Depressariinae
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__NOTOC__ The Depressariinae – sometimes spelled "Depressiinae" in error – are a subfamily of
moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...
s in the superfamily
Gelechioidea __NOTOC__ Gelechioidea (from the type genus ''Gelechia ''Gelechia'' is a genus of moths in the family Gelechiidae. The type species is '' Gelechia rhombella''. Species *Subgenus ''Gelechia'' **'' Gelechia asinella'' (Hubner, 1796) **'' Gel ...
. Like their relatives therein, their exact relationships are not yet very well resolved. It has been considered part of
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Elachistidae The Elachistidae (grass-miner moths) are a family of small moths in the superfamily Gelechioidea. Some authors lump about 3,300 species in eight subfamilies here, but this arrangement almost certainly results in a massively paraphyletic and comp ...
''
sensu lato ''Sensu'' is a Latin word meaning "in the sense of". It is used in a number of fields including biology, geology, linguistics, semiotics, and law. Commonly it refers to how strictly or loosely an expression is used in describing any particular c ...
'' or included in an expanded
Oecophoridae Oecophoridae (concealer moths) is a family of small moths in the superfamily Gelechioidea. The phylogeny and systematics of gelechoid moths are still not fully resolved, and the circumscription of the Oecophoridae is strongly affected by this. ...
.Scoble (1995), ABRS (2008), FE (2009), Wikispecies (24 March 2010), and see references in Savela (2009) In modern classifications they are treated as the distinct gelechioid family
Depressariidae Depressariidae is a family of moths. It has formerly been treated as a subfamily of Gelechiidae, but is now recognised as a separate family, comprising about 2,300 species worldwide.Heikkilä, M. ''et al''. 2014: Morphology reinforces proposed mo ...
. As regards subdivisions, the
Amphisbatinae __NOTOC__ The Amphisbatinae was a small subfamily of moths in the superfamily Gelechioidea. Like their relatives therein, their exact relationships are not yet very well resolved. The present lineage is often included in the Depressariinae as a ...
are often held to be as close to the Depressariinae as to be included there, in particular if the latter are raised to full family rank, but also otherwise (as a
tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide usage of the term in English is in the discipline of anthropology. This definition is contested, in part due to confli ...
Amphisbatini in the Depressariinae). If Depressariinae are treated as oecophorid subfamily, it is more common to include the group around '' Cryptolechia'', which is sometimes (especially in older arrangements) separated as tribe ''Cryptolechiini'' in subfamily ''Cryptolechiinae''. The lineage of '' Cacochroa'' and its closest relatives – typically included in the "Cryptolechiinae" if these are treated as independent oecophorid subfamily – actually seems to be closer to '' Orophia'', and would be part of tribe Orophiini if these are placed in the Oecophorinae. But they have also been included in the present group (as tribe(s) Orophiini and sometimes also Cacochroini), particularly if this is considered a full-fledged family. The same holds true for the
Xyloryctidae Xyloryctidae is a family of moths contained within the superfamily Gelechioidea described by Edward Meyrick in 1890. Most genera are found in the Indo-Australian region. While many of these moths are tiny, some members of the family grow to a wing ...
, a group treated here as family but previously thought to be a subfamily of the Oecophoridae. Finally, the '' Semioscopis'' lineage is occasionally separated from the rest of the Depressariinae as tribe ''Epigraphiini'' (after its junior synonym ''Epigraphia''). In all such treatments, the core group of Depressariinae becomes a tribe ''Depressariini''.


Distribution and ecology

Some 600
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
were placed here as of 1999 (but see the caveats about the group's
circumscription Circumscription may refer to: *Circumscribed circle * Circumscription (logic) *Circumscription (taxonomy) *Circumscription theory, a theory about the origins of the political state in the history of human evolution proposed by the American anthrop ...
above). New
taxa In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...
of Depressariinae are still being discovered and described on a regular basis. The group is found worldwide except on some
oceanic island An island (or isle) is an isolated piece of habitat that is surrounded by a dramatically different habitat, such as water. Very small islands such as emergent land features on atolls can be called islets, skerries, cays or keys. An island ...
s and frozen wasteland, but the diversity is highest in
temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout ...
regions and the group is presumably
Holarctic The Holarctic realm is a biogeographic realm that comprises the majority of habitats found throughout the continents in the Northern Hemisphere. It corresponds to the floristic Boreal Kingdom. It includes both the Nearctic zoogeographical reg ...
in origin – possibly
Palaearctic The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa. The realm consists of several bioregions: the Euro-Sibe ...
, as gelechoid diversity in the
Neotropics The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In bioge ...
is poor but e.g. in Australia almost 20 species are found. From Europe alone, more than 160 species and subspecies (mostly of
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
'' Agonopterix'') were known in 2009, with over 80 recorded from
Central Europe Central Europe is an area of Europe between Western Europe and Eastern Europe, based on a common historical, social and cultural identity. The Thirty Years' War (1618–1648) between Catholicism and Protestantism significantly shaped the a ...
. The caterpillars usually develop in leaves spun together with
silk Silk is a natural protein fiber, some forms of which can be woven into textiles. The protein fiber of silk is composed mainly of fibroin and is produced by certain insect larvae to form cocoons. The best-known silk is obtained from the ...
, as stem borers or as seed or flower feeders of dicotyledons. Recorded Depressariinae host plants are mainly
eurosids I The rosids are members of a large clade (monophyletic group) of flowering plants, containing about 70,000 species, more than a quarter of all angiosperms. The clade is divided into 16 to 20 orders, depending upon circumscription and classificatio ...
(e.g.
Betulaceae Betulaceae, the birch family, includes six genera of deciduous nut-bearing trees and shrubs, including the birches, alders, hazels, hornbeams, hazel-hornbeam, and hop-hornbeams numbering a total of 167 species. They are mostly natives of the ...
, Fabaceae, Fagaceae, Rosaceae,
Salicaceae The Salicaceae is the willow family of flowering plants. The traditional family (Salicaceae ''sensu stricto'') included the willows, poplar, aspen, and cottonwoods. Genetic studies summarized by the Angiosperm Phylogeny Group (APG) have greatly ...
,
Urticaceae The Urticaceae are a family, the nettle family, of flowering plants. The family name comes from the genus ''Urtica''. The Urticaceae include a number of well-known and useful plants, including nettles in the genus ''Urtica'', ramie (''Boehmeri ...
) but also from some other families (e.g.
Malvaceae Malvaceae, or the mallows, is a family of flowering plants estimated to contain 244 genera with 4225 known species. Well-known members of economic importance include okra, cotton, cacao and durian. There are also some genera containing familiar ...
and
Rutaceae The Rutaceae is a family, commonly known as the rueRUTACEAE
in BoDD – Botanical Der ...
eurosids II The rosids are members of a large clade ( monophyletic group) of flowering plants, containing about 70,000 species, more than a quarter of all angiosperms. The clade is divided into 16 to 20 orders, depending upon circumscription and classific ...
–, or
euasterids II In the APG IV system (2016) for the classification of flowering plants, the name asterids denotes a clade (a monophyletic group). Asterids is the largest group of flowering plants, with more than 80,000 species, about a third of the total floweri ...
like
Apiaceae Apiaceae or Umbelliferae is a family of mostly aromatic flowering plants named after the type genus '' Apium'' and commonly known as the celery, carrot or parsley family, or simply as umbellifers. It is the 16th-largest family of flowering plan ...
and
Asteraceae The family Asteraceae, alternatively Compositae, consists of over 32,000 known species of flowering plants in over 1,900 genera within the order Asterales. Commonly referred to as the aster, daisy, composite, or sunflower family, Compositae w ...
).


Genera

Genera of Depressariinae – with some notable
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
also listed – include:ABRS (2008), Wikispecies (24 March 2010), and see references in Savela (2009) *Tribe Fuchsiini **''
Fuchsia ''Fuchsia'' () is a genus of flowering plants that consists mostly of shrubs or small trees. The first to be scientifically described, ''Fuchsia triphylla'', was discovered on the Caribbean island of Hispaniola (Haiti and the Dominican Republic) ...
'' Spuler, 1910 *Tribe Telechrysidini **'' Telechrysis'' Toll, 1956 *Tribe Depressariini **'' Agonopterix'' Hübner, 1825 **'' Apachea'' Clarke, 1941 **'' Bibarrambla'' Clarke, 1941 **'' Depressaria'' **'' Exaeretia'' Stainton, 1849 **'' Himmacia'' Clarke, 1941 **'' Levipalpus'' Hannemann, 1953 **'' Luquetia'' Leraut, 1991 **'' Nites'' Hodges, 1974 **'' Semioscopis'' Hübner, 1825 *Tribe Amphisbatini **''
Eupragia ''Eupragia'' is a moth genus of the family Depressariidae Depressariidae is a family of moths. It has formerly been treated as a subfamily of Gelechiidae, but is now recognised as a separate family, comprising about 2,300 species worldwide.Heik ...
'' Walsingham, 1911 **''
Machimia ''Machimia'' is a genus of moths of the family Depressariidae described by James Brackenridge Clemens in 1860. Species * ''Machimia aethostola'' Meyrick, 1931 * ''Machimia albula'' Turner, 1946 * ''Machimia anthracospora'' Meyrick, 1934 * ''Mach ...
'' (Amphisbatinae?) **'' Psilocorsis'' Clemens, 1860 (Amphisbatinae?) *Unplaced to tribe **'' Acria'' Stephens, 1834 **'' Afdera'' Clarke, 1978 **'' Ancipita'' Busck, 1914 **'' Athrinacia'' Walsingham, 1911 **'' Barantola'' Walker, 1864 **'' Bleptochiton'' Turner, 1947 **'' Chariphylla'' Meyrick, 1921 **'' Comotechna'' Meyrick, 1920 **'' Costoma'' Busck, 1914 **'' Deloryctis'' Meyrick, 1934 **''
Doina The doina () is a Romanian musical tune style, possibly with Middle Eastern roots, customary in Romanian peasant music, as well as in Lăutărească. It was also adopted into klezmer music. Similar tunes are found throughout Eastern Europe an ...
'' Clarke, 1978 **'' Doleromima'' Meyrick, 1902 **'' Doshia'' Clarke, 1978 **'' Eclecta'' Meyrick, 1883 **'' Ectaga'' Walsingham, 1912 **'' Enchocrates'' Meyrick, 1883 **'' Enteremna'' Meyrick, 1917 **'' Erithyma'' Meyrick, 1914 **'' Euprionocera'' Turner, 1896 **'' Eutorna'' Meyrick, 1889 **'' Filinota'' Busck, 1911 **'' Gnathotona'' Meyrick, 1931 **'' Gonada'' Busck, 1911 **'' Gymnoceros'' Turner, 1946 **'' Habrophylax'' Meyrick, 1931 **'' Haereta'' Turner, 1947 **'' Hamadera'' Busck, 1914 **'' Hastamea'' Busck, 1940 **'' Heterobathra'' Lower, 1901 **'' Himotica'' Meyrick, 1912 **'' Hozbeka'' Özdikmen, 2009 (formerly ''Talitha'' Clarke, 1978 (''non'' Faure, 1958: preoccupied)) **'' Idiocrates'' Meyrick, 1909 **'' Iphimachaera'' Meyrick, 1931 **'' Lepidozancla'' Turner, 1916 **'' Loboptila'' Turner, 1919 **'' Lucyna'' Clarke, 1978 **'' Maesara'' Clarke, 1968 **'' Magniophaga'' Beéche, 2018 **'' Mimozela'' Meyrick, 1914 **''
Muna Muna may refer to: Places * Muna (Mikulovice), a World War II POW camp and ammunition factory in the Czech Republic * Muna, Estonia, village in Rõuge Parish, Võru County, Estonia * Muna, Iran, village in East Azerbaijan Province, Iran * Mu ...
'' Clarke, 1978 **'' Nedenia'' Clarke, 1978 **'' Nematochares'' Meyrick, 1931 **'' Notosara'' Meyrick, 1890 **'' Octasphales'' Meyrick, 1886 **'' Osmarina'' Clarke, 1978 **'' Palinorsa'' Meyrick, 1924 **'' Pedois'' Lower, 1894 **'' Peritornenta'' Turner, 1900 **'' Perzelia'' Clarke, 1978 **'' Philtronoma'' Meyrick, 1914 **'' Pholcobates'' Meyrick, 1931 **'' Phytomimia'' Walsingham, 1912 **'' Pisinidea'' Butler, 1883 **'' Profilinota'' Clarke, 1973 **'' Pseudocentris'' Meyrick, 1921 **'' Psittacastis'' Meyrick, 1909 **'' Psorosticha'' Lower, 1901 **'' Ptilobola'' Meyrick, 1933 **'' Rhindoma'' Busck, 1914 **'' Scoliographa'' Meyrick, 1916 **'' Scorpiopsis'' Turner, 1894 **'' Thalamarchella'' T. B. Fletcher, 1940 (formerly ''Thalamarchis'' Meyrick, 1904 (''non'' Meyrick, 1897: preoccupied)) **'' Thyromorpha'' Turner, 1917 **'' Tonica'' Walker, 1864 **'' Trycherodes'' Meyrick, 1914


Footnotes


References

See also Gelechioidea Talk page for comparison of some approaches to gelechioid systematics and taxonomy. * (2008): Australian Faunal Directory &ndash
Depressariidae
'sic''.html" ;"title="sic.html" ;"title="'sic">'sic''">sic.html" ;"title="'sic">'sic'' Version of 9 October 2008. Retrieved 24 April 2010. * (2009)
Depressariidae
'sic''.html" ;"title="sic.html" ;"title="'sic">'sic''">sic.html" ;"title="'sic">'sic'' Version 2.1, 22 December 2009. Retrieved 22 February 2010. * (1999): The Gelechioidea. ''In:'' : ''Handbuch der Zoologie/Handbook of Zoology'' (Volume IV – Arthropoda: Insecta. Part 35: Lepidoptera, Moths and Butterflies 1): 131–158. Walter de Gruyter, Berlin & New York. * (2010)
Depressariidae [''sic''
/nowiki> (Flachleibmotten) in Mitteleuropa] [in German]. Version of 28 March 2010. Retrieved 22 February 2010. * (2009): ''Lepidoptera and Some Other Life Forms'' &ndash
Depressiinae
'sic''.html"_;"title="sic.html"_;"title="'sic">'sic''">sic.html"_;"title="'sic">'sic''_Version_of_20_August_2009._Retrieved_22_April_2010. *_(1995):_''The_Lepidoptera:_Form,_Function_and_Diversity''._Oxford_University_Press,_Oxford._ {{Taxonbar.html" ;"title="sic">' 'sic''">sic.html"_;"title="'sic">'sic''_Version_of_20_August_2009._Retrieved_22_April_2010. *_(1995):_''The_Lepidoptera:_Form,_Function_and_Diversity''._Oxford_University_Press,_Oxford._ {{Taxonbar">from=Q1128027 Depressariinae.html" ;"title="sic''.html" ;"title="sic.html" ;"title="'sic">'sic''">sic.html" ;"title="'sic">'sic'' Version of 20 August 2009. Retrieved 22 April 2010. * (1995): ''The Lepidoptera: Form, Function and Diversity''. Oxford University Press, Oxford. {{Taxonbar">from=Q1128027 Depressariinae"> Depressariidae Moth subfamilies