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''Dianthus caryophyllus'' (), commonly known as the carnation or clove pink, is a species of '' Dianthus''. It is likely native to the Mediterranean region but its exact range is unknown due to extensive cultivation for the last 2,000 years.Med-Checklist
''Dianthus caryophyllus''
/ref>Flora Europaea
''Dianthus caryophyllus''
/ref>Blamey, M. & Grey-Wilson, C. (1989). ''Flora of Britain and Northern Europe''. Huxley, A., ed. (1992). ''New RHS Dictionary of Gardening''. Macmillan .


Taxonomy

Carnations were mentioned in Greek literature 2,000 years ago. The term ''dianthus'' was coined by Greek botanist
Theophrastus Theophrastus (; grc-gre, Θεόφραστος ; c. 371c. 287 BC), a Greek philosopher and the successor to Aristotle in the Peripatetic school. He was a native of Eresos in Lesbos.Gavin Hardy and Laurence Totelin, ''Ancient Botany'', Routle ...
, and is derived from the
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic pe ...
words for divine ("dios") and flower ("anthos"). The name "carnation" is believed to come from the Latin ''corona-ae'', a "wreath, garland, chaplet, crown",Cassell's Latin Dictionary, Marchant, J.R.V, & Charles, Joseph F., (Eds.), Revised Edition, 1928 as it was one of the flowers used in Greek and Roman ceremonial crowns, or possibly from the Latin ''caro'' (genitive ''carnis''), "flesh", which refers to the natural colour of the flower, or in Christian iconography ''incarnatio'', "
incarnation Incarnation literally means ''embodied in flesh'' or ''taking on flesh''. It refers to the conception and the embodiment of a deity or spirit in some earthly form or the appearance of a god as a human. If capitalized, it is the union of divinit ...
", God made flesh in the form of Jesus.
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, ...
described the carnation in volume one of his ''
Species Plantarum ' (Latin for "The Species of Plants") is a book by Carl Linnaeus, originally published in 1753, which lists every species of plant known at the time, classified into genera. It is the first work to consistently apply binomial names and was the ...
'' in 1753, giving it the name ''Dianthus caryophyllus'', Although originally applied to the species ''Dianthus caryophyllus'', the name carnation is also often applied to some of the other species of '' Dianthus'', and more particularly to garden hybrids between ''D. caryophyllus'' and other species in the genus.


Description

''Dianthus caryophyllus'' is a
herbaceous Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition o ...
perennial plant A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
growing up to tall. The
leaves A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, st ...
are
glaucous ''Glaucous'' (, ) is used to describe the pale grey or bluish-green appearance of the surfaces of some plants, as well as in the names of birds, such as the glaucous gull (''Larus hyperboreus''), glaucous-winged gull (''Larus glaucescens''), ...
greyish green to blue-green, slender, up to long. The
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechanis ...
s are produced singly or up to five together in a cyme; they are around diameter, and sweetly scented; the original natural flower color is bright pinkish-purple, but
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture ...
s of other colors, including red, white, yellow, blue and green, along with some white with colored striped variations have been developed. The fragrant,
hermaphrodite In reproductive biology, a hermaphrodite () is an organism that has both kinds of reproductive organs and can produce both gametes associated with male and female sexes. Many taxonomic groups of animals (mostly invertebrates) do not have ...
flowers have a radial symmetry. The four to six surrounding the
calyx Calyx or calyce (plural "calyces"), from the Latin ''calix'' which itself comes from the Ancient Greek ''κάλυξ'' (''kálux'') meaning "husk" or "pod", may refer to: Biology * Calyx (anatomy), collective name for several cup-like structures ...
, egg-shaped, sting-pointed scales leaves are only ¼ as long as the calyx tube.Flora of NW Europe
''Dianthus caryophyllus''


Distribution and habitat

The wild carnation is found in the Mediterranean countries of
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = '' Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , ...
,
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical ...
,
Croatia , image_flag = Flag of Croatia.svg , image_coat = Coat of arms of Croatia.svg , anthem = " Lijepa naša domovino"("Our Beautiful Homeland") , image_map = , map_caption = , capi ...
,
Albania Albania ( ; sq, Shqipëri or ), or , also or . officially the Republic of Albania ( sq, Republika e Shqipërisë), is a country in Southeastern Europe. It is located on the Adriatic and Ionian Seas within the Mediterranean Sea and share ...
,
Greece Greece,, or , romanized: ', officially the Hellenic Republic, is a country in Southeast Europe. It is situated on the southern tip of the Balkans, and is located at the crossroads of Europe, Asia, and Africa. Greece shares land borders wi ...
and
Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a transcontinental country located mainly on the Anatolian Peninsula in Western Asia, with a small portion on the Balkan Peninsula ...
.


Cultivation and uses

Carnations require well-drained, neutral to slightly alkaline soil, and full sun. Numerous
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture ...
s have been selected for garden planting. Typical examples include 'Gina Porto', 'Helen', 'Laced Romeo', and 'Red Rocket'. They are used for medical purposes, such as for upset stomach and fever. Their fragrance was historically used for
vinegar Vinegar is an aqueous solution of acetic acid and trace compounds that may include flavorings. Vinegar typically contains 5–8% acetic acid by volume. Usually, the acetic acid is produced by a double fermentation, converting simple sugars to eth ...
, beer, wine, sauces and salads. Crossbreeding ''D. caryophyllus'' with ''D. capitatus'' results in a hybrid that is resistant to bacterial wilt from '' Paraburkholderia caryophylli''. However, the flower is less attractive and so more breeding and backcrossing is needed to improve the flower. Carnation cultivars with no fragrance are often used by men as
boutonnière A boutonnière () or buttonhole (British English) is a floral decoration, typically a single flower or bud, worn on the lapel of a tuxedo or suit jacket. While worn frequently in the past, boutonnières are now usually reserved for special oc ...
s or "buttonholes".


Diseases


Symbolism


Traditional meanings

For the most part, carnations express love, fascination, and distinction, though there are many variations dependent on color. * Along with the red rose, the red carnation can be used as a symbol of
socialism Socialism is a left-wing economic philosophy and movement encompassing a range of economic systems characterized by the dominance of social ownership of the means of production as opposed to private ownership. As a term, it describes th ...
and the
labour movement The labour movement or labor movement consists of two main wings: the trade union movement (British English) or labor union movement (American English) on the one hand, and the political labour movement on the other. * The trade union movement ...
, and historically has often been used in demonstrations on
International Workers' Day International Workers' Day, also known as Labour Day in some countries and often referred to as May Day, is a celebration of labourers and the working classes that is promoted by the international labour movement and occurs every year on 1 May, ...
(
May Day May Day is a European festival of ancient origins marking the beginning of summer, usually celebrated on 1 May, around halfway between the spring equinox and summer solstice. Festivities may also be held the night before, known as May Eve. Tr ...
). * In
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's List of countries and dependencies by population, most populous country, with a Population of China, population exceeding 1.4 billion, slig ...
, the carnation flower is the most frequently used flower in weddings. * In
Portugal Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country whose mainland is located on the Iberian Peninsula of Southwestern Europe, and whose territory also includes the Atlantic archipelagos of th ...
, bright red carnations were used when in 1974 the authoritarian Estado Novo regime was overthrown; therefore, this transition (brought about by a combination of a ''coup d'état'' with
civil resistance Civil resistance is political action that relies on the use of nonviolent resistance by ordinary people to challenge a particular power, force, policy or regime. Civil resistance operates through appeals to the adversary, pressure and coercion: it ...
) is known as the
Carnation Revolution The Carnation Revolution ( pt, Revolução dos Cravos), also known as the 25 April ( pt, 25 de Abril, links=no), was a military coup by left-leaning military officers that overthrew the authoritarian Estado Novo regime on 25 April 1974 in Lisbo ...
. * Light red carnations represent admiration, while dark red denote deep love and affection. * White carnations represent pure love and good luck, while striped (variegated) carnations symbolise regret that a love cannot be shared. * White carnations, in the
Netherlands ) , anthem = ( en, "William of Nassau") , image_map = , map_caption = , subdivision_type = Sovereign state , subdivision_name = Kingdom of the Netherlands , established_title = Before independence , established_date = Spanish Netherl ...
are associated with Prince Bernhard. He wore one during
World War II World War II or the Second World War, often abbreviated as WWII or WW2, was a world war that lasted from 1939 to 1945. It involved the World War II by country, vast majority of the world's countries—including all of the great power ...
and in a gesture of defiance some of the Dutch population took up this gesture. After the war the white carnation became a sign of the Prince, veterans and remembrance of the resistance. * Purple carnations indicate capriciousness. In
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
, it is a traditional funeral flower, given in condolence for the death of a loved one. * Carnation is the
birth flower Floriography (language of flowers) is a means of cryptological communication through the use or arrangement of flowers. Meaning has been attributed to flowers for thousands of years, and some form of floriography has been practiced in tradition ...
for those born in the month of January. *Since Ottoman times, red carnations and
tulip Tulips (''Tulipa'') are a genus of spring-blooming perennial herbaceous bulbiferous geophytes (having bulbs as storage organs). The flowers are usually large, showy and brightly coloured, generally red, pink, yellow, or white (usually in warm ...
s are used in the interior wall paintings of
mosque A mosque (; from ar, مَسْجِد, masjid, ; literally "place of ritual prostration"), also called masjid, is a place of prayer for Muslims. Mosques are usually covered buildings, but can be any place where prayers ( sujud) are performed, ...
s in
Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a transcontinental country located mainly on the Anatolian Peninsula in Western Asia, with a small portion on the Balkan Peninsula ...
. It is often said that while tulips represent God, carnations is the symbol for
Muhammad Muhammad ( ar, مُحَمَّد;  570 – 8 June 632 CE) was an Arab religious, social, and political leader and the founder of Islam. According to Islamic doctrine, he was a prophet divinely inspired to preach and confirm the mon ...
. However these flower designs are not unique to mosques but also used in many other Ottoman traditional art forms. *In
Azerbaijan Azerbaijan (, ; az, Azərbaycan ), officially the Republic of Azerbaijan, , also sometimes officially called the Azerbaijan Republic is a transcontinental country located at the boundary of Eastern Europe and Western Asia. It is a part of th ...
, red carnations has turned into a symbol of mourning after the usage of the flower during
Black January Black January ( az, Qara Yanvar), also known as Black Saturday or the January Massacre, was a violent crackdown on the civilian population of Baku on 19–20 January 1990, as part of a state of emergency during the dissolution of the Soviet Uni ...
events of 1990, a violent crackdown on the civilian population of the country by USSR troops. *In the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country Continental United States, primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., ...
, former U.S. President
William McKinley William McKinley (January 29, 1843September 14, 1901) was the 25th president of the United States, serving from 1897 until his assassination in 1901. As a politician he led a realignment that made his Republican Party largely dominant in t ...
considered the red carnation to be his lucky flower and often wore one on his lapel. Following his assassination in 1901, the State of Ohio adopted the red carnation as its state flower to honor McKinley The formal name for carnation, ''dianthus'', comes from Greek for "heavenly flower", or the flower of Jove.


Holidays and events

Carnations are often worn on special occasions, especially
Mother's Day Mother's Day is a celebration honoring the mother of the family or individual, as well as motherhood, maternal bonds, and the influence of mothers in society. It is celebrated on different days in many parts of the world, most commonly in th ...
and weddings. In 1907,
Anna Jarvis Anna Maria Jarvis (May 1, 1864 – November 24, 1948) was the founder of Mother's Day in the United States. Her mother had frequently expressed a desire for the establishment of such a holiday, and after her mother's death, Jarvis led the moveme ...
chose a carnation as the emblem of Mother's Day because it was her mother's favourite flower. This tradition is now observed in the United States and Canada on the second Sunday in May. Ann Jarvis chose the white carnation because she wanted to represent the purity of a mother's love. This meaning has evolved over time, and now a red carnation may be worn if one's mother is alive, and a white one if she has died. In
Korea Korea ( ko, 한국, or , ) is a peninsular region in East Asia. Since 1945, it has been divided at or near the 38th parallel, with North Korea (Democratic People's Republic of Korea) comprising its northern half and South Korea (Republic ...
, carnations express admiration, love and gratitude. Red and pink carnations are worn on Parents Day (Korea does not separate Mother's Day or Father's Day, but has Parents Day on 8 May). Sometimes, parents wear a corsage of carnation(s) on their left chest on Parents Day. Carnations are also worn on Teachers Day (15 May). Red carnations are worn on
May Day May Day is a European festival of ancient origins marking the beginning of summer, usually celebrated on 1 May, around halfway between the spring equinox and summer solstice. Festivities may also be held the night before, known as May Eve. Tr ...
as a symbol of
socialism Socialism is a left-wing economic philosophy and movement encompassing a range of economic systems characterized by the dominance of social ownership of the means of production as opposed to private ownership. As a term, it describes th ...
and the
labour movement The labour movement or labor movement consists of two main wings: the trade union movement (British English) or labor union movement (American English) on the one hand, and the political labour movement on the other. * The trade union movement ...
in some countries, such as
Austria Austria, , bar, Östareich officially the Republic of Austria, is a country in the southern part of Central Europe, lying in the Eastern Alps. It is a federation of nine states, one of which is the capital, Vienna, the most populous ...
, Italy, and successor countries of the
former Yugoslavia The Socialist Federal Republic of Yugoslavia, commonly referred to as SFR Yugoslavia or simply as Yugoslavia, was a country in Central and Southeast Europe. It emerged in 1945, following World War II, and lasted until 1992, with the breakup of Yu ...
. The red carnation is also the symbol of the
Carnation Revolution The Carnation Revolution ( pt, Revolução dos Cravos), also known as the 25 April ( pt, 25 de Abril, links=no), was a military coup by left-leaning military officers that overthrew the authoritarian Estado Novo regime on 25 April 1974 in Lisbo ...
in
Portugal Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country whose mainland is located on the Iberian Peninsula of Southwestern Europe, and whose territory also includes the Atlantic archipelagos of th ...
. Green carnations are for St. Patrick's Day and were famously worn by the Irish writer
Oscar Wilde Oscar Fingal O'Flahertie Wills Wilde (16 October 185430 November 1900) was an Irish poet and playwright. After writing in different forms throughout the 1880s, he became one of the most popular playwrights in London in the early 1890s. He is ...
. The green carnation thence became a symbol of
homosexuality Homosexuality is Romance (love), romantic attraction, sexual attraction, or Human sexual activity, sexual behavior between members of the same sex or gender. As a sexual orientation, homosexuality is "an enduring pattern of emotional, romant ...
in the early 20th century, especially through the book '' The Green Carnation'' and
Noël Coward Sir Noël Peirce Coward (16 December 189926 March 1973) was an English playwright, composer, director, actor, and singer, known for his wit, flamboyance, and what ''Time'' magazine called "a sense of personal style, a combination of cheek and ...
's song, "We All Wear a Green Carnation" in his
operetta Operetta is a form of theatre and a genre of light opera. It includes spoken dialogue, songs, and dances. It is lighter than opera in terms of its music, orchestral size, length of the work, and at face value, subject matter. Apart from its ...
, ''
Bitter Sweet Bittersweet, bitter-sweet, or bitter sweet may refer to: Biology * A vine in the nightshade family, ''Solanum dulcamara'' * Some species of vines in the genus ''Celastrus'', including American bittersweet (''C. scandens'') and Oriental bitters ...
''. In communist
Czechoslovakia , rue, Чеськословеньско, , yi, טשעכאסלאוואקיי, , common_name = Czechoslovakia , life_span = 1918–19391945–1992 , p1 = Austria-Hungary , image_p1 ...
and in
Poland Poland, officially the Republic of Poland, is a country in Central Europe. It is divided into 16 administrative provinces called voivodeships, covering an area of . Poland has a population of over 38 million and is the fifth-most populou ...
in times of the
People's Republic of Poland The Polish People's Republic ( pl, Polska Rzeczpospolita Ludowa, PRL) was a country in Central Europe that existed from 1947 to 1989 as the predecessor of the modern Republic of Poland. With a population of approximately 37.9 million ne ...
, carnations were traditionally given to women on the widely celebrated Women's Day, together with commodities that were difficult to obtain due to the countries' communist system, such as
tights Tights are a kind of cloth garment, most often sheathing the body from the waist to the toe tips with a tight fit, hence the name. They come in absolute opaque, opaque, sheer and fishnet styles — or a combination, such as the original concep ...
, towels,
soap Soap is a salt of a fatty acid used in a variety of cleansing and lubricating products. In a domestic setting, soaps are surfactants usually used for washing, bathing, and other types of housekeeping. In industrial settings, soaps are us ...
and
coffee Coffee is a drink prepared from roasted coffee beans. Darkly colored, bitter, and slightly acidic, coffee has a stimulating effect on humans, primarily due to its caffeine content. It is the most popular hot drink in the world. Seeds of ...
. After the 1990 uprisings against Soviets in
Azerbaijan Azerbaijan (, ; az, Azərbaycan ), officially the Republic of Azerbaijan, , also sometimes officially called the Azerbaijan Republic is a transcontinental country located at the boundary of Eastern Europe and Western Asia. It is a part of th ...
in which 147 Azerbaijani civilians were killed, 800 people were injured and five people went missing, the carnation has become a symbol of the Black January tragedy associated with the carnations thrown into the puddles of blood shed in the streets of Azerbaijan subsequent to the massacre. At the
University of Oxford , mottoeng = The Lord is my light , established = , endowment = £6.1 billion (including colleges) (2019) , budget = £2.145 billion (2019–20) , chancellor ...
, carnations are traditionally worn to all examinations; white for the first exam, pink for exams in between, and red for the last exam. One story explaining this tradition relates that initially a white carnation was kept in a red inkpot between exams, so by the last exam it was fully red; the story is thought to originate in the late 1990s. Carnations are the traditional first wedding anniversary flower.


Symbols of territorial entities and organizations

The carnation is the
national flower In a number of countries, plants have been chosen as symbols to represent specific geographic areas. Some countries have a country-wide floral emblem; others in addition have symbols representing subdivisions. Different processes have been used to ...
of
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = '' Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , ...
,
Monaco Monaco (; ), officially the Principality of Monaco (french: Principauté de Monaco; Ligurian: ; oc, Principat de Mónegue), is a sovereign city-state and microstate on the French Riviera a few kilometres west of the Italian region of Lig ...
, and
Slovenia Slovenia ( ; sl, Slovenija ), officially the Republic of Slovenia (Slovene: , abbr.: ''RS''), is a country in Central Europe. It is bordered by Italy to the west, Austria to the north, Hungary to the northeast, Croatia to the southeast, and ...
, and the provincial flower of the autonomous community of the
Balearic Islands The Balearic Islands ( es, Islas Baleares ; or ca, Illes Balears ) are an archipelago in the Balearic Sea, near the eastern coast of the Iberian Peninsula. The archipelago is an autonomous community and a province of Spain; its capital is ...
. The state flower of
Ohio Ohio () is a U.S. state, state in the Midwestern United States, Midwestern region of the United States. Of the List of states and territories of the United States, fifty U.S. states, it is the List of U.S. states and territories by area, 34th-l ...
is a scarlet carnation, which was introduced to the state by
Levi L. Lamborn Levi Leslie Lamborn (October 10, 1829 – June 14, 1910) was an Ohio doctor, horticulturalist, and politician. Born in 1829 in Chester County, Pennsylvania, Lamborn was the son of Townsend Lamborn and Anna (Clayton) Lamborn.S. Lamborn, p. 180 Town ...
. The choice was made to honor
William McKinley William McKinley (January 29, 1843September 14, 1901) was the 25th president of the United States, serving from 1897 until his assassination in 1901. As a politician he led a realignment that made his Republican Party largely dominant in t ...
, Ohio governor and U.S. president, who was assassinated in 1901, and regularly wore a scarlet carnation on his lapel.


Colours

Carnations do not naturally produce the pigment delphinidin, thus a blue carnation cannot occur by
natural selection Natural selection is the differential survival and reproduction of individuals due to differences in phenotype. It is a key mechanism of evolution, the change in the heritable traits characteristic of a population over generations. Cha ...
or be created by traditional
plant breeding Plant breeding is the science of changing the traits of plants in order to produce desired characteristics. It has been used to improve the quality of nutrition in products for humans and animals. The goals of plant breeding are to produce cr ...
. It shares this characteristic with other widely sold flowers like
rose A rose is either a woody perennial flowering plant of the genus ''Rosa'' (), in the family Rosaceae (), or the flower it bears. There are over three hundred species and tens of thousands of cultivars. They form a group of plants that can be ...
s, lilies,
tulip Tulips (''Tulipa'') are a genus of spring-blooming perennial herbaceous bulbiferous geophytes (having bulbs as storage organs). The flowers are usually large, showy and brightly coloured, generally red, pink, yellow, or white (usually in warm ...
s,
chrysanthemum Chrysanthemums (), sometimes called mums or chrysanths, are flowering plants of the genus ''Chrysanthemum'' in the family Asteraceae. They are native to East Asia and northeastern Europe. Most species originate from East Asia and the cent ...
s and gerberas. Around 1996, a company, Florigene, used
genetic engineering Genetic engineering, also called genetic modification or genetic manipulation, is the modification and manipulation of an organism's genes using technology. It is a set of technologies used to change the genetic makeup of cells, including ...
to extract certain genes from petunia and snapdragon flowers to produce a blue-mauve carnation, which was commercialized as ''Moondust''. In 1998, a violet carnation called ''Moonshadow'' was commercialized.Phys.Org website. 4 April 200
Plant gene replacement results in the world's only blue rose
/ref> As of 2004, three additional blue-violet/purple varieties have been commercialized.


See also

* List of Award of Garden Merit dianthus


References


External links


Carnations and the Floriculture Industry: Records of the Colorado Flower Growers Association
{{Authority control caryophyllus Symbols of Ohio National symbols of Spain Garden plants Plants described in 1753 Taxa named by Carl Linnaeus