Club (zoology)
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In zoology, a club is a bony mass at the end of the
tail The tail is the section at the rear end of certain kinds of animals’ bodies; in general, the term refers to a distinct, flexible appendage to the torso. It is the part of the body that corresponds roughly to the sacrum and coccyx in mammal ...
of some
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
s and of some mammals, most notably the
ankylosaurids Ankylosauridae () is a family of armored dinosaurs within Ankylosauria, and is the sister group to Nodosauridae. The oldest known Ankylosaurids date to around 122 million years ago and went extinct 66 million years ago during the Cretaceous–Pal ...
and the
glyptodont Glyptodonts are an extinct subfamily of large, heavily armoured armadillos. They arose in South America around 48 million years ago and spread to southern North America after the continents became connected several million years ago. The best-k ...
s, as well as meiolaniid
turtle Turtles are an order of reptiles known as Testudines, characterized by a special shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Cryptodira (hidden necked t ...
s. It is thought that this was a form of defensive
armour Armour (British English) or armor (American English; see spelling differences) is a covering used to protect an object, individual, or vehicle from physical injury or damage, especially direct contact weapons or projectiles during combat, or f ...
or weapon that was used to defend against
predator Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill t ...
s, much in the same way as a thagomizer, possessed by stegosaurids, though at least in glyptodonts it is hypothesized it was used in fighting for mating rights. Among dinosaurs, the club was present mainly in ankylosaurids, although
sauropods Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
like ''
Shunosaurus ''Shunosaurus'', meaning "shu lizard", is a genus of sauropod dinosaur from Late Jurassic ( Oxfordian) beds in Sichuan Province in China, approximately 159±2 million years ago. The name derives from "Shu", an ancient name for the Sichuan p ...
'' also possessed a tail club. The tail club is most often depicted on ''
Ankylosaurus ''Ankylosaurus'' is a genus of armored dinosaur. Its fossils have been found in geological formations dating to the very end of the Cretaceous Period, about 68–66 million years ago, in western North America, making it among the last of th ...
'', especially in encounters with larger predators such as ''
Tyrannosaurus ''Tyrannosaurus'' is a genus of large theropoda, theropod dinosaur. The species ''Tyrannosaurus rex'' (''rex'' meaning "king" in Latin), often called ''T. rex'' or colloquially ''T-Rex'', is one of the best represented theropods. ''Tyrannosa ...
''.
Victoria Arbour Victoria Megan Arbour is a Canadian evolutionary biologist and vertebrate palaeontologist at Royal BC Museum, where she is Curator of Palaeontology. An "expert on the armoured dinosaurs known as ankylosaurs", Arbour analyzes fossils and creates ...
has established that ankylosaurid tails could generate enough force to break bone during impacts. In a separate study, Arbour suggested tail clubs as well as large armoured herbivores as a whole evolve when animals are too large to hide and too small to avoid predation by size alone.


See also

* Thagomizer


References

Mammal anatomy Dinosaur anatomy {{Vertebrate anatomy-stub