Desidae
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Desidae is a family of
spider Spiders (order Araneae) are air-breathing arthropods that have eight legs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species ...
s, some of which are known as intertidal spiders. The family is named for the genus '' Desis'', members of which live in a very unusual location — between the
tide Tides are the rise and fall of sea levels caused by the combined effects of the gravitational forces exerted by the Moon (and to a much lesser extent, the Sun) and are also caused by the Earth and Moon orbiting one another. Tide tables ...
s. The family has been reevaluated in recent years and now includes inland genera and species as well, such as ''
Badumna ''Badumna'' is a genus of intertidal spiders that was first described by Tamerlan Thorell in 1890. They are harmless spiders that can be found around human structures and buildings. The most well-known species is '' B. insignis'', also known a ...
'' and ''
Phryganoporus ''Phryganoporus'' is a genus of Australian intertidal spiders that was first described by Eugène Simon in 1908. Its five species only occur in Australia and Tasmania, with one species also occurring on Norfolk Island, south of New Caledonia. ...
''. In 2017, the family Amphinectidae was merged into Desidae. The family
Toxopidae Toxopidae is a small family of araneomorph spiders, first described in 1940. For many years it was sunk into Desidae as a subfamily, although doubts were expressed as to whether this was correct. A large-scale molecular phylogenetic study in 2016 ...
has been separated off. Those intertidal spiders that are truly marine commonly live in
barnacle A barnacle is a type of arthropod constituting the subclass Cirripedia in the subphylum Crustacea, and is hence related to crabs and lobsters. Barnacles are exclusively marine, and tend to live in shallow and tidal waters, typically in erosiv ...
shells, which they seal up with silk; this allows them to maintain an air bubble during high tide. They emerge at night to feed on various small
arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and cuticle made of chiti ...
s that live in the intertidal zone.


Distribution

As now circumscribed, the family Desidae is mainly found in
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sou ...
and
Australasia Australasia is a region that comprises Australia, New Zealand and some neighbouring islands in the Pacific Ocean. The term is used in a number of different contexts, including geopolitically, physiogeographically, philologically, and ecologic ...
, with some species reaching north to
Malaysia Malaysia ( ; ) is a country in Southeast Asia. The federal constitutional monarchy consists of thirteen states and three federal territories, separated by the South China Sea into two regions: Peninsular Malaysia and Borneo's East Mal ...
. ''
Metaltella simoni ''Metaltella simoni'' is a species of spider, native to South America (Brazil, Uruguay and Argentina), and introduced into the United States and Canada. Left palp of male, ventral view Description Females are about 8 to 9 mm long, males ab ...
'' has been introduced in a large part of the Southern United States (records exist from
California California is a state in the Western United States, located along the Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the most populous U.S. state and the 3rd largest by area. It is also the m ...
,
Louisiana Louisiana , group=pronunciation (French: ''La Louisiane'') is a state in the Deep South and South Central regions of the United States. It is the 20th-smallest by area and the 25th most populous of the 50 U.S. states. Louisiana is bord ...
,
Mississippi Mississippi () is a state in the Southeastern region of the United States, bordered to the north by Tennessee; to the east by Alabama; to the south by the Gulf of Mexico; to the southwest by Louisiana; and to the northwest by Arkansas. Miss ...
, and
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and ...
) and is considered an
invasive species An invasive species otherwise known as an alien is an introduced organism that becomes overpopulated and harms its new environment. Although most introduced species are neutral or beneficial with respect to other species, invasive species adv ...
in
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and ...
. It is feared that it could
extirpate Local extinction, also known as extirpation, refers to a species (or other taxon) of plant or animal that ceases to exist in a chosen geographic area of study, though it still exists elsewhere. Local extinctions are contrasted with global extinct ...
the native titanoecid species ''Titanoeca brunnea''.


Genera

, the
World Spider Catalog The World Spider Catalog (WSC) is an online searchable database concerned with spider taxonomy. It aims to list all accepted families, genera and species, as well as provide access to the related taxonomic literature. The WSC began as a series of ...
accepts the following genera: *''
Akatorea ''Akatorea'' is a genus of South Pacific intertidal spiders first described by Raymond Robert Forster & C. L. Wilton in 1973. it contains only two species, both found in New Zealand New Zealand ( mi, Aotearoa ) is an island country ...
'' Forster & Wilton, 1973 — New Zealand *''
Amphinecta ''Amphinecta'' is a genus of South Pacific intertidal spiders first described by Eugène Simon in 1898. Species it contains eleven species, all found in New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southweste ...
'' Simon, 1898 — New Zealand *''
Austmusia ''Austmusia'' is a genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below ...
'' Gray, 1983 — Australia *''
Badumna ''Badumna'' is a genus of intertidal spiders that was first described by Tamerlan Thorell in 1890. They are harmless spiders that can be found around human structures and buildings. The most well-known species is '' B. insignis'', also known a ...
'' Thorell, 1890 — Oceania, Asia, North America, Uruguay *''
Baiami ''Baiami'' is a genus of Australian intertidal spiders that was first described by Pekka T. Lehtinen in 1967. Originally placed with the Stiphidiidae, it was transferred to the Desidae after the results of a 2019 genetic analysis. Species it c ...
'' Lehtinen, 1967 — Australia *'' Bakala'' Davies, 1990 — Australia *''
Barahna ''Barahna'' is a genus of Australian intertidal spiders that was first described by V. T. Davies in 2003. The name is derived from ''baran-barahn'', the Bundjalung word for "spider". Originally placed with the Stiphidiidae, it was moved to the ...
'' Davies, 2003 — Australia *''
Buyina ''Buyina'' is a genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below ...
'' Davies, 1998 — Australia *'' Calacadia'' Exline, 1960 — Chile *''
Cambridgea ''Cambridgea'' (common name New Zealand sheetweb spider, bush spider) is a spider genus in the family Desidae and some of the first endemic spiders described from New Zealand. They are known for constructing large horizontal sheet webs measurin ...
'' L. Koch, 1872 — New Zealand *''
Canala Canala is a commune A commune is an alternative term for an intentional community. Commune or comună or comune or other derivations may also refer to: Administrative-territorial entities * Commune (administrative division), a municipality or ...
'' Gray, 1992 — New Caledonia *'' Cicirra'' Simon, 1886 — Australia *''
Colcarteria ''Colcarteria'' is a genus of Australia Australia, officially the Commonwealth of Australia, is a sovereign country comprising the mainland of the Australian continent, the island of Tasmania, and numerous smaller islands. With an ar ...
'' Gray, 1992 — Australia *''
Corasoides ''Corasoides'' is a genus of South Pacific intertidal spiders that was first described by Arthur Gardiner Butler in 1929. Originally placed with the Agelenidae, it was moved to the Stiphidiidae in 1973, and to the Desidae after a 2017 genetic st ...
'' Butler, 1929 — Papua New Guinea, Australia *'' Cunnawarra'' Davies, 1998 — Australia *'' Desis'' Walckenaer, 1837 — Africa, Ecuador, Asia, Oceania *''
Dunstanoides ''Dunstanoides'' is a genus of Oceania, South Pacific Desidae, intertidal spiders first described by Norman I. Platnick in 1989. Species it contains nine species, all found in New Zealand: *''Dunstanoides angustiae'' (Marples, 1959) – New Zea ...
'' Forster & Wilton, 1989 — New Zealand *''
Epimecinus ''Epimecinus'' is a genus of South Pacific intertidal spiders that was first described by Eugène Simon in 1908. Species it contains four species: *'' Epimecinus alkirna'' Gray, 1973 – Australia (Western Australia) *'' Epimecinus humilis'' B ...
'' Simon, 1908 — Australia *'' Forsterina'' Lehtinen, 1967 — Australia *'' Goyenia'' Forster, 1970 — New Zealand *'' Helsonia'' Forster, 1970 — New Zealand *'' Holomamoea'' Forster & Wilton, 1973 — New Zealand *''
Huara Huara is a Chilean town and commune in Tamarugal Province, Tarapacá Region. It is located or ( by road) northeast of Iquique. The village is crossed by the Pan-American Highway and is the crossing point for the road that goes to Oruro in Boli ...
'' Forster, 1964 — New Zealand *'' Ischalea'' L. Koch, 1872 — Madagascar, Mauritius, New Zealand *''
Jalkaraburra ''Jalkaraburra'' is a genus of Australian intertidal spiders containing the single species, ''Jalkaraburra alta''. It was first described by V. T. Davies in 1998, and has only been found in Australia Australia, officially the Commonw ...
'' Davies, 1998 — Australia *'' Keera'' Davies, 1998 — Australia *'' Lathyarcha'' Simon, 1908 — Australia *''
Magua Magua is a fictional character and the main antagonist in the 1826 novel ''The Last of the Mohicans'' by James Fenimore Cooper. This historical novel is set at the time of the French and Indian War. A Huron Indian chief, he is also known by the F ...
'' Davies, 1998 — Australia *''
Makora ''Makora'' is a genus of South Pacific intertidal spiders first described by Raymond Robert Forster & C. L. Wilton in 1973. Species it contains five species, all found in New Zealand New Zealand ( mi, Aotearoa ) is an island country ...
'' Forster & Wilton, 1973 — New Zealand *''
Mamoea ''Mamoea'' is a genus of Oceania, South Pacific Desidae, intertidal spiders first described by Raymond Robert Forster & C. L. Wilton in 1973. Species it contains nineteen species, all found in New Zealand: *''Mamoea assimilis'' Forster & Wilton ...
'' Forster & Wilton, 1973 — New Zealand *'' Mangareia'' Forster, 1970 — New Zealand *'' Maniho'' Marples, 1959 — New Zealand *'' Manjala'' Davies, 1990 — Australia *'' Matachia'' Dalmas, 1917 — New Zealand *''
Mesudus ''Mesudus'' is a genus of South Pacific intertidal spiders that was first described by H. Özdikmen in 2007. it contains only three species, all found in New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southweste ...
'' Özdikmen, 2007 — New Zealand *'' Metaltella'' Mello-Leitão, 1931 — South America, North America *'' Namandia'' Lehtinen, 1967 — Australia *'' Nanocambridgea'' Forster & Wilton, 1973 — New Zealand *''
Neororea ''Neororea'' is a genus of Oceania, South Pacific Desidae, intertidal spiders first described by Raymond Robert Forster & C. L. Wilton in 1973. it contains only two species, both found in New Zealand. References

Araneomorphae genera Des ...
'' Forster & Wilton, 1973 — New Zealand *'' Notomatachia'' Forster, 1970 — New Zealand *''
Nuisiana ''Nuisiana'' is a monotypic genus of South Pacific intertidal spiders containing the single species, ''Nuisiana arboris''. It was first described by Raymond Robert Forster & C. L. Wilton in 1973, and has only been found in New Zealand N ...
'' Forster & Wilton, 1973 — New Zealand *'' Oparara'' Forster & Wilton, 1973 — New Zealand *'' Panoa'' Forster, 1970 — New Zealand *'' Paramamoea'' Forster & Wilton, 1973 — New Zealand *''
Paramatachia ''Paramatachia'' is a genus of Australia Australia, officially the Commonwealth of Australia, is a sovereign country comprising the mainland of the Australian continent, the island of Tasmania, and numerous smaller islands. With an a ...
'' Dalmas, 1918 — Australia *'' Penaoola'' Davies, 1998 — Australia *''
Phryganoporus ''Phryganoporus'' is a genus of Australian intertidal spiders that was first described by Eugène Simon in 1908. Its five species only occur in Australia and Tasmania, with one species also occurring on Norfolk Island, south of New Caledonia. ...
'' Simon, 1908 — Australia *'' Pitonga'' Davies, 1984 — Northern Australia *'' Poaka'' Forster & Wilton, 1973 — New Zealand *'' Porteria'' Simon, 1904 — Chile *'' Quemusia'' Davies, 1998 — Australia *'' Rangitata'' Forster & Wilton, 1973 — New Zealand *'' Rapua'' Forster, 1970 — New Zealand *'' Reinga'' Forster & Wilton, 1973 — New Zealand *'' Rorea'' Forster & Wilton, 1973 — New Zealand *'' Syrorisa'' Simon, 1908 — Australia *'' Tanganoides'' Davies, 2005 — Australia *'' Taurongia'' Hogg, 1901 — Australia *'' Tuakana'' Forster, 1970 — New Zealand *''
Waterea ''Waterea'' is a genus of South Pacific intertidal spiders containing the single species, ''Waterea cornigera''. It was first described by Raymond Robert Forster & C. L. Wilton in 1973, and has only been found in New Zealand New Zealan ...
'' Forster & Wilton, 1973 — New Zealand


See also

*
List of Desidae species This page lists all described genera and species of the spider family Desidae. , the World Spider Catalog accepts 330 species in 60 genera: A ''Akatorea'' '' Akatorea'' Forster & Wilton, 1973 * '' Akatorea gracilis'' (Marples, 1959) ( type) — ...


References


External links

{{Taxonbar, from=Q10038 Araneomorphae families