Crambinae
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Crambinae is a large
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus Genus ( plural genera ) is a taxonomic rank used in the biological classifica ...
of the
lepidoptera Lepidoptera ( ) is an order of insects that includes butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 families and 46 superfamilies, 10 percent of the total described speci ...
n family
Crambidae The Crambidae are the grass moth family of lepidopterans. They are variable in appearance, the nominal subfamily Crambinae (grass moths) taking up closely folded postures on grass stems where they are inconspicuous, while other subfamilies includ ...
, the crambid snout
moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...
s. It currently includes over 1,800
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
worldwide. The larvae are root feeders or stem borers, mostly on grasses. A few species are pests of sod grasses, maize, sugar cane, rice, and other Poaceae. The
monophyly In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic ...
of this group is supported by the structure of the
tympanal organ A tympanal organ (or tympanic organ) is a hearing organ in insects, consisting of a membrane ( tympanum) stretched across a frame backed by an air sac and associated sensory neurons. Sounds vibrate the membrane, and the vibrations are sensed by ...
s and the phallus attached medially to the juxta, as well as genetic analyses. Taxonomists' opinions differ as to the correct placement of the
Crambidae The Crambidae are the grass moth family of lepidopterans. They are variable in appearance, the nominal subfamily Crambinae (grass moths) taking up closely folded postures on grass stems where they are inconspicuous, while other subfamilies includ ...
, some authorities treating them as a subfamily of the family Pyralidae. If this is done, the present group would be demoted to
tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide usage of the term in English is in the discipline of anthropology. This definition is contested, in part due to confl ...
status, as Crambini.


Crambinae taxonomy

*tribe ''
incertae sedis ' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertain ...
'' **'' Anaclastis'' Turner, 1904 **'' Aphrophantis'' Meyrick, 1933 **'' Argentochiloides'' Błeszyński, 1961 (= ''Argentochilo'' Błeszyński & Collins, 1962) **'' Australargyria'' Błeszyński, 1970 **'' Autarotis'' Meyrick, 1886 (= ''Pogonoptera'' Turner, 1911) **'' Batiana'' Walker, 1866 **'' Burmannia'' Bleszynski, 1965 **'' Catancyla'' Hampson, 1919 **'' Catharylla'' Zeller, 1863 **'' Charltona'' Swinhoe, 1886 **'' Charltoniada'' Strand, 1918 **'' Conocrambus'' Hampson, 1919 (= ''Anaresca'' Turner, 1947) **'' Corynophora'' Berg, 1898 (= ''Halterophora'' Meyrick, 1897) **'' Cypholomia'' Meyrick, 1933 **'' Diadexia'' Turner, 1905 **'' Diploptalis'' Hampson, 1919 (= ''Diplotalis'' Błeszyński, 1963) **'' Diploschistis'' Meyrick, 1937 **'' Elethyia'' Ragonot in de Joannis & Ragonot, 1889 **'' Epina'' Walker, 1866 (= ''Diatraenopsis'' Dyar & Heinrich, 1927) **'' Eurhythma'' Turner, 1904 (= ''Eurythma'' Błeszyński & Collins, 1962) **'' Hemiplatytes'' Barnes & Benjamin, 1924 (= ''Alamogordia'' Dyar & Heinrich, 1927) **'' Hemiptocha'' Dognin, 1905 **'' Idioblasta'' Warren, 1891 **'' Japonichilo'' Okano, 1962 **'' Kupea'' Philpott, 1930 **'' Libuna'' Moore, 1886 (= ''Bulina'' Walker, 1866) **'' Lyndia'' Savigny, 1816 **'' Maoricrambus'' Gaskin, 1975 **''
Mestolobes ''Mestolobes'' is a genus of moths of the family Crambidae described by Arthur Gardiner Butler in 1882. All species are endemic to the Hawaiian Islands. Species *''Mestolobes abnormis'' (Butler, 1882) *''Mestolobes amethystias'' Meyrick, 1899 ...
'' Butler, 1882 (= ''Promylaea'' Meyrick, 1899) **'' Metaeuchromius'' Błeszyński, 1960 (= ''Pseudeuchromius'' Błeszyński, 1965) **'' Micrelephas'' Dognin, 1905 **'' Microchilo'' Okano, 1962 **'' Micronix'' Amsel, 1956 **'' Microtalis'' Turner, 1911 **'' Neargyria'' Hampson, 1896 **'' Neargyrioides'' Błeszyński, 1970 **'' Nechilo'' Błeszyński, 1970 **'' Neobanepa'' Hampson, 1919 (= ''Pyralopsis'' Dognin, 1905) **'' Niveocatharylla'' Bassi, 1999 **'' Orthomecyna'' Butler, 1883 **'' Parancyla'' Hampson, 1919 **'' Paratraea'' Hampson, 1919 **'' Parerupa'' Hampson, 1919 (= ''Coenotalis'' Hampson, 1919) **'' Protyparcha'' Meyrick, 1909 **'' Pseudargyria'' Okano, 1962 **'' Pseudometachilo'' Błeszyński, 1962 **'' Ptochostola'' Meyrick, 1882 **'' Roxita'' Błeszyński, 1963 (= ''Modestia'' Błeszyński, 1965) **'' Schoenobiodes'' Hampson, 1917 (= ''Stenopydna'' Roepke, 1943) **'' Sphaerodeltis'' Meyrick, 1933 **'' Styxon'' Błeszyński, 1962 **'' Thalamarchis'' Meyrick, 1897 **'' Tomissa'' Walker, 1864 **'' Tulla'' Zimmerman, 1958 **'' Ubida'' Walker, 1863 (= ''Crunophila'' Meyrick, 1882) **'' Zacatecas'' Błeszyński, 1962 *tribe Ancylolomiini Ragonot, 1889 **''
Ancylolomia ''Ancylolomia'' is a genus of moths of the family Crambidae described by Jacob Hübner in 1825. Description Palpi porrect (stretched forward), thickly clothed with hair, and extending about three times the length of head. Maxillary palp trian ...
'' Hübner, 1825 (= ''Acylolomia'' Hampson, 1919, ''Jartheza'' Walker, 1863, ''Pseudoctenella'' Strand, 1907, ''Ctenus'' Mabille, 1906, ''Tollia'' Amsel, 1949) **'' Aurotalis'' Błeszyński, 1970 **'' Eufernaldia'' Hulst, 1900 **'' Mesolia'' Ragonot in de Joannis & Ragonot, 1889 (= ''Deuterolia'' Dyar, 1914, ''Eugrotea'' Fernald, 1896, ''Euparolia'' Dyar, 1914, ''Masolia'' Hampson, 1919) **'' Prionapteron'' Błeszyński, 1965 **'' Prionapteryx'' Stephens, 1834 (= ''Alloea'' Turner, 1947, ''Calarina'' Walker, 1866, ''Hypotomorpha'' Rebel, 1892, ''Nuarace'' Walker, 1863, ''Naurace'' Błeszyński & Collins, 1962, ''Pindicitora'' Walker, 1863, ''Pindicitra'' Shibuya, 1928, ''Platytesia'' Strand, 1918, ''Prionopteryx'' Zeller, 1863) **'' Prionotalis'' Hampson, 1919 **'' Pseudoschoenobius'' Fernald, 1891 **'' Surattha'' Walker, 1863 **'' Talis'' Guenée, 1845 (= ''Drasa'' Kapur, 1950, ''Prosmixis'' Zeller, 1846, ''Tulis'' Pagenstecher, 1909) **'' Zovax'' Błeszyński, 1962 *tribe Argyriini Munroe, 1995 **'' Argyria'' Hübner, 1818 **'' Urola'' Walker, 1863 **'' Vaxi'' Błeszyński, 1962 *tribe Calamotrophini Gaskin, 1988 **'' Calamotropha'' Zeller, 1863 (= ''Aurelianus'' Błeszyński, 1962, ''Myeza'' Walker, 1863) **'' Chrysocatharylla'' Bassi, 1999 **'' Classeya'' Błeszyński, 1960 **'' Pseudocatharylla'' Błeszyński, 1961 **'' Pseudoclasseya'' Błeszyński, 1964 *tribe Chiloini Heinemann, 1865 **'' Chilandrus'' Błeszyński, 1970 **''
Chilo Chilo may refer to: * Chilo of Sparta, one of the Seven Sages of Greece * Chilo, Ohio, a village in Clermont County, Ohio * Chilo, Bhutan * ''Chilo'' (moth), a genus of crambid moths * André Chilo (1898–1982), French rugby player * Chilo Rachal ...
'' Zincken, 1817 (= ''Borer'' Guenée, 1862, ''Chilona'' Sodoffsky, 1837, ''Chilotraea'' Kapur, 1950, ''Diphryx'' Grote, 1881, ''Hypiesta'' Hampson, 1919, ''Nephalia'' Turner, 1911, ''Silveria'' Dyar, 1925) **'' Chiqua'' Błeszyński, 1970 **'' Diatraea'' Guilding, 1828 (= ''Crambidiatraea'' Box & Capps, 1955, ''Diaraetria'' Grote, 1882, ''Diatraerupa'' Schaus, 1913, ''Diatrea'' Guilding, 1828, ''Eodiatraea'' Box, 1953, ''Iesta'' Dyar, 1909, ''Trinidadia'' Dyar & Heinrich, 1927, ''Zeadiatraea'' Box, 1955) **'' Eschata'' Walker, 1856 (= ''Chaerecla'' Walker, 1865) **'' Gadira'' Walker, 1866 (= ''Cryptomima'' Meyrick, 1882, ''Scenoploca'' Meyrick, 1882) **'' Hednota'' Meyrick, 1886 **''
Leonardo Leonardo is a masculine given name, the Italian, Spanish, and Portuguese equivalent of the English, German, and Dutch name, Leonard. People Notable people with the name include: * Leonardo da Vinci (1452–1519), Italian Renaissance scientist ...
'' Błeszyński, 1965 **'' Malgasochilo'' Błeszyński, 1970 **'' Myelobia'' Herrich-Schäffer, 1854 (= ''Chilopsis'' Hampson, 1919, ''Doratoperas'' Hampson, 1896, ''Protaphomia'' Meyrick, 1936, ''Xanthopherne'' Dyar & Heinrich, 1927) **'' Tauroscopa'' Meyrick, 1888 (= ''Oressaula'' Turner, 1913) *tribe Crambini Latreille, 1810 **'' Agriphila'' Hübner, 1825 (= ''Agrophila'' J. L. R. Agassiz, 1847, ''Alisa'' Ganev & Hacker, 1984) **'' Agriphiloides'' Błeszyński, 1965 **'' Almita'' B. Landry, 1995 **'' Alphacrambus'' Bassi, 1995 **'' Amselia'' Błeszyński, 1959 **'' Angustalius'' Marion, 1954 (= ''Bleszynskia'' Lattin, 1961, ''Crambopsis'' Lattin, 1952) **''
Arequipa Arequipa (; Aymara and qu, Ariqipa) is a city and capital of province and the eponymous department of Peru. It is the seat of the Constitutional Court of Peru and often dubbed the "legal capital of Peru". It is the second most populated city ...
'' Walker, 1863 **'' Aureocramboides'' Błeszyński, 1961 **'' Bassiknysna'' Kemal & Kocak, 2005 (= ''Knysna'' Bassi, 1999) **'' Caffrocrambus'' Błeszyński, 1961 (= ''Anomocrambus'' Błeszyński, 1961, ''Caffocrambus'' Bassi, 1994) **'' Catoptria'' Hübner, 1825 (= ''Exoria'' Hübner, 1825, ''Tetrachila'' Hübner, 1806) **'' Cervicrambus'' Błeszyński, 1966 **'' Chrysocrambus'' Błeszyński, 1957 (= ''Chrysocramboides'' Błeszyński, 1957) **'' Chrysoteuchia'' Hübner, 1825 (= ''Amphibolia'' Snellen, 1884, ''Veronese'' Błeszyński, 1962) **'' Conocramboides'' Błeszyński, 1970 **'' Crambixon'' Błeszyński, 1965 **'' Crambus'' Fabricius, 1798 (= ''Argyroteuchia'' Hübner, 1825, ''Chilus'' Billberg, 1820, ''Palparia'' Haworth, 1811, ''Tetrachila'' Hübner, 1822) **'' Culladia'' Moore, 1886 (= ''Araxes'' Walker, 1863, ''Crambidion'' Mabille, 1900, ''Nirmaladia'' Rose, 1983) **'' Culladiella'' Błeszyński, 1970 **'' Dimorphocrambus'' Gibeaux, 1987 **'' Epichilo'' Ragonot in de Joannis & Ragonot, 1889 **'' Fernandocrambus'' Aurivillius, 1922 (= ''Juania'' Aurivillius, 1922, ''Juanita'' Munroe, 1995) **'' Fissicrambus'' Błeszyński, 1963 **'' Flavocrambus'' Błeszyński, 1959 **'' Haplopediasia'' Błeszyński, 1963 **'' Haploplatytes'' Błeszyński, 1966 **'' Japonicrambus'' Okano, 1962 **'' La'' Błeszyński, 1966 **'' Loxocrambus'' Forbes, 1920 **'' Mesocrambus'' Błeszyński, 1957 **'' Mesopediasia'' Błeszyński, 1963 **'' Metacrambus'' Błeszyński, 1957 **'' Microcramboides'' Błeszyński, 1967 **'' Microcrambon'' Błeszyński, 1970 **'' Microcrambus'' Błeszyński, 1963 **'' Miraxis'' Błeszyński, 1962 **'' Miyakea'' Marumo, 1933 **'' Neocrambus'' Błeszyński, 1957 **'' Neoculladia'' Błeszyński, 1967 **'' Neodactria'' B. Landry, 1995 **'' Neopediasia'' Okano, 1962 **'' Novocrambus'' Amsel, 1956 **'' Orocrambus'' Purdie, 1884 (= ''Orocrambus'' Meyrick, 1885) **'' Parapediasia'' Błeszyński, 1966 (''Parapediasia'' Błeszyński, 1963) **'' Paraplatytes'' Błeszyński, 1965 **'' Pediasia'' Hübner, 1825 (= ''Carvanca'' Walker, 1856, ''Oseriates'' Fazekas, 1991, ''Pseudopediasia'' Ganev, 1987) **'' Platytes'' Guenée, 1845 (= ''Nagahama'' Marumo, 1933) **'' Precaffrocrambus'' Bassi, 2002 **'' Productalius'' Marion, 1954 **'' Pseudopediasia'' Błeszyński, 1963 **'' Raphiptera'' Hampson, 1896 **'' Sebrus'' Błeszyński, 1970 **'' Sericocrambus'' Wallengren, 1861 **'' Supercrambus'' Błeszyński, 1967 **'' Tawhitia'' Philpott, 1931 (= ''Velasquez'' Błeszyński, 1962) **'' Tehama'' Hulst, 1888 **'' Thaumatopsis'' Morrison, 1874 (= ''Propexus'' Grote, 1880) **'' Thisanotia'' Hübner, 1825 (= ''Thinasotia'' Heinemann, 1865, ''Thysanotia'' J. L. R. Agassiz, 1847) **'' Tortriculladia'' Błeszyński, 1967 **'' Xanthocrambus'' Błeszyński, 1957 (= ''Xanthocrambus'' Błeszyński, 1955) *tribe Diptychophorini Gaskin, 1972 **'' Cleoeromene'' Gaskin, 1986 **'' Diptychophora'' Zeller, 1866 (= ''Colimaea'' Dyar, 1925, ''Colimea'' Błeszyński, 1966, ''Mysticomima'' Meyrick, 1931, ''Scissolia'' Barnes & McDunnough, 1914) **'' Gargela'' Walker, 1864 (= ''Mixophyla'' Meyrick, 1887, ''Angonia'' Snellen, 1893, ''Mixophila'' Hampson, 1896) **'' Glaucocharis'' Meyrick, 1938 (= ''Pagmania'' Amsel, 1961, ''Pareromene'' Osthelder, 1941, ''Ditomoptera'' Hampson, 1893) **'' Incaeromene'' Gaskin, 1986 **'' Neoeromene'' Gaskin, 1986 **'' Peniculimius'' Schouten, 1994 **'' Steneromene'' Gaskin, 1986 **'' Tamsica'' Zimmerman, 1958 *tribe Euchromiini Léger, Landry & Nuss, 2019 **'' Euchromius'' Guenée, 1845 (= ''Eromene'' Hübner, 1825, ''Ommatopteryx'' Kirby, 1897, ''Pseudoancylolomia'' Ahmad, Zaidi & Kamaluddin, 1982) *tribe Haimbachiini B. Landry, 1995 **'' Achilo'' Amsel, 1957 (= ''Chilopsis'' Amsel, 1956) **'' Bissetia'' Kapur, 1950 (= ''Girdharia'' Kapur, 1950) **'' Coniesta'' Hampson, 1919 **'' Donacoscaptes'' Zeller, 1877 **''
Eoreuma ''Eoreuma'' is a genus of moths of the family Crambidae The Crambidae are the grass moth family of lepidopterans. They are variable in appearance, the nominal subfamily Crambinae (grass moths) taking up closely folded postures on grass stems w ...
'' Ely, 1910 **'' Friedlanderia'' Agnew, 1987 (= ''Chiloides'' Amsel, 1949, ''Chiloides'' Błeszyński, 1963) **'' Haimbachia'' Dyar, 1909 **'' Neogirdharia'' Song & Chen in Chen, Song & Yuan, 2004 **'' Occidentalia'' Dyar & Heinrich, 1927 **'' Pseudobissetia'' Błeszyński, 1959 **'' Thopeutis'' Hübner, 1818 (= ''Cephis'' Ragonot in Staudinger, 1892, ''Hombergia'' de Joannis, 1910, ''Stenochilo'' Hampson, 1896, ''Tetrachila'' Hübner, 1808, ''Topeutis'' Hübner, 1825) **'' Xubida'' Schaus, 1922


Former genera

*'' Araxates'' Ragonot in de Joannis & Ragonot, 1889 *'' Charitopepla'' Meyrick, 1933 *'' Neerupa'' Hampson, 1919 *'' Loxophantis'' Meyrick, 1935 *'' Welaka'' Hulst, 1888


See also

* List of crambid genera


References

{{Taxonbar, from=Q149407 Crambidae Taxa named by Pierre André Latreille Moth subfamilies