Caricaceae
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The Caricaceae are a
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants t ...
s in the order
Brassicales The Brassicales (or Cruciales) are an order of flowering plants, belonging to the eurosids II group of dicotyledons under the APG II system. One character common to many members of the order is the production of glucosinolate (mustard oil) compo ...
, found primarily in
tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...
regions of
Central Central is an adjective usually referring to being in the center of some place or (mathematical) object. Central may also refer to: Directions and generalised locations * Central Africa, a region in the centre of Africa continent, also known a ...
and
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sou ...
and
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
. They are usually short-lived
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, whic ...
pachycaul Pachycauls are plants with a disproportionately thick trunk for their height, and few branches. This can be the product of exceptional primary growth (as with palms and cycads) or disproportioate secondary growth as with Adansonia. The word is d ...
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from tree ...
s or small to medium-sized
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
s growing to 5–10 m tall. One species, ''
Vasconcellea horovitziana ''Vasconcellea horovitziana'' is a species of plant in the family Caricaceae. It is endemic to Ecuador. Its natural habitat is subtropical or tropical moist lowland forest. It is threatened by habitat loss. It was formerly placed in genus ''Car ...
'' is a liana and the three species of the genus '' Jarilla'' are herbs. Some species, such as the
papaya The papaya (, ), papaw, () or pawpaw () is the plant species ''Carica papaya'', one of the 21 accepted species in the genus '' Carica'' of the family Caricaceae. It was first domesticated in Mesoamerica, within modern-day southern Mexico and ...
, bear edible
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in partic ...
and produce
papain Papain, also known as papaya proteinase I, is a cysteine protease () enzyme present in papaya (''Carica papaya'') and mountain papaya (''Vasconcellea cundinamarcensis''). It is the namesake member of the papain-like protease family. It has wide ...
. Based on molecular analyses, this family has been proposed to have originated in
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
in the early
Cenozoic The Cenozoic ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterised by the dominance of mammals, birds and flowering plants, a cooling and drying climate, and the current configu ...
era, ~66 million years ago (mya). The dispersal from Africa to
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
occurred ~35 mya, possibly via ocean currents from the Congo delta. From Central America, the family reached
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sou ...
19-27 mya. The family comprises six genera and about 34-35 species: *''
Carica ''Carica'' is a genus of flowering plants in the family Caricaceae including the papaya (''C. papaya'' syn. ''C. peltata'', ''C. posoposa''), a widely cultivated fruit tree native to the American tropics. The genus was formerly treated as incl ...
'' – one species, ''Carica papaya'' (
papaya The papaya (, ), papaw, () or pawpaw () is the plant species ''Carica papaya'', one of the 21 accepted species in the genus '' Carica'' of the family Caricaceae. It was first domesticated in Mesoamerica, within modern-day southern Mexico and ...
), Americas *''
Cylicomorpha ''Cylicomorpha'' is a plant genus consisting of two species that are native to the African tropics. They are the only African representatives of the Caricaceae, and are consequently related to the papaya. Habit and appearance They have the habit ...
'' – two species, Africa *''
Horovitzia ''Horovitzia cnidoscoloides'' is a plant species in the family Caricaceae, endemic to the cloud forest of Sierra de Juarez in Oaxaca Oaxaca ( , also , , from nci, Huāxyacac ), officially the Free and Sovereign State of Oaxaca ( es, Estado Li ...
'' – one species, Mexico *''
Jacaratia ''Jacaratia'' is a genus of shrubs or trees in the family Caricaceae. They are native to South and Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the U ...
'' – eight species, Americas *'' Jarilla'' – four species, Americas *''
Vasconcellea ''Vasconcellea'' is a genus with 20 or 26 species of flowering plants in the family Caricaceae. Most were formerly treated in the genus ''Carica'', but have been split out on genetic evidence. The genus name has also been spelled "''Vasconcella'' ...
'' – twenty species, Americas


References


Ghent University: ''Cylicomorpha'' checkliste-Monograph of Caricaceae
Brassicales families Dioecious plants {{Brassicales-stub