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The Cyprinidae
Cyprinidae
are the family of freshwater fishes, collectively called cyprinids, that includes the carps, the true minnows, and their relatives (for example, the barbs and barbels). Also commonly called the "carp family", or "minnow family", Cyprinidae
Cyprinidae
is the largest known fish family and the largest vertebrate animal family in general, with about 3,000 living and extinct species in about 370 genera.[1][2] The family belongs to the ostariophysian order Cypriniformes, of whose genera and species the cyprinids make more than two-thirds.[1][2][3] The family name is derived from the Ancient Greek
Ancient Greek
kyprînos (κυπρῖνος, "carp").

Contents

1 Biology and ecology 2 Relationship with humans 3 Systematics 4 Phylogeny 5 Subfamilies
Subfamilies
and genera

5.1 Incertae sedis

6 See also 7 References 8 External links

Biology and ecology[edit] Cyprinids are stomachless fish with toothless jaws. Even so, food can be effectively chewed by the gill rakers of the specialized last gill bow. These pharyngeal teeth allow the fish to make chewing motions against a chewing plate formed by a bony process of the skull. The pharyngeal teeth are unique to each species and are used by scientists to identify species. Strong pharyngeal teeth allow fish such as the common carp and ide to eat hard baits such as snails and bivalves. Hearing is a well-developed sense in the cyprinids since they have the Weberian organ, three specialized vertebral processes that transfer motion of the gas bladder to the inner ear. The vertebral processes of the Weberian organ
Weberian organ
also permit a cyprinid to detect changes in motion of the gas bladder due to atmospheric conditions or depth changes. The cyprinids are considered physostomes because the pneumatic duct is retained in adult stages and the fish are able to gulp air to fill the gas bladder, or they can dispose excess gas to the gut.

Giant barbs ( Catlocarpio
Catlocarpio
siamensis) are the largest members of this family

Cyprinids are native to North America, Africa, and Eurasia. The largest known cyprinid is the giant barb ( Catlocarpio
Catlocarpio
siamensis), which may grow up to 3 m (9.8 ft) in length and 300 kg (660 lb) in weight.[4] Other very large species that can surpass 2 m (6.6 ft) are the golden mahseer (Tor putitora) and mangar ( Luciobarbus
Luciobarbus
esocinus).[5][6] The largest North American species is the Colorado pikeminnow
Colorado pikeminnow
( Ptychocheilus
Ptychocheilus
lucius), which can reach up to 1.8 m (5.9 ft) in length.[7] Conversely, many species are smaller than 5 cm (2 in). The smallest known fish is Paedocypris
Paedocypris
progenetica, reaching 10.3 mm (0.41 in) at the longest.[8] All fish in this family are egg-layers and most do not guard their eggs; however, a few species build nests and/or guard the eggs. The bitterlings of subfamily Acheilognathinae
Acheilognathinae
are notable for depositing their eggs in bivalve molluscs, where the young develop until able to fend for themselves. Most cyprinids feed mainly on invertebrates and vegetation, probably due to the lack of teeth and stomach; however, some species, like the asp, are predators that specialize in fish. Many species, such as the ide and the common rudd, prey on small fish when individuals become large enough. Even small species, such as the moderlieschen, are opportunistic predators that will eat larvae of the common frog in artificial circumstances. Some cyprinids, such as the grass carp, are specialized herbivores; others, such as the common nase, eat algae and biofilms, while others, such as the black carp, specialize in snails, and some, such as the silver carp, are specialized filter feeders. For this reason, cyprinids are often introduced as a management tool to control various factors in the aquatic environment, such as aquatic vegetation and diseases transmitted by snails. Unlike most fish species, cyprinids generally increase in abundance in eutrophic lakes. Here, they contribute towards positive feedback as they are efficient at eating the zooplankton that would otherwise graze on the algae, reducing its abundance. Relationship with humans[edit]

Wild capture of cyprinids by species in million tonnes, 1950–2009, as reported by the FAO[9]

Cyprinids are highly important food fish; they are fished and farmed across Eurasia. In land-locked countries in particular, cyprinids are often the major species of fish eaten because they make the largest part of biomass in most water types except for fast-flowing rivers. In Eastern Europe, they are often prepared with traditional methods such as drying and salting. The prevalence of inexpensive frozen fish products made this less important now than it was in earlier times. Nonetheless, in certain places, they remain popular for food, as well as recreational fishing, and have been deliberately stocked in ponds and lakes for centuries for this reason.[10] Cyprinids are popular for angling especially for match fishing (due to their dominance in biomass and numbers) and fishing for common carp because of its size and strength. Several cyprinids have been introduced to waters outside their natural ranges to provide food, sport, or biological control for some pest species. The common carp ( Cyprinus
Cyprinus
carpio) and the grass carp ( Ctenopharyngodon
Ctenopharyngodon
idella) are the most important of these, for example in Florida. In some cases, such as the Asian carp
Asian carp
in the Mississippi Basin, they have become invasive species that compete with native fishes or disrupt the environment. Carp
Carp
in particular can stir up sediment, reducing the clarity of the water and making it difficult for plants to grow.[11][12] Numerous cyprinids have become important in the aquarium and fishpond hobbies, most famously the goldfish, which was bred in China from the Prussian carp
Prussian carp
( Carassius
Carassius
(auratus) gibelio). First imported into Europe
Europe
around 1728, it was much fancied by Chinese nobility as early as 1150 AD and after it arrived there in 1502, also in Japan. In the latter country, from the 18th century onwards, the common carp was bred into the ornamental variety known as koi – or more accurately nishikigoi (錦鯉), as koi (鯉) simply means "common carp" in Japanese. Other popular aquarium cyprinids include danionins, rasborines, and true barbs.[13] Larger species are bred by the thousands in outdoor ponds, particularly in Southeast Asia, and trade in these aquarium fishes is of considerable commercial importance. The small rasborines and danionines are perhaps only rivalled by characids and poecilid livebearers in their popularity for community aquaria.[citation needed] One particular species of these small and undemanding danionines is the zebrafish ( Danio
Danio
rerio). It has become the standard model species for studying developmental genetics of vertebrates, in particular fish.[14] Habitat destruction
Habitat destruction
and other causes have reduced the wild stocks of several cyprinids to dangerously low levels; some are already entirely extinct. In particular, the cyprinids of the subfamily Leuciscinae from southwestern North America
North America
have been hit hard by pollution and unsustainable water use in the early to mid-20th century; most globally extinct cypriniform species are in fact leuciscinid cyprinids from the southwestern United States and northern Mexico. Systematics[edit] The massive diversity of cyprinids has so far made it difficult to resolve their phylogeny in sufficient detail to make assignment to subfamilies more than tentative in many cases. Some distinct lineages obviously exist – for example, the Cultrinae
Cultrinae
and Leuciscinae, regardless of their exact delimitation, are rather close relatives and stand apart from Cyprininae – but the overall systematics and taxonomy of the Cyprinidae
Cyprinidae
remain a subject of considerable debate. A large number of genera are incertae sedis, too equivocal in their traits and/or too little-studied to permit assignment to a particular subfamily with any certainty.[15][16][17] Part of the solution seems that the delicate rasborines are the core group, consisting of minor lineages that have not shifted far from their evolutionary niche, or have coevolved for millions of years. These are among the most basal lineages of living cyprinids. Other "rasborines" are apparently distributed across the diverse lineages of the family.[16] The validity and circumscription of proposed subfamilies like the Labeoninae
Labeoninae
or Squaliobarbinae
Squaliobarbinae
also remain doubtful, although the latter do appear to correspond to a distinct lineage. The sometimes-seen grouping of the large-headed carps (Hypophthalmichthyinae) with Xenocypris, though, seems quite in error. More likely, the latter are part of the Cultrinae.[16] The entirely paraphyletic "Barbinae" and the disputed Labeoninae
Labeoninae
might be better treated as part of the Cyprininae, forming a close-knit group whose internal relationships are still little known. The small African "barbs" do not belong in Barbus
Barbus
sensu stricto – indeed, they are as distant from the typical barbels and the typical carps (Cyprinus) as these are from Garra
Garra
(which is placed in the Labeoninae by most who accept the latter as distinct) and thus might form another as yet unnamed subfamily. However, as noted above, how various minor lineages tie into this has not yet been resolved; therefore, such a radical move, though reasonable, is probably premature.[15][18][19] The tench ( Tinca
Tinca
tinca), a significant food species farmed in western Eurasia
Eurasia
in large numbers, is unusual. It is most often grouped with the Leuciscinae, but even when these were rather loosely circumscribed, it always stood apart. A cladistic analysis of DNA sequence data of the S7 ribosomal protein intron 1 supports the view that it is distinct enough to constitute a monotypic subfamily. It also suggests it may be closer to the small East Asian Aphyocypris, Hemigrammocypris, and Yaoshanicus. They would have diverged roughly at the same time from cyprinids of east-central Asia, perhaps as a result of the Alpide orogeny
Alpide orogeny
that vastly changed the topography of that region in the late Paleogene, when their divergence presumably occurred.[17] A DNA-based analysis of these fish places the Rasborinae as the basal lineage with the Cyprininae as a sister clade to the Leuciscinae.[20] The subfamilies Acheilognathinae, Gobioninae, and Leuciscinae
Leuciscinae
are monophyletic. Phylogeny[edit]

Phylogeny
Phylogeny
of living Cyprinoidei[21][22] with clade names from van der Laan 2017.[23]

Psilorhynchidae

Psilorhynchus

Cyprinidae

Probarbinae

Labeoninae

Parapsilorhynchini

Labeonini

Garrini

Torinae

Smiliogastrinae

Cyprininae

Cyprinini

Rohteichthyini

Acrossocheilini

Spinibarbini

Schizothoracini

Schizopygopsini

Barbini

Leuciscidae

Danioninae

?Paedocypridini

?Sundadanionini

Rasborini

Danionini

Chedrini

Leptobarbinae

Leptobarbus

Xenocypridinae

Squaliobarbini

Opsariichthyini

Oxygastrini

Hypophthalmichthyini

Xenocypridini

Tincinae

Tinca

Acheilognathinae

Gobioninae

Hemibarbus- Squalidus
Squalidus
clade

Sarcocheilichthyini

Gobionini

Tanichthyinae

Tanichthys

Leuciscinae

Phoxinini

Laviniini

Plagiopterini

Leuciscini

Pogonichthyini

Subfamilies
Subfamilies
and genera[edit]

Rainbow shark, Epalzeorhynchos
Epalzeorhynchos
frenatum, a somewhat aggressive aquarium fish

Acheilognathus
Acheilognathus
longipinnis: Acheilognathinae

Blue danio, Danio
Danio
kerri: Danioninae

Pseudogobio esocinus
Pseudogobio esocinus
, Gobioninae

Silver carp, Hypophthalmichthys
Hypophthalmichthys
molitrix: Xenocyprinae, alternatively Hypophthalmichthyinae

Rohu, Labeo
Labeo
rohita, of the disputed Labeoninae

The tench, Tinca
Tinca
tinca, is of unclear affiliations and often placed in a subfamily of its own.

Subfamily
Subfamily
Probarbinae

Catlocarpio Probarbus

Subfamily
Subfamily
Labeoninae

Tribe Parapsilorhynchini

Diplocheilichthys Neorohita Parapsilorhynchus Longanalus Protolabeo Sinilabeo

Tribe Labeonini

Altigena Bangana Cirrhinus
Cirrhinus
(mud carps) Gymnostomus Incisilabeo Labeo
Labeo
(labeos) Speolabeo Schismatorhynchos Vinalabeo

Tribe Garrini

Garra Paracrossocheilus Tariqilabeo Osteochilus
Osteochilus
clade

Barbichthys Crossocheilus Epalzeorhynchos Henicorhynchus Labiobarbus Lobocheilos Osteochilus Thynnichthys

Semilabeo clade

Ageneiogarra Cophecheilus Discogobio Hongshuia Linichthys Mekongina Paraqianlabeo [24] Parasinilabeo Placocheilus Prolixicheilus Pseudocrossocheilus Pseudogyrinocheilus Ptychidio Qianlabeo Rectoris Semilabeo Sinigarra [25] Sinocrossocheilus Stenorynchoacrum [26] Vinagarra

Subfamily
Subfamily
Torinae

Acapoeta Arabibarbus
Arabibarbus
[27] Barbopsis
Barbopsis
(Somalian blind barb) Carasobarbus Hypselobarbus Labeobarbus
Labeobarbus
(yellowfish) Lepidopygopsis

Mesopotamichthys Naziritor (Zhobi mahseers) Neolissochilus
Neolissochilus
(mahseers) Osteochilichthys Sanagia Tor (mahseers) Pterocapoeta

Subfamily
Subfamily
Smiliogastrinae

Barbodes Barboides Caecobarbus
Caecobarbus
(Congo blind barb) Chagunius Clypeobarbus Coptostomabarbus Dawkinsia
Dawkinsia
[28] Desmopuntius
Desmopuntius
[29] Eechathalakenda Enteromius Haludaria
Haludaria
[30] Hampala Oliotius
Oliotius
[29] Oreichthys

Osteobrama Pethia
Pethia
[28] Prolabeo Prolabeops Pseudobarbus
Pseudobarbus
(redfins) Puntigrus
Puntigrus
[29] Puntius
Puntius
(spotted barbs) Rohtee
Rohtee
(Vatani rohtee) Sahyadria Striuntius
Striuntius
[29] Systomus Xenobarbus

Subfamily
Subfamily
Cyprininae [incl. Barbinae]

Tribe Cyprinini

Aaptosyax
Aaptosyax
(giant salmon carp) Carassioides Carassius
Carassius
(Crucian carps and goldfish) Cyprinus
Cyprinus
(typical carps) Luciocyprinus Paraspinibarbus Parator Procypris Pseudosinocyclocheilus Sinibarbus Sinocyclocheilus (golden-line fish) Typhlobarbus

Tribe Rohteichthyini

Albulichthys Amblyrhynchichthys Anematichthys Balantiocheilos Barbonymus
Barbonymus
(tinfoil barbs) Cosmochilus Cyclocheilichthys Cyclocheilos Discherodontus Eirmotus Hypsibarbus Kalimantania Laocypris Mystacoleucus Parasikukia Poropuntius Puntioplites Rohteichthys Sawbwa (Sawbwa barb) Scaphognathops Sikukia Troglocyclocheilus

Tribe Acrossocheilini

Acrossocheilus Folifer Onychostoma

Tribe Spinibarbini

Spinibarbus Spinibarbichthys

Tribe Schizothoracini

Aspiorhynchus Percocypris Schizopyge (snowtrouts) Schizothorax
Schizothorax
(snowtrouts)

Tribe Schizopygopsini

Chuanchia Diptychus Gymnocypris Gymnodiptychus Oreinus Oxygymnocypris Platypharodon Ptychobarbus Schizopygopsis
Schizopygopsis
(snowtrouts)

Tribe Barbini

Aulopyge
Aulopyge
(Dalmatian barbelgudgeon) Barbus
Barbus
(typical barbels and barbs) †Hsianwenia[31] Caecocypris Capoeta
Capoeta
(khramulyas) Cyprinion Kantaka Luciobarbus Scaphiodonichthys Schizocypris (snowtrouts) Semiplotus

Subfamily
Subfamily
Danioninae

Tribe Paedocypridini

Paedocypris

Tribe Sundadanionini

Fangfangia [32] Sundadanio

Tribe Rasborini

Amblypharyngodon
Amblypharyngodon
(carplets) Boraras
Boraras
(rasboras) Brevibora Horadandia Kottelatia Pectenocypris Rasbora Rasboroides Rasbosoma
Rasbosoma
(dwarf scissortail rasbora) Trigonopoma Trigonostigma

Tribe Danionini

Betadevario Brachydanio Celestichthys Chela Danio
Danio
(danios) Danionella Devario Inlecypris Laubuka Microdevario Microrasbora

Tribe Chedrini

Barilius Bengala Cabdio [Aspidoparia] Chelaethiops Engraulicypris Esomus
Esomus
(flying barbs) Leptocypris Luciosoma Malayochela Nematabramis Neobola Opsaridium Opsarius Raiamas Rastrineobola
Rastrineobola
(silver cyprinid) Salmostoma
Salmostoma
(razorbelly minnows) Securicula Thryssocypris

Subfamily
Subfamily
Leptobarbinae

Leptobarbus

Flame chub
Flame chub
Hemitremia
Hemitremia
flammea, one of the chubs in the Leuciscinae)

Ide, Leuciscus
Leuciscus
idus , one of the Eurasian daces

Sailfin shiner, Notropis
Notropis
hypselopterus, a small and colorful shiner of the Leuciscinae

Rhynchocypris
Rhynchocypris
oxycephalus, a minnow related to some North American daces

Sarmarutilus
Sarmarutilus
rubilio, a European roach

Trigonostigma
Trigonostigma
somphongsi, a rasbora, a relative of the blue danio above

Black carp, Mylopharyngodon
Mylopharyngodon
piceus: Squaliobarbinae

Subfamily
Subfamily
Xenocypridinae [incl. Cultrinae
Cultrinae
& Squaliobarbinae]

Tribe Squaliobarbini

Squaliobarbus

Tribe Opsariichthyini

Candidia Nipponocypris Opsariichthys Parazacco Xenocyprioides

Tribe Oxygastrini

Aphyocypris Araiocypris Gymnodanio Hemigrammocypris Macrochirichthys
Macrochirichthys
(long pectoral-fin minnow) Metzia Oxygaster Parachela Paralaubuca Rasborichthys

Tribe Hypophthalmichthyini

Atrilinea Ctenopharyngodon
Ctenopharyngodon
(grass carp) Elopichthys Hypophthalmichthys
Hypophthalmichthys
(bighead carps) Luciobrama Mylopharyngodon
Mylopharyngodon
(black carp) Ochetobius

Tribe Xenocypridini

Subtribe Xenocypridina

Distoechodon Plagiognathops Pseudobrama Xenocypris

Subtribe Cultrina

Anabarilius Chanodichthys Culter Ischikauia Longiculter Megalobrama Parabramis (white Amur bream) Pogobrama Sinibrama

Hemiculter
Hemiculter
clade

Hainania Hemiculter
Hemiculter
(sharpbellies) Pseudohemiculter Pseudolaubuca Toxabramis

Subfamily
Subfamily
Tincinae

Tinca

Subfamily
Subfamily
Acheilognathinae
Acheilognathinae
(bitterlings)

 ? Acanthorhodeus (Khanka spiny bitterling) Acheilognathus Paratanakia Pseudorhodeus Rhodeus Tanakia

Subfamily
Subfamily
Gobioninae

Hemibarbus- Squalidus
Squalidus
clade

Belligobio Hemibarbus
Hemibarbus
(steeds) Squalidus

Tribe Gobionini

Subtribe Gobiobotiina

Gobiobotia Xenophysogobio

Subtribe Gobionina

Gobio
Gobio
(typical gudgeons) Mesogobio Romanogobio

Subtribe Armatogobionina

Abbottina
Abbottina
(false gudgeons) Biwia  ?Huigobio Microphysogobio Platysmacheilus Pseudogobio Saurogobio

Tribe Sarcocheilichthyini

Coreius Coreoleuciscus (Korean splendid dace) Gnathopogon Gobiocypris Ladislavia Paracanthobrama Paraleucogobio  ?Parasqualidus Pseudopungtungia Pseudorasbora Pungtungia Rhinogobio Sarcocheilichthys

Subfamily
Subfamily
Tanichthyinae

Tanichthys

Subfamily
Subfamily
Leuciscinae
Leuciscinae
[incl. Alburninae]

Tribe Phoxinini

Oreoleuciscus Phoxinus
Phoxinus
(Eurasian minnows and daces) Pseudaspius

Tribe Laviniini

Subtribe Chrosomina

Chrosomus
Chrosomus
(typical daces)

Subtribe Laviniina

Eremichthys
Eremichthys
(desert dace) Gila (western chubs) Hesperoleucus
Hesperoleucus
(California roach) Klamathella Lavinia (hitch) Mylopharodon
Mylopharodon
(hardhead) Orthodon
Orthodon
(Sacramento blackfish) Ptychocheilus
Ptychocheilus
(pikeminnows) Relictus
Relictus
(relict dace) Siphateles

Tribe Leuciscini

Pachychilon clade

Pachychilon

Alburnoides
Alburnoides
clade

Alburnoides

Primitive Leuciscine clade

Delminichthys Leucalburnus Notemigonus
Notemigonus
(golden shiner) Pelasgus

Subtribe Leuciscina

Aspiolucius
Aspiolucius
(pike asp) Leuciscus
Leuciscus
(Eurasian daces) Pelecus
Pelecus
(sabre carp)

Subtribe Abramina

Abramis
Abramis
(common bream) Acanthobrama
Acanthobrama
(bleaks) Capoetobrama Mirogrex Vimba
Vimba
(Vimbas)

Subtribe Chondrostomina

Achondrostoma Alburnus
Alburnus
(bleaks) Anaecypris Chondrostoma
Chondrostoma
(typical nases) Iberochondrostoma Leucaspius
Leucaspius
(moderlieschen) Leucos
Leucos
[33] Parachondrostoma Petroleuciscus (Ponto-Caspian chubs and daces) Phoxinellus Protochondrostoma
Protochondrostoma
(South European nase) Pseudochondrostoma Pseudophoxinus Rutilus
Rutilus
(roaches) Sarmarutilus
Sarmarutilus
[33] Scardinius
Scardinius
(rudds) Squalius
Squalius
(European chubs) Telestes Tropidophoxinellus

Tribe Plagiopterini

Couesius
Couesius
(lake chub) Hemitremia
Hemitremia
(flame chub) Lepidomeda (spinedaces) Margariscus (pearl daces) Meda (pikedace) Plagopterus
Plagopterus
(woundfin) Rhynchocypris
Rhynchocypris
(Eurasian minnows) Semotilus
Semotilus
(creek chubs) † Stypodon
Stypodon
(stumptooth minnow)

Tribe Pogonichthyini

Subtribe Pogonichthyina

Clinostomus
Clinostomus
(redside daces) Iotichthys
Iotichthys
(least chub) Mylocheilus
Mylocheilus
(peamouth) Pogonichthys
Pogonichthys
(splittails) Richardsonius
Richardsonius
(redside shiners)

Subtribe Exoglossina

Exoglossum
Exoglossum
(cutlips minnows) Oregonichthys
Oregonichthys
(Oregon chubs) Pararhinichthys
Pararhinichthys
(cheat minnow) Rhinichthys
Rhinichthys
(riffle daces, loach minnows) Tiaroga

Subtribe Campostomina

Campostoma
Campostoma
(stonerollers) Nocomis
Nocomis
(hornyhead chubs)

Subtribe Hybognathina

Agosia
Agosia
(longfin dace) Alburnops Algansea (Mexican chubs)  ? Aztecula
Aztecula
(Aztec chub)  ? Ballerus
Ballerus
(breams)  ? Blicca
Blicca
(silver bream) Codoma (ornate shiner) Cyprinella
Cyprinella
(satinfin shiners) Dionda
Dionda
(desert minnows)  ? Ericymba
Ericymba
(longjaw minnows) Erimonax Erimystax
Erimystax
(slender chubs) † Evarra
Evarra
(Mexican daces) Graodus Hudsonius Hybognathus
Hybognathus
(silvery minnows) Hybopsis
Hybopsis
(bigeye chubs)  ?Iberocypris  ?Ladigesocypris Luxilus
Luxilus
(highscale shiners) Lythrurus
Lythrurus
(finescale shiners) Macrhybopsis (blacktail chubs) Miniellus  ?Moapa (moapa dace) Notropis
Notropis
(eastern shiners) Opsopoeodus
Opsopoeodus
(pugnose minnow) Phenacobius
Phenacobius
(suckermouth minnows) Pimephales
Pimephales
(bluntnose minnows) Platygobio
Platygobio
(flathead chub) Pteronotropis
Pteronotropis
(flagfin shiners)  ? Snyderichthys (spinedaces) Tampichthys  ?Tribolodon  ?Yuriria

Incertae sedis[edit]

Hemigrammocypris
Hemigrammocypris
rasborella, of uncertain relationship: Possibly related to Aphyocypris.

Acanthalburnus
Acanthalburnus
(bleaks) Acanthogobio Acrocheilus
Acrocheilus
(chiselmouth) Ancherythroculter Anchicyclocheilus Gibelion (catla) (some authorities consider this species to belong in the genus Catla) Cultrichthys Discocheilus Discolabeo Hemiculterella Herzensteinia

Horalabiosa Megarasbora Neobarynotus Paracrossochilus Phreatichthys
Phreatichthys
(Somalian cavefish) Placogobio Scardinius
Scardinius
(rudds) Tropidophoxinellus Typhlogarra
Typhlogarra
(Iraq blind barb) Vinalabeo
Vinalabeo
[34] Zacco

See also[edit]

List of fish families Media related to Cyprinidae
Cyprinidae
at Wikimedia Commons Data related to Cyprinidae
Cyprinidae
at Wikispecies

References[edit]

^ a b Froese, Rainer, and Daniel Pauly, eds. (2015). "Cyprinidae" in FishBase. July 2015 version. ^ a b Eschmeyer, W.N.; Fong, J.D. (2015). " Species
Species
by family/subfamily". Catalog of Fishes. California Academy of Science. Retrieved 2 July 2015.  ^ Nelson, Joseph (2006). Fishes of the World. Chichester: John Wiley & Sons. ISBN 0-471-25031-7.  ^ Froese, Rainer and Pauly, Daniel, eds. (2015). "Catlocarpio siamensis" in FishBase. March 2015 version. ^ Froese, Rainer and Pauly, Daniel, eds. (2017). "Tor putitora" in FishBase. March 2017 version. ^ Froese, Rainer and Pauly, Daniel, eds. (2017). "Luciobarbus esocinus" in FishBase. March 2017 version. ^ Froese, Rainer and Pauly, Daniel, eds. (2015). "Ptychocheilus lucius" in FishBase. March 2015 version. ^ Froese, Rainer and Pauly, Daniel, eds. (2015). "Paedocypris progenetica" in FishBase. March 2015 version. ^ Based on data sourced from the FishStat database ^ MacMahon, Alexander Francis Magri (1946). Fishlore: British Freshwater Fishes. Pelican Books. 161. Penguin Books. pp. 149–152.  ^ Gulf States Marine Fisheries Commission
Gulf States Marine Fisheries Commission
(3 August 2005). "Cyprinus carpio (Linnaeus, 1758)". Archived from the original on 18 August 2007. Retrieved 3 May 2007.  ^ Florida
Florida
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External links[edit]

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Carp

Carp
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groups

Cyprinidae
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(the carp family) Asian carp

True carp species

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Other carp species

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Diseases

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Recreational fishing

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As food

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Related topics

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Taxon identifiers

Wd: Q35047 ADW: Cyprinidae EoL: 3198 EPPO: 1CYPNF Fauna Europaea: 12021 Fossilworks: 64192 GBIF: 7336 ITIS: 163342 NCBI: 7953 WoRMS: 154163

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