Conchiolins (sometimes referred to as conchins) are complex
protein
Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residues. Proteins perform a vast array of functions within organisms, including catalysing metabolic reactions, DNA replication, res ...
s which are secreted by a mollusc's outer epithelium (the mantle).
These proteins are part of a matrix of organic macromolecules, mainly proteins and polysaccharides, that assembled together form the microenvironment where crystals nucleate and grow. This organic matrix also holds and binds to the crystals of
aragonite
Aragonite is a carbonate mineral, one of the three most common naturally occurring crystal forms of calcium carbonate, (the other forms being the minerals calcite and vaterite). It is formed by biological and physical processes, including pre ...
which give such shells their stiffness.
The
ion
An ion () is an atom or molecule with a net electrical charge.
The charge of an electron is considered to be negative by convention and this charge is equal and opposite to the charge of a proton, which is considered to be positive by conve ...
s necessary to form calcium carbonate are also secreted by the mantle, but it is the tailored environment created by the organic matrix which causes aragonite (rather than calcite) crystals to nucleate, in much the same way that collagen nucleates hydroxyapatite crystals.
Conchiolin serves as a relatively flexible, crack-deflecting matrix for the mineral aggregate particles; its strength and the strong bonding of perlucin can in some cases (such as in the formation of nacre) give the finished material an impressive level of
toughness
In materials science and metallurgy, toughness is the ability of a material to absorb energy and plastically deform without fracturing.periostracum which is composed of the protein conchiolin. Some land snails (especially the taxa that have become adapted to living on acidic soils) have very thin, transparent, tan-colored shells, even as adults, and those shells are composed entirely of conchiolin.
Corneous
Corneous is a biological and medical term meaning horny, in other words made out of a substance similar to that of horns and hooves in some mammals.
The word is generally used to describe natural or pathological anatomical structures made out ...
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Mollusc shell
The mollusc (or molluskOften spelled mollusk shell in the USA; the spelling "mollusc" are preferred by ) shell is typically a calcareous exoskeleton which encloses, supports and protects the soft parts of an animal in the phylum Mollusca, wh ...
Sporopollenin
270px, SEM image of pollen grains
Sporopollenin is one of the most chemically inert biological polymers. It is a major component of the tough outer (exine) walls of plant spores and pollen grains. It is chemically very stable and is usually well ...