Crotalus molossus
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The black-tailed rattlesnake (''Crotalus molossus'') is a
venomous Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
pit viper The Crotalinae, commonly known as pit vipers,Mehrtens JM (1987). ''Living Snakes of the World in Color''. New York: Sterling Publishers. 480 pp. . crotaline snakes (from grc, κρόταλον ''krotalon'' castanet), or pit adders, are a subfa ...
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
found in the southwestern
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country Continental United States, primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., ...
and
Mexico Mexico (Spanish language, Spanish: México), officially the United Mexican States, is a List of sovereign states, country in the southern portion of North America. It is borders of Mexico, bordered to the north by the United States; to the so ...
. Four
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. Not all specie ...
are currently recognized, including the nominate subspecies described here.


Taxonomy

A recent revision showed that eastern populations from Texas and central and eastern New Mexico form a distinct species separate from ''C. molossus'': ''Crotalus ornatus'' Hallowell 1854. Alternate common names are green rattler,Campbell JA, Lamar WW. 2004. The Venomous Reptiles of the Western Hemisphere. Comstock Publishing Associates, Ithaca and London. 870 pp. 1500 plates. . and Northern black-tailed rattlesnake. Wright AH, Wright AA. 1957. ''Handbook of Snakes of the United States and Canada''. Comstock Publishing Associates. Ithaca and London. (7th printing, 1985). 1105 pp. .


Subspecies


Description

This medium-sized species averages from in length. Conant R. 1975. ''A Field Guide to Reptiles and Amphibians of Eastern and Central North America. Second Edition.'' irst published in 1958 Houghton Mifflin Company. Boston. 429 pp. 48 plates. (hc), (pb). Large specimens are usually not much more than long, although lengths of ( Gloyd, 1940), ( Klauber, 1972), and (Shaw & Campbell, 1974) have been reported. The females tend to be larger than the males. They range in color from yellows and olive greens to browns and black. As their name implies, one of their most distinguishing features is, despite variations in body color, entirely black tail scales. Often, this rattlesnake has a black band across its eyes extending diagonally down to the corners of its mouth, forming a sort of facial 'mask'. Like other rattlesnakes, ''C. molossus'' has a rattle composed of
keratin Keratin () is one of a family of structural fibrous proteins also known as ''scleroproteins''. Alpha-keratin (α-keratin) is a type of keratin found in vertebrates. It is the key structural material making up Scale (anatomy), scales, hair, Nail ...
on the end of its tail. Each time the
snake Snakes are elongated, limbless, carnivorous reptiles of the suborder Serpentes . Like all other squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales. Many species of snakes have skulls with several more ...
sheds its skin, a new segment is added to the rattle. A snake can shed its skin several times a year, and the rattle is fairly fragile and can be broken, so that the length of a
rattlesnake Rattlesnakes are venomous snakes that form the genera ''Crotalus'' and ''Sistrurus'' of the subfamily Crotalinae (the pit vipers). All rattlesnakes are vipers. Rattlesnakes are predators that live in a wide array of habitats, hunting small an ...
's rattle is not an accurate measure of its age, unless the terminal button is intact.


Distribution and habitat

The black-tailed rattlesnake is found in the southwestern
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country Continental United States, primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., ...
in
Arizona Arizona ( ; nv, Hoozdo Hahoodzo ; ood, Alĭ ṣonak ) is a state in the Southwestern United States. It is the 6th largest and the 14th most populous of the 50 states. Its capital and largest city is Phoenix. Arizona is part of the Fou ...
,
New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe, New Mexico, Santa Fe , LargestCity = Albuquerque, New Mexico, Albuquerque , LargestMetro = Albuquerque metropolitan area, Tiguex , Offi ...
and west and central
Texas Texas (, ; Spanish: ''Texas'', ''Tejas'') is a state in the South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2020, it is the second-largest U.S. state by ...
, and
Mexico Mexico (Spanish language, Spanish: México), officially the United Mexican States, is a List of sovereign states, country in the southern portion of North America. It is borders of Mexico, bordered to the north by the United States; to the so ...
as far south as
Oaxaca Oaxaca ( , also , , from nci, Huāxyacac ), officially the Free and Sovereign State of Oaxaca ( es, Estado Libre y Soberano de Oaxaca), is one of the 32 states that compose the Federative Entities of Mexico. It is divided into 570 municipaliti ...
. Also it is found in the
Gulf of California The Gulf of California ( es, Golfo de California), also known as the Sea of Cortés (''Mar de Cortés'') or Sea of Cortez, or less commonly as the Vermilion Sea (''Mar Bermejo''), is a marginal sea of the Pacific Ocean that separates the Baja C ...
on San Estéban and
Tiburón Island Tiburón Island is the largest island in the Gulf of California and the largest island in Mexico, with an area of . It was made a nature reserve in 1963 by President Adolfo López Mateos. Etymology is Spanish for ' shark'. Although the Seri ...
s. Its distribution reaches a maximum elevation of 2930 m, although it has been recorded at as high as 6900 feet at the McDonald Observatory in the Davis Mountains of west Texas. The type locality given is "Fort Webster, St. Rita del Cobre, N. Mex." (Fort Webster, Santa Rita del Cobre, Grant County, New Mexico, USA).


Conservation status

This species is classified as Least Concern
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biolo ...
of Threatened Species (v3.1, 2001). Species are listed as such owing to their wide distribution or presumed large population or because they are unlikely to be declining fast enough to qualify for listing in a more threatened category. The population trend was stable when assessed in 2007.


Behavior

All
rattlesnakes Rattlesnakes are venomous snakes that form the genera '' Crotalus'' and '' Sistrurus'' of the subfamily Crotalinae (the pit vipers). All rattlesnakes are vipers. Rattlesnakes are predators that live in a wide array of habitats, hunting small an ...
are
carnivorous A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose food and energy requirements derive from animal tissues (mainly muscle, fat and other ...
, their primary food sources being
rodents Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are na ...
, other small
mammals Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur o ...
,
birds Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
, and small
reptiles Reptiles, as most commonly defined are the animals in the class Reptilia ( ), a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates ( lizards and snakes) and rhynchoceph ...
(such as small
lizard Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia altho ...
s).https://animaldiversity.org/accounts/Crotalus_molossus/ The behavior of northern black-tailed
rattlesnakes Rattlesnakes are venomous snakes that form the genera '' Crotalus'' and '' Sistrurus'' of the subfamily Crotalinae (the pit vipers). All rattlesnakes are vipers. Rattlesnakes are predators that live in a wide array of habitats, hunting small an ...
varies over the course of a year. In the spring and fall, they are primarily diurnal. In the summer, they shift to a
nocturnal Nocturnality is an animal behavior characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnal meaning the opposite. Nocturnal creatures generally have highly developed sens ...
behavior, to avoid the heat of summer. In the winter, they hibernate in dens created and abandoned by other animals, often with other species of
snake Snakes are elongated, limbless, carnivorous reptiles of the suborder Serpentes . Like all other squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales. Many species of snakes have skulls with several more ...
s. They are variable in their form of locomotion depending on what substrate they need to traverse and will actively change between sidewinding or rectilinear movement. Although it is an able climber and expert swimmer, ''C. molossus'' is primarily a terrestrial species and inhabits grasslands, desert areas, and rocky and mountainous areas, as well as high-altitude forested


Docility and defense

''Crotalus molossus'' is considered to be one of the most docile rattlesnakes because of its calm demeanor and curious nature. Bites are accordingly fairly rare. The snake relies mostly on camouflage to avoid discovery. It normally tries to slither away when confronted, but will rattle when cornered.


Venom

The venom of the ''C. molossus'' is primarily
hemotoxic Hemotoxins, haemotoxins or hematotoxins are toxins that destroy red blood cells, disrupt blood clotting, and/or cause organ degeneration and generalized tissue damage. The term ''hemotoxin'' is to some degree a misnomer since toxins that damage t ...
, like that of most crotalids. However, it is only about two-thirds as toxic as western diamondback venom and is generally not fatal to humans.
CroFab Crotalidae polyvalent immune fab, sold under the brandname CroFab, is a snake antivenin, indicated for minimal or moderate North American Crotalid (Rattlesnake, Copperhead and Cottonmouth/Water moccasin) snake envenomation. CroFab is composed of ...
antivenom Antivenom, also known as antivenin, venom antiserum, and antivenom immunoglobulin, is a specific treatment for envenomation. It is composed of antibodies and used to treat certain venomous bites and stings. Antivenoms are recommended only if th ...
is often used to treat bites. ''Crotalus molossus'' has larger venom glands than most rattlesnakes in its region. Because its venom is less toxic than that of most other crotalids, it needs to inject large quantities into its prey to be effective.


Reproduction

Breeding occurs in the spring when males follow the
pheromone A pheromone () is a secreted or excreted chemical factor that triggers a social response in members of the same species. Pheromones are chemicals capable of acting like hormones outside the body of the secreting individual, to affect the behavio ...
trails of the females. Copulation can sometimes last for hours and occur multiple times over a period of days. After mating, the male often stays near the female for several days to prevent any other males from mating with her. The female gives birth to live young in the summer, and the babies stay with the mother only until they wander off on their own, usually in less than a day or two. Females are believed to breed every year, and can have litters as large as 10-12 young but usually averaging four to six. Their lifespans average 15–20 years.


References


External links

*
Crotalus molossus
a
Herps of Texas
Accessed 24 February 2007.

a
Centennial Museum & Chihuahuan Desert Gardens Natural and Cultural History
Accessed 24 February 2007.

a
California Reptiles and Amphibians
Accessed 24 February 2007.

a
Munich AntiVenom INdex
Accessed 24 February 2007. {{Taxonbar, from=Q2254464 molossus Tiburón Island Reptiles described in 1853 Taxa named by Spencer Fullerton Baird