Delacour's langur
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Delacour's langur or Delacour's lutung (''Trachypithecus delacouri'') is a critically endangered species of
lutung The lutungs, langurs, or leaf monkeys are a group of Old World monkeys in the genus ''Trachypithecus'' (derived from Greek , meaning "rough" and , meaning "monkey"). Their range is much of Southeast Asia (northeast India, Vietnam, southern Chin ...
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found else ...
to northern
Vietnam Vietnam or Viet Nam ( vi, Việt Nam, ), officially the Socialist Republic of Vietnam,., group="n" is a country in Southeast Asia, at the eastern edge of mainland Southeast Asia, with an area of and population of 96 million, making i ...
. It is named for French-American ornithologist Jean Théodore Delacour.


Physical description

Delacour's langur is somewhat larger than its two closest relatives,
François' langur François' langur (''Trachypithecus francoisi''), also known as Francois' leaf monkey, the Tonkin leaf monkey, or the white side-burned black langur is a species of lutung and the type species of its species group. It is one of the least studied ...
and the
Laotian langur The Laotian langur or white-browed black langur (''Trachypithecus laotum'') is a species of primate in the family Cercopithecidae. It is endemic to Laos. Its natural habitat is subtropical or tropical dry forests. Genetically the Indochinese bla ...
, but in other respects has a similar appearance. Adults measure from in head-body length, with a tail long. Males weigh between , while the females are slightly smaller, weighing between . Their fur is predominantly black, with white markings on the face and distinctive creamy-white fur over the rump and the outer thighs, while females also have a patch of pale fur in the pubic area. Like other closely related lutungs, Delacour's langur has a crest of long, upright, hair over the forehead and crown; this is, however, somewhat taller and narrower than in other species.


Distribution and habitat

Delacour's langur is
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found else ...
to Vietnam, where it is found only in an area around in the provinces of Ninh Bình,
Hà Nam Hà is a Vietnamese surname. The name is transliterated as He in Chinese and Ha in Korean. Ha is the anglicized variation of the surname Hà. It is also the anglicized variation of Hạ. Notable people with the surname Hà * Hà Kiều Anh, ...
, Hòa Bình,
Thanh Hóa Thanh Hóa () is the capital of Thanh Hóa Province. The city is situated in the east of the province on the Ma River (Sông Mã), about 150 kilometers (93 miles) south of Hanoi and 1560 kilometers (969 miles) north of Ho Chi Minh City. Thanh ...
, and Hà Tây in the north of the country. The largest surviving population is thought to live in Van Long Nature Reserve in Ninh Bình, where the it inhabits open forest up to elevations of in terrain dominated by
limestone Limestone ( calcium carbonate ) is a type of carbonate sedimentary rock which is the main source of the material lime. It is composed mostly of the minerals calcite and aragonite, which are different crystal forms of . Limestone forms whe ...
karst.


Behaviour

Delacour's langurs are diurnal, often spending the day sleeping in limestone caves, although they sleep on bare rocky surfaces if no caves are available. They are
folivorous In zoology, a folivore is a herbivore that specializes in eating leaves. Mature leaves contain a high proportion of hard-to-digest cellulose, less energy than other types of foods, and often toxic compounds.Jones, S., Martin, R., & Pilbeam, D. (1 ...
, with about 78% of the diet reportedly consisting of foliage, although they also eat fruit, seeds, and flowers. The monkeys have been reported to eat leaves from a wide range of different plant species, indicating that their apparent dependence on limestone habitats is not related to their diet. In previous decades, Delacour's langurs were reported to live in troops of up to 30 individuals, often including a mix of males and females, although single-male groups are more common, and some small all-male groups have also been reported. In more recent years, the typical group size seems to be much smaller, with only about four to 16 members each. Males defend the troop's territory from outsiders, often standing watch on rocky outcrops. When potential rivals are spotted, the males in a troop initially try to intimidate them with loud hoots and visual displays, resorting to chasing and fighting if this fails. Within the group, social bonds are maintained by grooming and play. Despite living in forested habitats, Delacour's langurs are primarily terrestrial, only occasionally venturing into the trees. They swing by their hands when travelling through trees, and use their tails for balance when scrambling over steep rocky terrain, which may allow them to move more rapidly than other related lutungs.


Reproduction

Females give birth to a single young after a gestation period of 170 to 200 days. The young are born with orange fur, and are
precocial In biology, altricial species are those in which the young are underdeveloped at the time of birth, but with the aid of their parents mature after birth. Precocial species are those in which the young are relatively mature and mobile from the mome ...
, with open eyes and strong arms. The fur begins to turn black at around four months, and the young are probably weaned at 19 to 21 months, when the mother is likely ready to breed again. However, the full adult coat pattern is not achieved for around three years. Females reach sexual maturity at four years, and males at five years; the total life expectancy is around 20 years.


Conservation

The population of Delacour's langurs has declined rapidly in recent years. As of 2006, only 19 populations were known, following a dramatic decline in the total population of approximately 20% between 1999 and 2004. Since that time, two of the populations have been
extirpated Local extinction, also known as extirpation, refers to a species (or other taxon) of plant or animal that ceases to exist in a chosen geographic area of study, though it still exists elsewhere. Local extinctions are contrasted with global extinct ...
, and only that in the Van Long Nature Reserve may still be large enough to remain viable. Classified as critically endangered by the IUCN, the primary threat to the species is hunting for
traditional medicine Traditional medicine (also known as indigenous medicine or folk medicine) comprises medical aspects of traditional knowledge that developed over generations within the folk beliefs of various societies, including indigenous peoples, before the ...
, with loss of forest habitat and the local development of tourism also being a potential risk. As of 2010, less than 250 animals were believed to remain in the wild, with nineteen in captivity.


References


External links

*ARKive
images and information on Delacour’s langur ''(Trachypithecus delacouri)''
{{Taxonbar, from=Q681546 Delacour's langur Endemic fauna of Vietnam Mammals of Vietnam Primates of Southeast Asia Critically endangered fauna of Asia Delacour's langur Species endangered by human consumption for medicinal or magical purposes Taxa named by Wilfred Hudson Osgood