Cornus nuttallii
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''Cornus nuttallii'', the Pacific dogwood, western dogwood, or mountain dogwood, is a species of
dogwood ''Cornus'' is a genus of about 30–60 species of woody plants in the family Cornaceae, commonly known as dogwoods, which can generally be distinguished by their blossoms, berries, and distinctive bark. Most are deciduous trees or ...
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
native to western
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and th ...
.


Description

It is a small to medium-sized
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, ...
tree, reaching tall, often with a canopy spread of . Its
habit A habit (or wont as a humorous and formal term) is a routine of behavior that is repeated regularly and tends to occur subconsciously.
varies based on the level of sunlight; in full sun it will have a short trunk with a crown as wide as it is tall, while under a canopy it will have a tapered trunk with a short, slender crown. The trunk attains in diameter. The
bark Bark may refer to: * Bark (botany), an outer layer of a woody plant such as a tree or stick * Bark (sound), a vocalization of some animals (which is commonly the dog) Places * Bark, Germany * Bark, Warmian-Masurian Voivodeship, Poland Arts, e ...
is reddish brown. The branches have fine hairs and the young bark is thin and smooth, becoming scale-like with ridges as it ages. The
leaves A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, st ...
are opposite, simple, oval, long, and broad. They are green with stiff, appressed hairs on top, and hairier and lighter on the bottom. They turn orange to purplish in autumn. The
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechanis ...
s are individually small and inconspicuous, across, produced in a dense, rounded, greenish-white
flower head A pseudanthium (Greek for "false flower"; ) is an inflorescence that resembles a flower. The word is sometimes used for other structures that are neither a true flower nor a true inflorescence. Examples of pseudanthia include flower heads, compos ...
in diameter; the 4–8 large white 'petals' are actually
bract In botany, a bract is a modified or specialized leaf, especially one associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves. They may be smaller, larger, or o ...
s, each bract long and broad, creating the appearance of a larger flower head. The flowers commonly bloom twice per season, once in the spring and again in late summer or early fall. The
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in partic ...
is a compound pink-red or orange
drupe In botany, a drupe (or stone fruit) is an indehiscent fruit in which an outer fleshy part ( exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pit'', ''stone'', or ''pyrena'') of hardened endocarp with a seed (''kernel ...
about long, in clusters containing 20–40 drupelets, each of which contains two seeds. They appear in September or October.


Similar species

The eastern United States' ''
Cornus florida ''Cornus florida'', the flowering dogwood, is a species of flowering tree in the family Cornaceae native to eastern North America and northern Mexico. An endemic population once spanned from southernmost coastal Maine south to northern Florida ...
'' (flowering dogwood) is similar in appearance and possibly in chemical composition. ''
Cornus canadensis ''Cornus canadensis'' is a species of flowering plant in the dogwood family Cornaceae, native to eastern Asia and North America. Common names include Canadian dwarf cornel, Canadian bunchberry, quatre-temps, crackerberry, and creeping dogwood. Un ...
'' has similar blossoms but grows as a
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from tree ...
.


Etymology

In 1806,
Meriwether Lewis Meriwether Lewis (August 18, 1774 – October 11, 1809) was an American explorer, soldier, politician, and public administrator, best known for his role as the leader of the Lewis and Clark Expedition, also known as the Corps of Discovery, wit ...
noted that the species is similar in appearance to ''C. florida''. However, when Scottish botanist David Douglas encountered ''C. nuttalli'' on his expedition to the Pacific Northwest in the 1820s, he mistook it for ''C. florida'' and did not send seeds back to England. English botanist
Thomas Nuttall Thomas Nuttall (5 January 1786 – 10 September 1859) was an English botanist and zoologist who lived and worked in America from 1808 until 1841. Nuttall was born in the village of Long Preston, near Settle in the West Riding of Yorkshire and ...
was the first to describe the species for science while staying at
Fort Vancouver Fort Vancouver was a 19th century fur trading post that was the headquarters of the Hudson's Bay Company's Columbia Department, located in the Pacific Northwest. Named for Captain George Vancouver, the fort was located on the northern bank of ...
in the autumn of 1834. It was named ''nuttallii'' after him by his friend
John James Audubon John James Audubon (born Jean-Jacques Rabin; April 26, 1785 – January 27, 1851) was an American self-trained artist, naturalist, and ornithologist. His combined interests in art and ornithology turned into a plan to make a complete pictori ...
. The common names comes from that of ''
Cornus sanguinea ''Cornus sanguinea'', the common dogwood or bloody dogwood, is a species of dogwood native to most of Europe and western Asia, from England and central Scotland east to the Caspian Sea. It is widely grown as an ornamental plant. Description It ...
'', the hard wood of which Northern Europeans used to make nails ("dags") during the medieval era.


Distribution and habitat

It occurs from the lowlands of southern
British Columbia British Columbia (commonly abbreviated as BC) is the westernmost province of Canada, situated between the Pacific Ocean and the Rocky Mountains. It has a diverse geography, with rugged landscapes that include rocky coastlines, sandy beaches, for ...
to the mountains of southern
California California is a state in the Western United States, located along the Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the most populous U.S. state and the 3rd largest by area. It is also the m ...
. There exists an inland population in central
Idaho Idaho ( ) is a U.S. state, state in the Pacific Northwest region of the Western United States. To the north, it shares a small portion of the Canada–United States border with the province of British Columbia. It borders the states of Monta ...
, where it is considered
critically imperiled The NatureServe conservation status system, maintained and presented by NatureServe in cooperation with the Natural Heritage Network, was developed in the United States in the 1980s by The Nature Conservancy (TNC) as a means for ranking or categ ...
. It occurs predominantly below in elevation. Cultivated examples are found as far north as
Haida Gwaii Haida Gwaii (; hai, X̱aaydag̱a Gwaay.yaay / , literally "Islands of the Haida people") is an archipelago located between off the northern Pacific coast of Canada. The islands are separated from the mainland to the east by the shallow Heca ...
. It has high flood tolerance, and is common along streams with moist but well-drained soils, often on gentle slopes. Soil composition can range from clay to sandy loam, and it prefers a high humus content, moderate to high nutrient levels, and acidic soils with a pH from 5.5 to 6. It has low frost tolerance, and is usually found in low-elevation temperate or mesothermal climates. It is hardy to USDA zone 7.


Ecology


Value to animals

New sprouts are good
browse Browsing is a kind of orienting strategy. It is supposed to identify something of relevance for the browsing organism. When used about human beings it is a metaphor taken from the animal kingdom. It is used, for example, about people browsing o ...
for both wild and domesticated
ungulate Ungulates ( ) are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. These include odd-toed ungulates such as horses, rhinoceroses, and tapirs; and even-toed ungulates such as cattle, pigs, giraffes, ...
s, especially after a recent fire, but the mature foliage is usually ignored by all species except
slugs Slug, or land slug, is a common name for any apparently shell-less terrestrial molluscs, terrestrial gastropod mollusc. The word ''slug'' is also often used as part of the common name of any gastropod mollusc that has no shell, a very reduced ...
. The fruit are eaten by
deer mice Deer or true deer are hoofed ruminant mammals forming the family Cervidae. The two main groups of deer are the Cervinae, including the muntjac, the elk (wapiti), the red deer, and the fallow deer; and the Capreolinae, including the reindeer ...
,
pileated woodpecker The pileated woodpecker (''Dryocopus pileatus'') is a large, mostly black woodpecker native to North America. An insectivore, it inhabits deciduous forests in eastern North America, the Great Lakes, the boreal forests of Canada, and parts of the ...
s, the
band-tailed pigeon The band-tailed pigeon (''Patagioenas fasciata'') is a medium-sized bird of the Americas. Its closest relatives are the Chilean pigeon and the ring-tailed pigeon, which form a clade of ''Patagioenas'' with a terminal tail band and iridescent p ...
, and bears. It provides habitat and cover to small birds such as
Wilson's warbler Wilson's warbler (''Cardellina pusilla'') is a small New World warbler. It is greenish above and yellow below, with rounded wings and a long, slim tail. The male has a black crown patch; depending on the subspecies, that mark is reduced or absent ...
, and small mammals including the red tree vole.


Diseases

Like the related ''Cornus florida'', it is very susceptible to dogwood
anthracnose A plant canker is a small area of dead tissue, which grows slowly, often over years. Some cankers are of only minor consequence, but others are ultimately lethal and therefore can have major economic implications for agriculture and horticultur ...
, a
disease A disease is a particular abnormal condition that negatively affects the structure or function of all or part of an organism, and that is not immediately due to any external injury. Diseases are often known to be medical conditions that a ...
caused by the fungus ''
Discula destructiva ''Discula destructiva'' is a fungus in the family Gnomoniaceae which causes dogwood anthracnose, affecting populations of dogwood trees native to North America. It was introduced to the United States in 1978 and is distributed throughout the Ea ...
''. Fungal activity is greatest from May to July, although it can be active any time conditions are moist and the plant is growing. Infected leaves become blotched and drop, and defoliation can be extreme. Twigs and leaf buds are also impacted. This has killed many of the larger plants in the wild and has also restricted its use as an ornamental tree, to the point where it is considered threatening to the species in its native range.


Successional status

It is present in all stages of both primary and secondary
succession Succession is the act or process of following in order or sequence. Governance and politics *Order of succession, in politics, the ascension to power by one ruler, official, or monarch after the death, resignation, or removal from office of ...
from new colonization on glacial outwash or areas destroyed by the 1980 eruption of Mount St. Helens, to late seral and even climax communities. It is
shade tolerant In ecology, shade tolerance is a plant's ability to tolerate low light levels. The term is also used in horticulture and landscaping, although in this context its use is sometimes imprecise, especially in labeling of plants for sale in commercial ...
but prefers sunlight in sufficiently humid conditions. It is adapted to a wide variety of
fire regime A fire regime is the pattern, frequency, and intensity of the bushfires and wildfires that prevail in an area over long periods of time. It is an integral part of fire ecology, and renewal for certain types of ecosystems. A fire regime describes th ...
s, with intervals ranging from just one year on dry sites, to 500 years or more in moist, riparian zones. The tree can survive low severity wildfires which are not hot enough to kill buds protected by bark. After being severely burned, ''Cornus nuttali'' typically resprouts from the root crownhowever, the resulting shoots are so palatable to
mule deer The mule deer (''Odocoileus hemionus'') is a deer indigenous to western North America; it is named for its ears, which are large like those of the mule. Two subspecies of mule deer are grouped into the black-tailed deer. Unlike the related whi ...
that they are at risk of being killed by over-browsing.


Uses

Some
Plateau Indian Indigenous peoples of the Northwest Plateau, also referred to by the phrase Indigenous peoples of the Plateau, and historically called the Plateau Indians (though comprising many groups) are Indigenous peoples of the Americas, indigenous peoples ...
tribes such as the Nlaka'pamux used the bark as a brown dye. Those groups also used the bark for medicinal purposes as a blood purifier, lung strengthener, stomach treatment,
laxative Laxatives, purgatives, or aperients are substances that loosen stools and increase bowel movements. They are used to treat and prevent constipation. Laxatives vary as to how they work and the side effects they may have. Certain stimulant, lubri ...
, and
emetic Vomiting (also known as emesis and throwing up) is the involuntary, forceful expulsion of the contents of one's stomach through the mouth and sometimes the nose. Vomiting can be the result of ailments like food poisoning, gastroenteritis ...
. The berries are edible by humans, though not very palatable. It is mostly prized as a cultivated ornamental.


Culture

It has been the provincial flower of
British Columbia British Columbia (commonly abbreviated as BC) is the westernmost province of Canada, situated between the Pacific Ocean and the Rocky Mountains. It has a diverse geography, with rugged landscapes that include rocky coastlines, sandy beaches, for ...
since 1956. It was once protected by law in the province (in an act which also protected '' Rhododendron macrophyllum'' and ''
Trillium ovatum ''Trillium ovatum'', the Pacific trillium, also known as the western wakerobin, western white trillium, or western trillium, is a species of flowering plant in the family Melanthiaceae. It is found in western North America, from southern British ...
''), but this was repealed in 2002.


References


External links


Jepson Flora Project: ''Cornus nuttallii''

Plants of British Columbia: ''Cornus nuttallii''

Interactive Distribution Map of ''Cornus nuttallii''
* {{Taxonbar, from=Q220777 nuttallii Plants described in 1840 Trees of British Columbia Trees of the Northwestern United States Trees of the Southwestern United States Ornamental trees Provincial symbols of British Columbia Trees of mild maritime climate Flora without expected TNC conservation status