Cixiidae
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The Cixiidae are a family of fulgoroid insects, one of many families commonly known as planthoppers, distributed worldwide and comprising more than 2,000 species from over 150 genera. The genera are placed into three subfamilies, Borystheninae, Bothriocerinae and
Cixiinae Cixiinae is a planthopper subfamily in the family Cixiidae The Cixiidae are a family of Fulgoroidea, fulgoroid insects, one of many families commonly known as planthoppers, distributed worldwide and comprising more than 2,000 species from over ...
with sixteen tribes currently accepted in Cixiinae.


Description

Cixiid species are typically comparatively small (body size less than a centimeter) and usually inconspicuous. The face is longer than wide and the head is narrower than the
pronotum The prothorax is the foremost of the three segments in the thorax of an insect, and bears the first pair of legs. Its principal sclerites (exoskeletal plates) are the pronotum ( dorsal), the prosternum (ventral), and the propleuron (lateral) on e ...
. The forewings are at least partly
transparent Transparency, transparence or transparent most often refer to: * Transparency (optics), the physical property of allowing the transmission of light through a material They may also refer to: Literal uses * Transparency (photography), a still, ...
and the veins bear minute
setae In biology, setae (singular seta ; from the Latin word for " bristle") are any of a number of different bristle- or hair-like structures on living organisms. Animal setae Protostomes Annelid setae are stiff bristles present on the body. ...
. The hind tibiae end in a cluster of spines and may sometimes have spines along their length.
Nymphs A nymph ( grc, νύμφη, nýmphē, el, script=Latn, nímfi, label=Modern Greek; , ) in ancient Greek folklore is a minor female nature deity. Different from Greek goddesses, nymphs are generally regarded as personifications of nature, are ...
live underground, feeding on roots. Adults feed on herbs, shrubs and/or trees; some are polyphagous, while others are specialised on their host plants (monophagous). A couple of species are cavernicolous, feeding on roots in volcanic caves. Females occasionally bear impressive "wax tails" produced by wax-producing plates at the tip of their abdomen. Several species are of economic importance (e.g. ''
Hyalesthes obsoletus ''Hyalesthes obsoletus'' is a bug species in the genus '' Hyalesthes''. ''H. obsoletus'' is the vector of the black wood disease of grapevine. References Insect vectors of plant pathogens Insects described in 1865 Cixiidae {{Fulgo ...
'', ''
Haplaxius crudus ''Haplaxius crudus'' is a planthopper species in the genus '' Haplaxius''. ''H. crudus'' is the vector of the coconut lethal yellowing/16SrIV-A.Brown, S.E., Been, B.O. & McLaughlin, W.A. (2006). Detection and variability of the lethal yellowing ...
'').
Phytoplasma Phytoplasmas are obligate intracellular parasites of plant phloem tissue and of the insect vectors that are involved in their plant-to-plant transmission. Phytoplasmas were discovered in 1967 by Japanese scientists who termed them mycoplasma-lik ...
are common
parasite Parasitism is a close relationship between species, where one organism, the parasite, lives on or inside another organism, the host, causing it some harm, and is adapted structurally to this way of life. The entomologist E. O. Wilson has ...
s in these insects, causing diseases in
coconut palm The coconut tree (''Cocos nucifera'') is a member of the palm tree family (Arecaceae) and the only living species of the genus ''Cocos''. The term "coconut" (or the archaic "cocoanut") can refer to the whole coconut palm, the seed, or t ...
s and foliage,
grape A grape is a fruit, botanically a berry, of the deciduous woody vines of the flowering plant genus '' Vitis''. Grapes are a non- climacteric type of fruit, generally occurring in clusters. The cultivation of grapes began perhaps 8,000 years a ...
vines, sugar beets, and
lilies ''Lilium'' () is a genus of herbaceous flowering plants growing from bulbs, all with large prominent flowers. They are the true lilies. Lilies are a group of flowering plants which are important in culture and literature in much of the world. M ...
.


Fossil record

The
fossil record A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
of Cixiidae is limited, and a number of taxa which have placed into the family may need to be reexamined and moved to different families. The oldest confirmed taxa are from the Early Cretaceous with '' ‘Cixius’ petrinus'' described from
Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 129.4 ± 1.5 Ma (million years ago) and 121.4 ± 1.0 Ma). It is a subdivision of the Early Cretaceous Epoch (or Lower Cretaceous Series). It is precede ...
deposits in
England England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe b ...
, '' Karebodopoides aptianus'' from
Hauterivian The Hauterivian is, in the geologic timescale, an age in the Early Cretaceous Epoch or a stage in the Lower Cretaceous Series. It spans the time between 132.9 ± 2 Ma and 129.4 ± 1.5 Ma (million years ago). The Hauterivian is preceded by the ...
to Aptian
Lebanese amber Lebanese amber is fossilized resin found in Lebanon and southwest Syria. It dates back approximately 130-125 million years to the Barremian of the Early Cretaceous. It formed on what was then the northern coast of Gondwana, believed to be a tropi ...
and '' Cretofennahia cretacea'' plus an unnamed specimen from the Aptian of
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
. Due to the abundant nature of Cixiidae as inclusions in
Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', " ...
Baltic amber The Baltic region is home to the largest known deposit of amber, called Baltic amber or succinite. It was produced sometime during the Eocene epoch, but exactly when is controversial. It has been estimated that these forests created more than ...
a number of taxa have been described, including '' Glisachaemus jonasdamzeni'' and '' Autrimpus sambiorum''. Several taxa have also been described from
Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
Dominican amber Dominican amber is amber from the Dominican Republic derived from resin of the extinct tree '' Hymenaea protera''. Dominican amber differentiates itself from Baltic amber by being nearly always transparent, and it has a higher number of fossil inc ...
including '' Oligocixia electrina'' and '' Oliarius kulickae''.


See also

*
List of Cixiidae genera These 232 genera belong to the family Cixiidae, cixiid planthoppers. There are at least 2,500 described species in Cixiidae. Cixiidae genera * '' Achaebana'' Attie, Bourgoin & Bonfils, 2002 * ''Achaemenes'' Stål, 1866 * '' Adolendana'' Distant, ...
File:Cixiid Bandipur.jpg File:Cixiid WG.jpg File:CixiidaeNilgiris.jpg, ''
Nilgiris The Nilgiri Mountains form part of the Western Ghats in northwestern Tamil Nadu, Southern Karnataka, and eastern Kerala in India. They are located at the trijunction of three states and connect the Western Ghats with the Eastern Ghats. At le ...
'' File:Brixia albomaculata.jpg, Female ''Brixia albomaculata'' showing wax filaments (India) File:Tachycixius pilosus (Cixiidae), Moergestel, the Netherlands.jpg, ''Tachycixius pilosus'' File:Another cixiid hopper - Flickr - treegrow.jpg, ''Bothriocera cognita''


References


External links


Cixiidae species list
(parts are in German)
FLOW - Fulgoromorpha Lists On the Web
Auchenorrhyncha families Fulgoromorpha {{Fulgoromorpha-stub