of small, edible saltwater clams, marine bivalve molluscs. The genus is sometimes known as bean clams or wedge shells; however, ''Donax'' species have numerous different common names in different parts of the world. In the southeastern U.S. they are known as " coquina", a word that is also used for the hard
limestone Limestone ( calcium carbonate ) is a type of carbonate sedimentary rock which is the main source of the material lime. It is composed mostly of the minerals calcite and aragonite, which are different crystal forms of . Limestone forms whe ...
concretions of their shells and those of other marine organisms.


Ecology

Species of ''Donax'' live, sometimes in high concentrations, vertically aligned in the sand on exposed beaches, on tropical and temperate coasts worldwide. When the waves wash these small clams out of the sand, they can dig back in again quite rapidly. They are filter feeders. Some species, such as '' Donax variabilis'', migrate vertically and horizontally with changes in the
tide Tides are the rise and fall of sea levels caused by the combined effects of the gravitational forces exerted by the Moon (and to a much lesser extent, the Sun) and are also caused by the Earth and Moon orbiting one another. Tide tables ...
s. These coquina clams are found extensively on the east coast beaches of Trinidad (Mayaro) and widely available in Venezuela. They are called "Chip Chip" in Trinidad & Tobago and "Chipi Chipi" in Venezuela.


Species

Species within the genus ''Donax'' include: * '' Donax asper'' Hanley 1845eol Encyclopedia of Life
/ref> * '' Donax assimilis'' Hanley, 1845 * '' Donax baliregteri'' Huber, 2012 * ''
Donax bertini ''Donax'' is the scientific name of two genera of organisms and may refer to: * ''Donax'' (bivalve), a genus of clams in the family Donacidae * ''Donax'' (plant), a genus of plants in the family Marantaceae {{Genus disambiguation ...
'' Pilsbry, 1901 * '' Donax bipartitus'' Sowerby III, 1892 * '' Donax brazieri''
Smith Smith may refer to: People * Metalsmith, or simply smith, a craftsman fashioning tools or works of art out of various metals * Smith (given name) * Smith (surname), a family name originating in England, Scotland and Ireland ** List of people wi ...
, 1892
* '' Donax bruneirufi'' Huber, 2012 * '' Donax burnupi'' Sowerby III, 1894 * '' Donax caelatus'' Carpenter, 1857 * '' Donax californicus'' Conrad, 1837 * '' Donax carinatus'' Hanley, 1843 * '' Donax clathratus'' Reeve, 1855 * '' Donax culter'' Hanley, 1845 * '' Donax cuneatus''
Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
, 1758
* '' Donax denticulatus''
Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
, 1758
* '' Donax dentifer'' Hanley, 1843 * '' Donax ecuadorianus'' Olsson, 1961 * '' Donax erythraeensis'' Bertin, 1881 * '' Donax faba'' Gmelin, 1791 * '' Donax fossor'' Say, 1822 - was thought at one time to be a northern form of '' D. variabilis'' * '' Donax francisensis'' (Cotton & Godfrey, 1938) * '' Donax gemmula'' Morrison, 1971 * '' Donax gouldii'' Dall, 1921 - The bean clam, aka ''Donax gracilis'' Hanley 1845 * '' Donax hanleyanus'' ( Philippi, 1847) * '' Donax kindermanni'' ( Philippi, 1847) * '' Donax lubricus'' Hanley, 1845 * '' Donax madagascariensis''
Wood Wood is a porous and fibrous structural tissue found in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulose fibers that are strong in tension and embedded in a matrix of lignin ...
, 1828
* '' Donax nitidus'' Deshayes, 1855 * '' Donax obesulus'' Reeve 1854 * '' Donax obesus'' d'Orbigny, 1845 * '' Donax oweni'' Hanley, 1843 * '' Donax paxillus'' Reeve, 1854 * '' Donax punctatostriatus'' Hanley, 1843 * '' Donax rothi'' Coan, 1983 * '' Donax semistriatus'' Poli, 1795 * '' Donax serra'' Röding, 1798 * '' Donax simplex'' Sowerby III, 1897 * '' Donax sordidus'' Hanley, 1845 * '' Donax striatus''
Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
, 1767
* '' Donax texasianus'' Philippi, 1847 * '' Donax townsendi'' Sowerby III, 1894 * '' Donax transversus'' Sowerby I, 1825 * '' Donax trunculus''
Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
, 1758
- Locally known as ''tellin'' in French, ''
tellina ''Tellina'' is a widely distributed genus of marine bivalve molluscs, in the family Tellinidae.Gofas, S. (2011). Tellina Linnaeus, 1758. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id ...
'' or ''telline'' in Italian, ''coquina'' in Spanish and ''conquilha'' or ''cadelinha'' in Portuguese. * '' Donax variabilis'' Say, 1822 - Known in Florida under the Spanish word "coquina", is the common and colorful ''Donax'' species of the southern Atlantic and Gulf shores from Virginia to the Caribbean. This species is locally abundant on beaches with fine sand from the mid-Atlantic coast to
Texas Texas (, ; Spanish: ''Texas'', ''Tejas'') is a state in the South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2020, it is the second-largest U.S. state by ...
. * '' Donax variegatus'' ( Gmelin, 1791) * '' Donax vellicatus'' Reeve, 1855 * '' Donax veneriformis''
Lamarck Jean-Baptiste Pierre Antoine de Monet, chevalier de Lamarck (1 August 1744 – 18 December 1829), often known simply as Lamarck (; ), was a French naturalist, biologist, academic, and soldier. He was an early proponent of the idea that biolo ...
, 1818
* ''
Donax veruinus ''Donax'' is the scientific name of two genera of organisms and may refer to: * ''Donax'' (bivalve), a genus of clams in the family Donacidae * ''Donax'' (plant), a genus of plants in the family Marantaceae {{Genus disambiguation ...
'' Hanley, 1913 * '' Donax vittatus'' (
da Costa Costa (, , , , ), sometimes Costas, da Costa, Da Costa, or Dalla Costa, is an Italian, Portuguese and Spanish surname. The surname spread throughout the world through colonization. It is also a surname chosen by Jews and other religious groups due ...
, 1778)
The empty small (15 to 25 mm) shells of ''Donax variabilis'' and ''Donax fossor'' may be found washed up on the beach, especially at low tide. The living animals can often be seen where the waves wash the sand around in the shallowest part of the
littoral zone The littoral zone or nearshore is the part of a sea, lake, or river that is close to the shore. In coastal ecology, the littoral zone includes the intertidal zone extending from the high water mark (which is rarely inundated), to coastal a ...
as the tidal level changes. These clams can use the action of waves to move themselves up and down the beach, quickly burrowing into a new location before they can be swept away (the so-called "dance of the coquina").


References


Wildlife of Sydney - Fact File - Pipi, Goolwa Cockle, Surf Clam Fact File


Further reading


The Nautilus
''On the species of Donax of Eastern North America'', March 1892, pp. 125-126
Wayne A. O'Connor & Stephan J. O'Connor, "Early ontogeny of the pipi, Donax (Plebidonax) deltoides (Donacidae; Bivalvia)"
''Molluscan Research'' 31(1): 53–56; {{Taxonbar, from=Q2302897 Donacidae Bivalve genera