Coccothrinax
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''Coccothrinax'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of palms in the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Arecaceae The Arecaceae is a family of perennial flowering plants in the monocot order Arecales. Their growth form can be climbers, shrubs, tree-like and stemless plants, all commonly known as palms. Those having a tree-like form are called palm trees ...
. There are more than 50
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
described in the genus, plus many
synonyms A synonym is a word, morpheme, or phrase that means exactly or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are ...
and
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. Not all specie ...
. A new species (''
Coccothrinax spirituana ''Coccothrinax spirituana'' is a fan palm which is palm endemic to central Cuba. Populations have been found in Sancti Spíritus and Ciego de Ávila provinces. Its leaves are ash-grey in colour on their upper and lower surfaces, a characteristic ...
'') was described as recently as 2017. Many ''Coccothrinax'' produce thatch. In Spanish-speaking countries, ''guano'' is a common name applied to ''Coccothrinax'' palms. The species are native throughout the
Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean ...
, the
Bahamas The Bahamas (), officially the Commonwealth of The Bahamas, is an island country within the Lucayan Archipelago of the West Indies in the North Atlantic. It takes up 97% of the Lucayan Archipelago's land area and is home to 88% of the a ...
, extreme southern
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and ...
and southeastern
Mexico Mexico (Spanish language, Spanish: México), officially the United Mexican States, is a List of sovereign states, country in the southern portion of North America. It is borders of Mexico, bordered to the north by the United States; to the so ...
, but most of the species are known only from
Cuba Cuba ( , ), officially the Republic of Cuba ( es, República de Cuba, links=no ), is an island country comprising the island of Cuba, as well as Isla de la Juventud and several minor archipelagos. Cuba is located where the northern Caribb ...
.


Description

''Coccothrinax'' is a genus of small to medium-sized, fan palms with relatively slender stems and 8 to 22
palmate The following is a list of terms which are used to describe leaf morphology in the description and taxonomy of plants. Leaves may be simple (a single leaf blade or lamina) or compound (with several leaflets). The edge of the leaf may be regular ...
leaves. The stems are initially covered by fibrous leaf sheaths. These break down into a network of fibres or spines, eventually leaving a bare trunk covered with leaf scars. The undersides of the leaflets are often silvery-grey; this is reflected in the common name "silver palm", which is given to many species of ''Coccothrinax''. The base of the petiole is not split longitudinally. The absence of this trait is a distinguishing character that separates ''Coccothrinax'' from ''Thrinax''. ''Coccothrinax'' species bear branched
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are forme ...
s that are located among the leaves. The bisexual flowers, which are borne on short stalks, have between 6 and 13
stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the fila ...
s and a single
carpel Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) '' pistils' ...
. The fruit are small, single-seeded, and range in colour from purple-red to purple-black, to brown. The seeds are deeply grooved "and resemble a brain".


Taxonomy

The genus ''Coccothrinax'' was first described by American botanist
Charles Sprague Sargent Charles Sprague Sargent (April 24, 1841 – March 22, 1927) was an American botanist. He was appointed in 1872 as the first director of Harvard University's Arnold Arboretum in Boston, Massachusetts, and held the post until his death. He pub ...
in 1899. He split the genus away from ''
Thrinax ''Thrinax'' is a genus in the palm family, native to the Caribbean. It is closely related to the genera '' Coccothrinax'', '' Hemithrinax'' and ''Zombia''. Flowers are small, bisexual and are borne on small stalks. Taxonomy In the first editi ...
'' based on characteristics of the fruit and seeds. The genus was based on Sargent's description of ''C. jucunda'' (now '' C. argentata'') and ''C. garberi'' (also synonymised with ''C. argentata''). He also assigned '' Thrinax argentea'' and '' T. radiata'' to the genus; while the former is now recognised as part of ''Coccothrinax'', the latter remains in ''Thrinax''. The generic epithet combines "coccus", the Latin word for berry, with ''Thrinax''. ''Coccothrinax'' is the most diverse genus of Caribbean palms. Although Andrew Henderson and colleagues only recognised fourteen species in their 1995 ''Field Guide to the Palms of the Americas'', the current ''World Checklist of Arecaceae'' recognises 52 or 53 species. ''Coccothrinax'' is placed in the tribe
Cryosophileae Cryosophileae is a tribe of palms in the subfamily Coryphoideae. The tribe ranges from southern South America, through Central America, into Mexico and the Caribbean. It includes New World genera formerly included in the tribe Thrinacinae, ...
.


Distribution

''Coccothrinax'' is a primarily Caribbean genus—it is found throughout the insular Caribbean, and in adjacent areas of
Mexico Mexico (Spanish language, Spanish: México), officially the United Mexican States, is a List of sovereign states, country in the southern portion of North America. It is borders of Mexico, bordered to the north by the United States; to the so ...
and
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and ...
. Species are usually found in dry, open or exposed habitats, on
limestone Limestone ( calcium carbonate ) is a type of carbonate sedimentary rock which is the main source of the material lime. It is composed mostly of the minerals calcite and aragonite, which are different crystal forms of . Limestone forms w ...
, serpentine or
sand Sand is a granular material composed of finely divided mineral particles. Sand has various compositions but is defined by its grain size. Sand grains are smaller than gravel and coarser than silt. Sand can also refer to a textural class ...
y soils. ''
Coccothrinax argentata ''Coccothrinax argentata'', commonly called the Florida silver palm, is a species of palm tree. It is native to south Florida, southeast Mexico, Colombia and to the West Indies, where it is found in the Bahamas, the southwest Caribbean and the T ...
'' ranges from
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and ...
and the
Florida Keys The Florida Keys are a coral cay archipelago located off the southern coast of Florida, forming the southernmost part of the continental United States. They begin at the southeastern coast of the Florida peninsula, about south of Miami, and e ...
, through the
Bahamas The Bahamas (), officially the Commonwealth of The Bahamas, is an island country within the Lucayan Archipelago of the West Indies in the North Atlantic. It takes up 97% of the Lucayan Archipelago's land area and is home to 88% of the a ...
and San Andrés Island to southeastern
Mexico Mexico (Spanish language, Spanish: México), officially the United Mexican States, is a List of sovereign states, country in the southern portion of North America. It is borders of Mexico, bordered to the north by the United States; to the so ...
. '' Coccothrinax barbadensis'' ranges through the
Lesser Antilles The Lesser Antilles ( es, link=no, Antillas Menores; french: link=no, Petites Antilles; pap, Antias Menor; nl, Kleine Antillen) are a group of islands in the Caribbean Sea. Most of them are part of a long, partially volcanic island arc be ...
to
Trinidad and Tobago Trinidad and Tobago (, ), officially the Republic of Trinidad and Tobago, is the southernmost island country in the Caribbean. Consisting of the main islands Trinidad and Tobago, and numerous much smaller islands, it is situated south of ...
and the
Netherlands Antilles nl, In vrijheid verenigd"Unified by freedom" , national_anthem = , common_languages = Dutch English Papiamento , demonym = Netherlands Antillean , capital = Willemstad , year_start = 1954 , year_end = 2010 , date_start = 15 December , ...
. The remaining species have narrower distributions; many are known from single populations in Cuba or Hispaniola.


Botanic gardens

Large collections are grown in
Fairchild Tropical Botanic Garden Fairchild Tropical Botanic Garden is an botanic garden with extensive collections of rare tropical plants including palms, cycads, flowering trees, and vines. It is located in the city of Coral Gables, Miami-Dade County, just south of Miami, s ...
(more than 250 plants, more than 23 species), in
Miami, Florida Miami ( ), officially the City of Miami, known as "the 305", "The Magic City", and "Gateway to the Americas", is a coastal metropolis and the county seat of Miami-Dade County in South Florida, United States. With a population of 442,241 at ...
, Jardín Botánico Nacional (15 species) in
Havana Havana (; Spanish: ''La Habana'' ) is the capital and largest city of Cuba. The heart of the La Habana Province, Havana is the country's main port and commercial center.
,
Cuba Cuba ( , ), officially the Republic of Cuba ( es, República de Cuba, links=no ), is an island country comprising the island of Cuba, as well as Isla de la Juventud and several minor archipelagos. Cuba is located where the northern Caribb ...
, and Palmetum of Santa Cruz de Tenerife (more than 1,000 ''Thrinax'' and ''Coccothrinx'' plants), in the
Canary Islands The Canary Islands (; es, :es:Canarias, Canarias, ), also known informally as the Canaries, are a Spanish Autonomous communities of Spain, autonomous community and archipelago in the Atlantic Ocean, in Macaronesia. At their closest point to ...
,
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = '' Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , ...
.


References


External links

* * {{Taxonbar, from=Q137792 Arecaceae genera Neotropical realm flora Taxa named by Charles Sprague Sargent