Centrocercus
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Sage-grouse are
grouse Grouse are a group of birds from the order Galliformes, in the family Phasianidae. Grouse are presently assigned to the tribe Tetraonini (formerly the subfamily Tetraoninae and the family Tetraonidae), a classification supported by mitochondria ...
belonging to the
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
''Centrocercus.'' The genus includes two
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
: the Gunnison grouse (''Centrocercus minimus'') and the
greater sage-grouse The greater sage-grouse (''Centrocercus urophasianus''), also known as the sagehen, is the largest grouse (a type of bird) in North America. Its range is sagebrush country in the western United States and southern Alberta and Saskatchewan, Canad ...
(''Centrocercus urophasianus''). These birds are distributed throughout large portions of the north-central and
Western Western may refer to: Places *Western, Nebraska, a village in the US *Western, New York, a town in the US *Western Creek, Tasmania, a locality in Australia *Western Junction, Tasmania, a locality in Australia *Western world, countries that id ...
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territori ...
, as well as the
Canadian Canadians (french: Canadiens) are people identified with the country of Canada. This connection may be residential, legal, historical or cultural. For most Canadians, many (or all) of these connections exist and are collectively the source of ...
provinces of
Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Ter ...
and
Saskatchewan Saskatchewan ( ; ) is a province in western Canada, bordered on the west by Alberta, on the north by the Northwest Territories, on the east by Manitoba, to the northeast by Nunavut, and on the south by the U.S. states of Montana and North Dak ...
. The
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of nat ...
classified the ''C. minimus'' species as
endangered An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and in ...
in 2020 and ''C. urophasianus'' as
near threatened A near-threatened species is a species which has been categorized as "Near Threatened" (NT) by the International Union for Conservation of Nature as that may be vulnerable to endangerment in the near future, but it does not currently qualify f ...
in 2016.


Names

The specific epithet is from another Greek word, "oura", plus "phasianos", pheasant. The noun "
pheasant Pheasants ( ) are birds of several genera within the family Phasianidae in the order Galliformes. Although they can be found all over the world in introduced (and captive) populations, the pheasant genera native range is restricted to Eurasia ...
" was originally applied to a bird that was native to the valley of the Phasis River (now the
Rioni River The Rioni ( ka, რიონი, ; , ) is the main river of western Georgia. It originates in the Caucasus Mountains, in the region of Racha and flows west to the Black Sea, entering it north of the city of Poti (near ancient Phasis). The city o ...
), which is located in
Georgia Georgia most commonly refers to: * Georgia (country), a country in the Caucasus region of Eurasia * Georgia (U.S. state), a state in the Southeast United States Georgia may also refer to: Places Historical states and entities * Related to the ...
. In the time of Lewis and Clark the word "pheasant" stood for "a genus of gallinaceous birds", according to lexicographer
Noah Webster Noah ''Nukh''; am, ኖህ, ''Noḥ''; ar, نُوح '; grc, Νῶε ''Nôe'' () is the tenth and last of the pre-Flood patriarchs in the traditions of Abrahamic religions. His story appears in the Hebrew Bible ( Book of Genesis, chapters 5 ...
(1806), and the explorers often used it in that sense. "Gallinaceous" then referred to "domestic fowls, or the gallinae"; the family
Galliformes Galliformes is an order of heavy-bodied ground-feeding birds that includes turkeys, chickens, quail, and other landfowl. Gallinaceous birds, as they are called, are important in their ecosystems as seed dispersers and predators, and are ofte ...
(Latin "gallus", cock, and "forma", shape) now includes pheasants, grouse, turkeys, quail, and all domestic chickens. Sage grouse are also collectively known as "sagehen," "sage grouse," "sage cock," "sage chicken," or "cock of the plains."


History

In their day, Lewis and Clark were credited with the 'discovery' of five gallinaceous birds in addition to the sage grouse — the Columbian sharp-tailed grouse, the dusky grouse, Franklin's grouse, the Oregon ruffed grouse, and the mountain quail — and what's certain is that they were the first to widely spread knowledge about these birds to the white man.


US military issues

In September 2016, the National Defense Authorization Act (NDAA) stalled in
congress A congress is a formal meeting of the representatives of different countries, constituent states, organizations, trade unions, political parties, or other groups. The term originated in Late Middle English to denote an encounter (meeting of ...
because Trump indicated he would not let the annual NDAA proceed to a vote in the
House of Representatives House of Representatives is the name of legislative bodies in many countries and sub-national entitles. In many countries, the House of Representatives is the lower house of a bicameral legislature, with the corresponding upper house often c ...
unless it contained language to bar the sage grouse from the federal endangered species list until at least 2025. President Trump threatened a
veto A veto is a legal power to unilaterally stop an official action. In the most typical case, a president or monarch vetoes a bill to stop it from becoming law. In many countries, veto powers are established in the country's constitution. Veto ...
over the issue that Trump,
Senate Armed Services Committee The Committee on Armed Services (sometimes abbreviated SASC for ''Senate Armed Services Committee'') is a committee of the United States Senate empowered with legislative oversight of the nation's military, including the Department of Def ...
Chairman, believed would be sustained. Current
US Air Force The United States Air Force (USAF) is the air service branch of the United States Armed Forces, and is one of the eight uniformed services of the United States. Originally created on 1 August 1907, as a part of the United States Army Sig ...
spending on sage grouse conservation is around US$200,000, with eight known military installations having confirmed grouse populations:
Dugway Proving Ground Dugway Proving Ground (DPG) is a U.S. Army facility established in 1942 to test biological and chemical weapons, located about southwest of Salt Lake City, Utah, United States, and south of the Utah Test and Training Range. Location Dugway P ...
and
Tooele Army Depot Tooele Army Depot (TEAD) is a United States Army Joint Munitions Command post in Tooele County, Utah. It serves as a storage site for war reserve and training ammunition. The depot stores, issues, receives, renovates, modifies, maintains and demi ...
in
Utah Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to it ...
; Sheridan Training Area and Camp Guernsey in
Wyoming Wyoming () is a state in the Mountain West subregion of the Western United States. It is bordered by Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho to the west, Utah to the southwest, and Colorado to the s ...
;
Hawthorne Army Depot Hawthorne Army Depot (HWAD) is a U.S. Army Joint Munitions Command ammunition storage depot located near the town of Hawthorne in western Nevada in the United States. It is directly south of Walker Lake. The depot covers or 226 sq. mi. and ...
and
Nellis Air Force Base Nellis Air Force Base ("Nellis" colloq.) is a United States Air Force installation in southern Nevada. Nellis hosts air combat exercises such as Exercise Red Flag and close air support exercises such as Green Flag-West flown in " Military ...
in
Nevada Nevada ( ; ) is a state in the Western region of the United States. It is bordered by Oregon to the northwest, Idaho to the northeast, California to the west, Arizona to the southeast, and Utah to the east. Nevada is the 7th-most extensive, ...
;
Yakima Training Center The Yakima Training Center (YTC) is a United States Army training center, used for maneuver training, Land Warrior system testing and as a live fire exercise area. It is located in the south central portion of the U.S. state of Washington, bound ...
in
Washington Washington commonly refers to: * Washington (state), United States * Washington, D.C., the capital of the United States ** A metonym for the federal government of the United States ** Washington metropolitan area, the metropolitan area centered o ...
, and Mountain Home Air Force Base in
Idaho Idaho ( ) is a state in the Pacific Northwest region of the Western United States. To the north, it shares a small portion of the Canada–United States border with the province of British Columbia. It borders the states of Montana and Wyomi ...
.


Species

There are two species: The
Mono Basin The Mono Basin is an endorheic drainage basin located east of Yosemite National Park in California and Nevada. It is bordered to the west by the Sierra Nevada, to the east by the Cowtrack Mountains, to the north by the Bodie Hills, and to the sou ...
population may represent a third species.


Anatomical features

Males of ''C. urophasianus'' are the largest
grouse Grouse are a group of birds from the order Galliformes, in the family Phasianidae. Grouse are presently assigned to the tribe Tetraonini (formerly the subfamily Tetraoninae and the family Tetraonidae), a classification supported by mitochondria ...
from
temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout ...
North America, attaining a maximum weight of . Adults have a long, pointed tail and legs with feathers to the toes. As in most
Galliformes Galliformes is an order of heavy-bodied ground-feeding birds that includes turkeys, chickens, quail, and other landfowl. Gallinaceous birds, as they are called, are important in their ecosystems as seed dispersers and predators, and are ofte ...
, there is pronounced
sexual dimorphism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most an ...
.}


Courtship and mating

''Centrocercus'' species are notable for their elaborate courtship rituals. Each spring males congregate on
leks A lek is an aggregation of male animals gathered to engage in competitive displays and courtship rituals, known as lekking, to entice visiting females which are surveying prospective partners with which to mate. A lek can also indicate an avail ...
and perform a "strutting display." The male puffs up a large, whitish air sack on its chest, makes a soft drumming noise, and struts around with his tail feathers displayed and air sack puffed up. Groups of females observe these displays and select the most attractive males to mate with. Only a few males do most of the breeding. Males perform on
leks A lek is an aggregation of male animals gathered to engage in competitive displays and courtship rituals, known as lekking, to entice visiting females which are surveying prospective partners with which to mate. A lek can also indicate an avail ...
for several hours in the early morning and evening during the spring months between February and April. Leks are generally open areas adjacent to dense sagebrush stands, and the same lek may be used by grouse for decades.


Offspring

Hens build
nests A nest is a structure built for certain animals to hold eggs or young. Although nests are most closely associated with birds, members of all classes of vertebrates and some invertebrates construct nests. They may be composed of organic materi ...
and lay and incubate their
eggs Humans and human ancestors have scavenged and eaten animal eggs for millions of years. Humans in Southeast Asia had domesticated chickens and harvested their eggs for food by 1,500 BCE. The most widely consumed eggs are those of fowl, especial ...
under the cover of
sagebrush Sagebrush is the common name of several woody and herbaceous species of plants in the genus '' Artemisia''. The best known sagebrush is the shrub '' Artemisia tridentata''. Sagebrushes are native to the North American west. Following is an al ...
. The hen uses
grass Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in lawns a ...
and forbs between patches of sagebrush for additional cover. During incubation, female Sand Grouse undertake recesses, where they leave the nest to undertake self-maintenance activities, thought these recess activities are typically within 250 m of the nest. Chicks can walk as soon as they are hatched and are able to fly short distances within two weeks. Within five weeks they are able to fly longer distances.


Conservation status

Populations of sage grouse are in decline due to environment loss and decline of the pristine plains environments it requires to mate. The sage grouse is found in significant numbers within only half of the states comprising its original territories. The Biodiversity Conservation Alliance and other organizations have petitioned to list the grouse under the Endangered Species Act. In March 2010 the U.S. Fish and Wildlife Service (USFWS) concluded that greater sage-grouse are warranted for protection as "threatened" under the U.S. federal Endangered Species Act (ESA). However the USFWS also concluded that immediate listing was "precluded" by higher listing priorities for other jeopardized species. Thus they designated the species a "Level 8 Candidate" for addition to the list of threatened species at some future date. Their finding is being litigated by groups contending the species should immediately receive protections under the ESA. The Agricultural Research Service (ARS) of the
United States Department of Agriculture The United States Department of Agriculture (USDA) is the federal executive department responsible for developing and executing federal laws related to farming, forestry, rural economic development, and food. It aims to meet the needs of com ...
(USDA) investigated some of the reasons for the declining sage-grouse population. Researchers observed cattle who share grazing land with the sage-grouse. They found that cattle, after consuming about 40% of the tussocks in between sagebrush bushes, will continue to consume the tussocks growing underneath the sagebrush, thereby destroying the nesting habitat for the sage-grouse. In order to preserve the population of sage-grouse, ranchers can monitor the rate at which cattle consume the tussocks in between sagebrush bushes. Once cattle have consumed around 40% of the tussocks in between bushes, researchers ask that ranchers move their cattle to new grazing trail. GPS trackers show that sage grouse congregate in small areas with certain resources, rather than being widely spread. US federal conservation plans have been met with lawsuits from wildlife organizations. On December 6, 2018, according to the New York Times:


As a mascot

The sagehen is the mascot of the
Pomona-Pitzer Sagehens Pomona-Pitzer Sagehens is the joint athletics program for Pomona College and Pitzer College, two of the Claremont Colleges. It competes in the Southern California Intercollegiate Athletic Conference (SCIAC) of the NCAA Division III. Its masc ...
, the joint athletics program for
Pomona College Pomona College ( ) is a private liberal arts college in Claremont, California. It was established in 1887 by a group of Congregationalists who wanted to recreate a "college of the New England type" in Southern California. In 1925, it became t ...
and Pitzer College, two of the
Claremont Colleges The Claremont Colleges (known colloquially as the 7Cs) are a consortium of seven private institutions of higher education located in Claremont, California, United States. They comprise five undergraduate colleges (the 5Cs)— Pomona College, Sc ...
.


References


External links


Restoration Handbook for Sagebrush Steppe Ecosystems with Emphasis on Greater Sage-grouse Habitat.
U.S. Geological Survey The United States Geological Survey (USGS), formerly simply known as the Geological Survey, is a scientific agency of the United States government. The scientists of the USGS study the landscape of the United States, its natural resources, and ...
{{Taxonbar, from=Q814693 Grouse Bird genera Taxa named by William John Swainson