Camel farming in Sudan
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A camel (from: la, camelus and grc-gre, κάμηλος (''kamēlos'') from Hebrew or Phoenician: גָמָל ''gāmāl''.) is an
even-toed ungulate The even-toed ungulates (Artiodactyla , ) are ungulates—hoofed animals—which bear weight equally on two (an even number) of their five toes: the third and fourth. The other three toes are either present, absent, vestigial, or pointing poster ...
in the genus ''Camelus'' that bears distinctive fatty deposits known as "humps" on its back. Camels have long been domesticated and, as
livestock Livestock are the domesticated animals raised in an agricultural setting to provide labor and produce diversified products for consumption such as meat, eggs, milk, fur, leather, and wool. The term is sometimes used to refer solely to animal ...
, they provide food (
milk Milk is a white liquid food produced by the mammary glands of mammals. It is the primary source of nutrition for young mammals (including breastfed human infants) before they are able to digest solid food. Immune factors and immune-modula ...
and meat) and textiles (fiber and felt from hair). Camels are
working animal A working animal is an animal, usually domesticated, that is kept by humans and trained to perform tasks instead of being slaughtered to harvest animal products. Some are used for their physical strength (e.g. oxen and draft horses) or for ...
s especially suited to their desert habitat and are a vital means of transport for passengers and cargo. There are three surviving species of camel. The one-humped
dromedary The dromedary (''Camelus dromedarius'' or ;), also known as the dromedary camel, Arabian camel, or one-humped camel, is a large even-toed ungulate, of the genus ''Camelus'', with one hump on its back. It is the tallest of the three species of ...
makes up 94% of the world's camel population, and the two-humped
Bactrian camel The Bactrian camel (''Camelus bactrianus''), also known as the Mongolian camel or domestic Bactrian camel, is a large even-toed ungulate native to the steppes of Central Asia. It has two humps on its back, in contrast to the single-humped dro ...
makes up 6%. The
Wild Bactrian camel The wild Bactrian camel (''Camelus ferus'') is a critically endangered species of camel living in parts of northwestern China and southwestern Mongolia. It is closely related to the Bactrian camel (''Camelus bactrianus''). Both are large, doub ...
is a separate species and is now critically endangered. The word ''camel'' is also used informally in a wider sense, where the more correct term is "camelid", to include all seven species of the family
Camelidae Camelids are members of the biological family Camelidae, the only currently living family in the suborder Tylopoda. The seven extant members of this group are: dromedary camels, Bactrian camels, wild Bactrian camels, llamas, alpacas, vicuñas, ...
: the true camels (the above three species), along with the "New World" camelids: the
llama The llama (; ) (''Lama glama'') is a domesticated South American camelid, widely used as a meat and pack animal by Andean cultures since the Pre-Columbian era. Llamas are social animals and live with others as a herd. Their wool is soft ...
, the
alpaca The alpaca (''Lama pacos'') is a species of South American camelid mammal. It is similar to, and often confused with, the llama. However, alpacas are often noticeably smaller than llamas. The two animals are closely related and can success ...
, the
guanaco The guanaco (; ''Lama guanicoe'') is a camelid native to South America, closely related to the llama. Guanacos are one of two wild South American camelids, the other being the vicuña, which lives at higher elevations. Etymology The guanaco ...
, and the
vicuña The vicuña (''Lama vicugna'') or vicuna (both , very rarely spelled ''vicugna'', its former genus name) is one of the two wild South American camelids, which live in the high alpine areas of the Andes, the other being the guanaco, which live ...
, which belong to the separate tribe
Lamini Lamini (members are called ''laminoids'') is a tribe of the subfamily Camelinae. It contains one extant genus with four species, all exclusively from South America: llamas, alpacas, vicuñas, and guanacos. The former two are domesticated specie ...
. Camelids originated in North America during the
Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', " ...
, with the ancestor of modern camels, ''
Paracamelus ''Paracamelus'' is an extinct genus of camel in the family Camelidae. It originated in North America during the Middle Miocene but crossed the Beringian land bridge into Eurasia during the Late Miocene, approximately 7.5–6.5  million years ...
'', migrating across the
Bering land bridge Beringia is defined today as the land and maritime area bounded on the west by the Lena River in Russia; on the east by the Mackenzie River in Canada; on the north by 72 degrees north latitude in the Chukchi Sea; and on the south by the tip of ...
into Asia during the late
Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
, around 6 million years ago.


Taxonomy


Extant species

3 species are extant:


Biology

The average
life expectancy Life expectancy is a statistical measure of the average time an organism is expected to live, based on the year of its birth, current age, and other demographic factors like sex. The most commonly used measure is life expectancy at birth ...
of a camel is 40 to 50 years. A full-grown adult dromedary camel stands at the shoulder and at the hump. Bactrian camels can be a foot taller. Camels can run at up to in short bursts and sustain speeds of up to . Bactrian camels weigh and dromedaries . The widening toes on a camel's hoof provide supplemental grip for varying soil sediments. The male dromedary camel has an organ called a dulla in his throat, a large, inflatable sac he extrudes from his mouth when in rut to assert dominance and attract females. It resembles a long, swollen, pink tongue hanging out of the side of his mouth. Camels mate by having both male and female sitting on the ground, with the male mounting from behind. The male usually
ejaculates Ejaculation is the discharge of semen (the ''ejaculate''; normally containing sperm) from the male reproductory tract as a result of an orgasm. It is the final stage and natural objective of male sexual stimulation, and an essential componen ...
three or four times within a single mating session. Camelids are the only ungulates to mate in a sitting position.


Ecological and behavioral adaptations

Camels do not directly store water in their humps; they are reservoirs of fatty tissue. When this tissue is metabolized, it yields more than one gram of water for every gram of fat processed. This fat metabolization, while releasing energy, causes water to evaporate from the lungs during
respiration Respiration may refer to: Biology * Cellular respiration, the process in which nutrients are converted into useful energy in a cell ** Anaerobic respiration, cellular respiration without oxygen ** Maintenance respiration, the amount of cellul ...
(as oxygen is required for the metabolic process): overall, there is a net decrease in water. Camels have a series of physiological adaptations that allow them to withstand long periods of time without any external source of water. The dromedary camel can drink as seldom as once every 10 days even under very hot conditions, and can lose up to 30% of its body mass due to dehydration. Unlike other mammals, camels'
red blood cell Red blood cells (RBCs), also referred to as red cells, red blood corpuscles (in humans or other animals not having nucleus in red blood cells), haematids, erythroid cells or erythrocytes (from Greek ''erythros'' for "red" and ''kytos'' for "holl ...
s are oval rather than circular in shape. This facilitates the flow of red blood cells during dehydration and makes them better at withstanding high
osmotic Osmosis (, ) is the spontaneous net movement or diffusion of solvent molecules through a selectively-permeable membrane from a region of high water potential (region of lower solute concentration) to a region of low water potential (region o ...
variation without rupturing when drinking large amounts of water: a camel can drink of water in three minutes. Camels are able to withstand changes in
body temperature Thermoregulation is the ability of an organism to keep its body temperature within certain boundaries, even when the surrounding temperature is very different. A thermoconforming organism, by contrast, simply adopts the surrounding temperature ...
and water consumption that would kill most other mammals. Their temperature ranges from at dawn and steadily increases to by sunset, before they cool off at night again. In general, to compare between camels and the other livestock, camels lose only 1.3 liters of fluid intake every day while the other livestock lose 20 to 40 liters per day. Maintaining the brain temperature within certain limits is critical for animals; to assist this, camels have a
rete mirabile A rete mirabile (Latin for "wonderful net"; plural retia mirabilia) is a complex of arteries and veins lying very close to each other, found in some vertebrates, mainly warm-blooded ones. The rete mirabile utilizes countercurrent blood flow within ...
, a complex of arteries and veins lying very close to each other which utilizes countercurrent blood flow to cool blood flowing to the brain.
Inside Nature's Giants ''Inside Nature's Giants'' is a British science documentary, first broadcast in June 2009 by Channel 4. The documentary shows experts performing dissection on some of nature's largest animals, including whales and elephants. The programme is ...
. Channel 4 (UK) documentary. Transmitted 30 August 2011
Camels rarely sweat, even when ambient temperatures reach . Any sweat that does occur evaporates at the skin level rather than at the surface of their coat; the heat of vaporization therefore comes from body heat rather than ambient heat. Camels can withstand losing 25% of their body weight in water, whereas most other mammals can withstand only about 12–14% dehydration before
cardiac failure Heart failure (HF), also known as congestive heart failure (CHF), is a syndrome, a group of signs and symptoms caused by an impairment of the heart's blood pumping function. Symptoms typically include shortness of breath, excessive fatigue, a ...
results from circulatory disturbance. When the camel exhales,
water vapor (99.9839 °C) , - , Boiling point , , - , specific gas constant , 461.5 J/( kg·K) , - , Heat of vaporization , 2.27 MJ/kg , - , Heat capacity , 1.864 kJ/(kg·K) Water vapor, water vapour or aqueous vapor is the gaseous p ...
becomes trapped in their
nostrils A nostril (or naris , plural ''nares'' ) is either of the two orifices of the nose. They enable the entry and exit of air and other gasses through the nasal cavities. In birds and mammals, they contain branched bones or cartilages called turbi ...
and is reabsorbed into the body as a means to conserve water. Camels eating green herbage can ingest sufficient moisture in milder conditions to maintain their bodies' hydrated state without the need for drinking. The camel's thick coat insulates it from the intense heat radiated from desert sand; a shorn camel must sweat 50% more to avoid overheating. During the summer the coat becomes lighter in color, reflecting light as well as helping avoid sunburn. The camel's long legs help by keeping its body farther from the ground, which can heat up to . Dromedaries have a pad of thick tissue over the
sternum The sternum or breastbone is a long flat bone located in the central part of the chest. It connects to the ribs via cartilage and forms the front of the rib cage, thus helping to protect the heart, lungs, and major blood vessels from injury. Sha ...
called the ''pedestal''. When the animal lies down in a sternal recumbent position, the pedestal raises the body from the hot surface and allows cooling air to pass under the body. Camels' mouths have a thick leathery lining, allowing them to chew thorny desert plants. Long eyelashes and ear hairs, together with nostrils that can close, form a barrier against sand. If sand gets lodged in their eyes, they can dislodge it using their transparent third eyelid (also known as the nictitating membrane). The camels' gait and widened feet help them move without sinking into the sand. The
kidney The kidneys are two reddish-brown bean-shaped organs found in vertebrates. They are located on the left and right in the retroperitoneal space, and in adult humans are about in length. They receive blood from the paired renal arteries; blo ...
s and
intestines The gastrointestinal tract (GI tract, digestive tract, alimentary canal) is the tract or passageway of the digestive system that leads from the mouth to the anus. The GI tract contains all the major organs of the digestive system, in humans and ...
of a camel are very efficient at reabsorbing water. Camels' kidneys have a 1:4
cortex Cortex or cortical may refer to: Biology * Cortex (anatomy), the outermost layer of an organ ** Cerebral cortex, the outer layer of the vertebrate cerebrum, part of which is the ''forebrain'' *** Motor cortex, the regions of the cerebral cortex i ...
to medulla ratio. Thus, the medullary part of a camel's kidney occupies twice as much area as a cow's kidney. Secondly,
renal corpuscles A renal corpuscle (also called malpighian body) is the blood-filtering component of the nephron of the kidney. It consists of a glomerulus - a tuft of capillaries composed of endothelial cells, and a glomerular capsule known as Bowman's capsul ...
have a smaller diameter, which reduces surface area for filtration. These two major anatomical characteristics enable camels to conserve water and limit the volume of urine in extreme desert conditions.Rehan S and AS Qureshi, 2006. Microscopic evaluation of the heart, kidneys and adrenal glands of one-humped camel calves (Camelus dromedarius) using semi automated image analysis system. J Camel Pract Res. 13(2): 123 Camel urine comes out as a thick syrup, and camel faeces are so dry that they do not require drying when the
Bedouins The Bedouin, Beduin, or Bedu (; , singular ) are nomadic Arab tribes who have historically inhabited the desert regions in the Arabian Peninsula, North Africa, the Levant, and Mesopotamia. The Bedouin originated in the Syrian Desert and Ar ...
use them to fuel fires. The camel
immune system The immune system is a network of biological processes that protects an organism from diseases. It detects and responds to a wide variety of pathogens, from viruses to parasitic worms, as well as cancer cells and objects such as wood splint ...
differs from those of other mammals. Normally, the Y-shaped
antibody An antibody (Ab), also known as an immunoglobulin (Ig), is a large, Y-shaped protein used by the immune system to identify and neutralize foreign objects such as pathogenic bacteria and viruses. The antibody recognizes a unique molecule of the ...
molecules consist of two heavy (or long) chains along the length of the Y, and two light (or short) chains at each tip of the Y. Camels, in addition to these, also have antibodies made of only two heavy chains, a trait that makes them smaller and more durable. These "heavy-chain-only" antibodies, discovered in 1993, are thought to have developed 50 million years ago, after camelids split from ruminants and pigs. Camels suffer from surra caused by ''
Trypanosoma evansi ''Trypanosoma evansi'' is a parasitic species of excavate trypanosome in the genus ''Trypanosoma'' that causes one form of surra in animals. Discovered by Griffith Evans in 1880 at Dera Ismail Khan (British India), it is the first known tryp ...
'' wherever camels are domesticated in the world, and resultantly camels have evolved trypanolytic antibodies as with many mammals. In the future, nanobody/
single-domain antibody A single-domain antibody (sdAb), also known as a nanobody, is an antibody fragment consisting of a single monomeric variable antibody domain. Like a whole antibody, it is able to bind selectively to a specific antigen. With a molecular weight of ...
therapy will surpass natural camel antibodies by reaching locations currently unreachable due to natural antibodies' larger size. Such therapies may also be suitable for other mammals. Tran ''et al.'' 2009 provides a new reference test for surra (''T. evansi'') of camel. They use recombinant (rISG75, an Invariant Surface Glycoprotein) and
ELISA The enzyme-linked immunosorbent assay (ELISA) (, ) is a commonly used analytical biochemistry assay, first described by Eva Engvall and Peter Perlmann in 1971. The assay uses a solid-phase type of enzyme immunoassay (EIA) to detect the presen ...
. The Tran test has high test specificity and appears likely to work just as well for ''T. evansi'' in other hosts, and for a pan-''
Trypanozoon ''Trypanosoma'' is a genus of kinetoplastids (class Trypanosomatidae), a monophyletic group of unicellular parasitic flagellate protozoa. Trypanosoma is part of the phylum Sarcomastigophora. The name is derived from the Greek ''trypano-'' ( ...
'' test, which would also be useful for '' T. b. brucei'', '' T. b. gambiense'', '' T. b. rhodesiense'', and '' T. equiperdum''.


Genetics

The karyotypes of different camelid species have been studied earlier by many groups, but no agreement on chromosome nomenclature of camelids has been reached. A 2007 study flow sorted camel chromosomes, building on the fact that camels have 37 pairs of chromosomes (2n=74), and found that the karyotype consisted of one
metacentric Metacentric may refer to: * Metacentric height The metacentric height (GM) is a measurement of the initial static stability of a floating body. It is calculated as the distance between the centre of gravity of a ship and its metacentre. A larger ...
, three submetacentric, and 32 acrocentric autosomes. The Y is a small metacentric chromosome, while the X is a large metacentric chromosome. The
hybrid camel A hybrid camel is a hybrid between a Bactrian camel (''Camelus bactrianus'') and dromedary (''Camelus dromedarius''). Names Hybrid camels have different names depending on zone and language. Some names include turkoman, tülu, bukht, nar, in ...
, a hybrid between Bactrian and dromedary camels, has one hump, though it has an indentation deep that divides the front from the back. The hybrid is at the shoulder and tall at the hump. It weighs an average of and can carry around , which is more than either the dromedary or Bactrian can. According to molecular data, the wild Bactrian camel (''C. ferus'') separated from the domestic Bactrian camel (''C. bactrianus'') about 1 million years ago. New World and Old World camelids diverged about 11 million years ago. In spite of this, these species can hybridize and produce viable offspring. The cama is a camel-llama hybrid bred by scientists to see how closely related the parent species are. Scientists collected semen from a camel via an artificial vagina and inseminated a llama after stimulating ovulation with gonadotrophin injections. The cama is halfway in size between a camel and a llama and lacks a hump. It has ears intermediate between those of camels and llamas, longer legs than the llama, and partially
cloven hooves A cloven hoof, cleft hoof, divided hoof or split hoof is a hoof split into two toes. This is found on members of the mammalian order Artiodactyla. Examples of mammals that possess this type of hoof are cattle, deer, pigs, antelopes, gazelles, ...
. Like the mule, camas are sterile, despite both parents having the same number of chromosomes.


Evolution

The earliest known camel, called '' Protylopus'', lived in North America 40 to 50 million years ago (during the
Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', " ...
). It was about the size of a rabbit and lived in the open woodlands of what is now
South Dakota South Dakota (; Sioux: , ) is a U.S. state in the North Central region of the United States. It is also part of the Great Plains. South Dakota is named after the Lakota and Dakota Sioux Native American tribes, who comprise a large porti ...
. By 35 million years ago, the ''
Poebrotherium ''Poebrotherium'' ( ) is an extinct genus of camelid, endemic to North America. They lived from the Eocene to Miocene epochs, 46.3—13.6 mya, existing for approximately . Discovery and history ''Poebrotherium'' was first named by scientist J ...
'' was the size of a goat and had many more traits similar to camels and llamas. The hoofed '' Stenomylus'', which walked on the tips of its toes, also existed around this time, and the long-necked ''
Aepycamelus ''Aepycamelus'' is an extinct genus of camelids that lived during the Miocene 20.6–4.9 million years ago, existing for about . Its name is derived from the Homeric Greek , "high and steep" and κάμηλος – "camel"; thus, "high camel"; ''a ...
'' evolved in the
Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
. The ancestor of modern camels, ''
Paracamelus ''Paracamelus'' is an extinct genus of camel in the family Camelidae. It originated in North America during the Middle Miocene but crossed the Beringian land bridge into Eurasia during the Late Miocene, approximately 7.5–6.5  million years ...
'', migrated into Eurasia from North America via Beringia during the late Miocene, between 7.5 and 6.5 million years ago. During the Pleistocene, around 3 to 1 million years ago, the North American Camelidae spread to South America as part of the Great American Interchange via the newly formed Isthmus of Panama, where they gave rise to
guanacos The guanaco (; ''Lama guanicoe'') is a camelid native to South America, closely related to the llama. Guanacos are one of two wild South American camelids, the other being the vicuña, which lives at higher elevations. Etymology The guanaco g ...
and related animals. Populations of ''Paracamelus'' continued to exist in the North American Arctic into the Early Pleistocene. This creature is estimated to have stood around tall. The Bactrian camel diverged from the dromedary about 1 million years ago, according to the fossil record. The last camel native to North America was ''
Camelops hesternus ''Camelops''Being occasionally called ''Western Camel'' or ''Yesterday's Camel''. is an extinct genus of camels that lived in North and Central America, ranging from Alaska to Guatemala, from the middle Pliocene to the end of the Pleistocene. It ...
'', which vanished along with
horse The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 million yea ...
s,
short-faced bear The Tremarctinae or short-faced bears is a subfamily of Ursidae that contains one living representative, the spectacled bear (''Tremarctos ornatus'') of South America, and several extinct species from four genera: the Florida spectacled bear ('' ...
s,
mammoth A mammoth is any species of the extinct elephantid genus ''Mammuthus'', one of the many genera that make up the order of trunked mammals called proboscideans. The various species of mammoth were commonly equipped with long, curved tusks an ...
s and
mastodon A mastodon ( 'breast' + 'tooth') is any proboscidean belonging to the extinct genus ''Mammut'' (family Mammutidae). Mastodons inhabited North and Central America during the late Miocene or late Pliocene up to their extinction at the end of th ...
s, ground sloths, sabertooth cats, and many other megafauna, coinciding with the migration of humans from Asia at the end of the Pleistocene, around 15–11,000 years ago. File:Stenomylus.jpg, alt=A drawing of two early camels, Stenomylus illustration File:NMNH-USNMV16601Stenomylus.tif, Stenomylus skeleton File:NMNH-USNMV15917Poebrotherium.jpg,
Poebrotherium ''Poebrotherium'' ( ) is an extinct genus of camelid, endemic to North America. They lived from the Eocene to Miocene epochs, 46.3—13.6 mya, existing for approximately . Discovery and history ''Poebrotherium'' was first named by scientist J ...
skeleton File:NMNH-USNM244271 2.jpg,
Procamelus ''Procamelus'' is an extinct genus of Camelini, camel endemic to North America. It lived from the Middle to Late Miocene 16.0—5.3 Annum, mya, existing for approximately . The name is derived from the Greek πρό, meaning "before" or denoting p ...
skull File:Camelops hesternus.jpg, alt=, ''
Camelops hesternus ''Camelops''Being occasionally called ''Western Camel'' or ''Yesterday's Camel''. is an extinct genus of camels that lived in North and Central America, ranging from Alaska to Guatemala, from the middle Pliocene to the end of the Pleistocene. It ...
'', the last true camel native to North America


Domestication

Like
horse The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 million yea ...
s, camels originated in North America and eventually spread across Beringia to Asia. They survived in the Old World, and eventually humans domesticated them and spread them globally. Along with many other megafauna in North America, the original wild camels were wiped out during the spread of the first
indigenous peoples of the Americas The Indigenous peoples of the Americas are the inhabitants of the Americas before the arrival of the European settlers in the 15th century, and the ethnic groups who now identify themselves with those peoples. Many Indigenous peoples of the A ...
from Asia into North America, 10 to 12,000 years ago; although fossils have never been associated with definitive evidence of hunting. Most camels surviving today are domesticated. Although
feral A feral () animal or plant is one that lives in the wild but is descended from domesticated individuals. As with an introduced species, the introduction of feral animals or plants to non-native regions may disrupt ecosystems and has, in some ...
populations exist in Australia, India and Kazakhstan, wild camels survive only in the
wild Bactrian camel The wild Bactrian camel (''Camelus ferus'') is a critically endangered species of camel living in parts of northwestern China and southwestern Mongolia. It is closely related to the Bactrian camel (''Camelus bactrianus''). Both are large, doub ...
population of the Gobi Desert.


History

When humans first domesticated camels is disputed. Dromedaries may have first been domesticated by humans in
Somalia Somalia, , Osmanya script: 𐒈𐒝𐒑𐒛𐒐𐒘𐒕𐒖; ar, الصومال, aṣ-Ṣūmāl officially the Federal Republic of SomaliaThe ''Federal Republic of Somalia'' is the country's name per Article 1 of thProvisional Constituti ...
or South Arabia sometime during the
3rd millennium BC The 3rd millennium BC spanned the years 3000 through 2001 BC. This period of time corresponds to the Early to Middle Bronze Age, characterized by the early empires in the Ancient Near East. In Ancient Egypt, the Early Dynastic Period is followe ...
, the Bactrian in
central Asia Central Asia, also known as Middle Asia, is a subregion, region of Asia that stretches from the Caspian Sea in the west to western China and Mongolia in the east, and from Afghanistan and Iran in the south to Russia in the north. It includes t ...
around 2,500 BC, as at Shar-i Sokhta (also known as the Burnt City),
Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...
. Martin Heide's 2010 work on the domestication of the camel tentatively concludes that humans had domesticated the Bactrian camel by at least the middle of the third millennium somewhere east of the
Zagros Mountains The Zagros Mountains ( ar, جبال زاغروس, translit=Jibal Zaghrus; fa, کوه‌های زاگرس, Kuh hā-ye Zāgros; ku, چیاکانی زاگرۆس, translit=Çiyakani Zagros; Turkish: ''Zagros Dağları''; Luri: ''Kuh hā-ye Zāgr ...
, with the practice then moving into Mesopotamia. Heide suggests that mentions of camels "in the patriarchal narratives may refer, at least in some places, to the Bactrian camel", while noting that the camel is not mentioned in relationship to
Canaan Canaan (; Phoenician: 𐤊𐤍𐤏𐤍 – ; he, כְּנַעַן – , in pausa – ; grc-bib, Χανααν – ;The current scholarly edition of the Greek Old Testament spells the word without any accents, cf. Septuaginta : id est Vetus T ...
. Recent excavations in the
Timna Valley The Timna Valley (תִּמְנָע, ) is located in southern Israel in the southwestern Arava/Arabah, approximately north of the Gulf of Aqaba and the city of Eilat. The area is rich in copper ore and has been mined since the 5th millennium ...
by Lidar Sapir-Hen and Erez Ben-Yosef discovered what may be the earliest domestic camel bones yet found in Israel or even outside the Arabian Peninsula, dating to around 930 BC. This garnered considerable media coverage, as it is strong evidence that the stories of
Abraham Abraham, ; ar, , , name=, group= (originally Abram) is the common Hebrew patriarch of the Abrahamic religions, including Judaism, Christianity, and Islam. In Judaism, he is the founding father of the special relationship between the Je ...
,
Jacob Jacob (; ; ar, يَعْقُوب, Yaʿqūb; gr, Ἰακώβ, Iakṓb), later given the name Israel, is regarded as a patriarch of the Israelites and is an important figure in Abrahamic religions, such as Judaism, Christianity, and Islam. J ...
,
Esau Esau ''Ēsaû''; la, Hesau, Esau; ar, عِيسَوْ ''‘Īsaw''; meaning "hairy"Easton, M. ''Illustrated Bible Dictionary'', (, , 2006, p. 236 or "rough".Mandel, D. ''The Ultimate Who's Who in the Bible'', (.), 2007, p. 175 is the elder son o ...
, and
Joseph Joseph is a common male given name, derived from the Hebrew Yosef (יוֹסֵף). "Joseph" is used, along with "Josef", mostly in English, French and partially German languages. This spelling is also found as a variant in the languages of the mo ...
were written after this time. The existence of camels in Mesopotamia—but not in the eastern Mediterranean lands—is not a new idea. The historian
Richard Bulliet Richard W. Bulliet (born 1940) is a professor of history at Columbia University who specializes in the history of Islamic society and institutions, the history of technology, and the history of the role of animals in human society. Early life ...
did not think that the occasional mention of camels in the Bible meant that the domestic camels were common in the Holy Land at that time. The archaeologist
William F. Albright William Foxwell Albright (May 24, 1891– September 19, 1971) was an American archaeologist, biblical scholar, philologist, and expert on ceramics. He is considered "one of the twentieth century's most influential American biblical scholars." ...
, writing even earlier, saw camels in the Bible as an anachronism. The official report by Sapir-Hen and Ben-Joseph notes:
The introduction of the dromedary camel (Camelus dromedarius) as a pack animal to the
southern Levant The Southern Levant is a geographical region encompassing the southern half of the Levant. It corresponds approximately to modern-day Israel, Palestine, and Jordan; some definitions also include southern Lebanon, southern Syria and/or the Sinai P ...
... substantially facilitated trade across the vast deserts of Arabia, promoting both economic and social change (e.g., Kohler 1984; Borowski 1998: 112–116; Jasmin 2005). This ... has generated extensive discussion regarding the date of the earliest domestic camel in the southern Levant (and beyond) (e.g., Albright 1949: 207; Epstein 1971: 558–584; Bulliet 1975; Zarins 1989; Köhler-Rollefson 1993; Uerpmann and Uerpmann 2002; Jasmin 2005; 2006; Heide 2010; Rosen and Saidel 2010; Grigson 2012). Most scholars today agree that the dromedary was exploited as a pack animal sometime in the early
Iron Age The Iron Age is the final epoch of the three-age division of the prehistory and protohistory of humanity. It was preceded by the Stone Age ( Paleolithic, Mesolithic, Neolithic) and the Bronze Age ( Chalcolithic). The concept has been mostl ...
(not before the 12th century C
and concludes:
Current data from copper smelting sites of the Aravah Valley enable us to pinpoint the introduction of domestic camels to the southern Levant more precisely based on stratigraphic contexts associated with an extensive suite of radiocarbon dates. The data indicate that this event occurred not earlier than the last third of the 10th century Cand most probably during this time. The coincidence of this event with a major reorganization of the copper industry of the region—attributed to the results of the campaign of Pharaoh Shoshenq I—raises the possibility that the two were connected, and that camels were introduced as part of the efforts to improve efficiency by facilitating trade.
File:Camel cart.JPG, alt= A camel harnessed to a cart loaded with branches and twigs, A camel serving as a draft animal in
Pakistan Pakistan ( ur, ), officially the Islamic Republic of Pakistan ( ur, , label=none), is a country in South Asia. It is the world's fifth-most populous country, with a population of almost 243 million people, and has the world's second-lar ...
(2009) File:A camel with its rider playing kettle drums..jpg, alt= A painting of a man sitting on a camel and playing the drums, A camel in a ceremonial procession, its rider playing
kettledrum Timpani (; ) or kettledrums (also informally called timps) are musical instruments in the percussion family. A type of drum categorised as a hemispherical drum, they consist of a membrane called a head stretched over a large bowl traditionally ...
s,
Mughal Empire The Mughal Empire was an early-modern empire that controlled much of South Asia between the 16th and 19th centuries. Quote: "Although the first two Timurid emperors and many of their noblemen were recent migrants to the subcontinent, the d ...
(c. 1840) File:Negev camel petroglyph.jpg, Petroglyph of a camel,
Negev The Negev or Negeb (; he, הַנֶּגֶב, hanNegév; ar, ٱلنَّقَب, an-Naqab) is a desert and semidesert region of southern Israel. The region's largest city and administrative capital is Beersheba (pop. ), in the north. At its sout ...
, southern
Israel Israel (; he, יִשְׂרָאֵל, ; ar, إِسْرَائِيل, ), officially the State of Israel ( he, מְדִינַת יִשְׂרָאֵל, label=none, translit=Medīnat Yīsrāʾēl; ), is a country in Western Asia. It is situated ...
(prior to c. 5300 BC) File:Bartholomeus Breenbergh 002.jpg, ''Joseph Sells Grain'' by
Bartholomeus Breenbergh Bartholomeus Breenbergh (before 13 November 1598 – after 3 October 1657) was a Dutch Golden Age painter of Italian and Italianate landscapes, in Rome (1619-1630) and Amsterdam (1630-1657). Biography Little is known of his early life. In his ...
(1655), showing camel with rider at left


Textiles

Desert tribes and Mongolian nomads use camel hair for tents,
yurt A yurt (from the Turkic languages) or ger ( Mongolian) is a portable, round tent covered and insulated with skins or felt and traditionally used as a dwelling by several distinct nomadic groups in the steppes and mountains of Central Asia ...
s, clothing, bedding and accessories. Camels have outer guard hairs and soft inner down, and the fibers are sorted by color and age of the animal. The guard hairs can be felted for use as waterproof coats for the herdsmen, while the softer hair is used for premium goods. The fiber can be spun for use in weaving or made into yarns for hand knitting or crochet. Pure camel hair is recorded as being used for western garments from the 17th century onwards, and from the 19th century a mixture of wool and camel hair was used.


Military uses

By at least 1200 BC the first camel saddles had appeared, and
Bactrian camels The Bactrian camel (''Camelus bactrianus''), also known as the Mongolian camel or domestic Bactrian camel, is a large even-toed ungulate native to the steppes of Central Asia. It has two humps on its back, in contrast to the single-humped drome ...
could be ridden. The first saddle was positioned to the back of the camel, and control of the Bactrian camel was exercised by means of a stick. However, between 500 and 100 BC, Bactrian camels came into military use. New saddles, which were inflexible and bent, were put over the humps and divided the rider's weight over the animal. In the seventh century BC the military Arabian saddle evolved, which again improved the saddle design slightly. Military forces have used camel cavalries in wars throughout Africa, the Middle East, and into the modern-day
Border Security Force The Border Security Force (BSF) is India's border guarding organisation on its border with Pakistan and Bangladesh. It is one of the seven Central Armed Police Forces (CAPF) of India, and was raised in the wake of the 1965 war on 1 December 1 ...
(BSF) of
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
(though as of July 2012, the BSF planned the replacement of camels with ATVs). The first documented use of camel cavalries occurred in the
Battle of Qarqar The Battle of Qarqar (or Ḳarḳar) was fought in 853 BC when the army of the Neo-Assyrian Empire led by Emperor Shalmaneser III encountered an allied army of eleven kings at Qarqar led by Hadadezer, called in Assyrian ''Adad-idir'' and possi ...
in 853 BC. Armies have also used camels as freight animals instead of horses and mules. The
East Roman Empire The Byzantine Empire, also referred to as the Eastern Roman Empire or Byzantium, was the continuation of the Roman Empire primarily in its eastern provinces during Late Antiquity and the Middle Ages, when its capital city was Constantinopl ...
used
auxiliary Auxiliary may refer to: * A backup site or system In language * Auxiliary language (disambiguation) * Auxiliary verb In military and law enforcement * Auxiliary police * Auxiliaries, civilians or quasi-military personnel who provide support of ...
forces known as '' dromedarii'', whom the Romans recruited in desert provinces. The camels were used mostly in combat because of their ability to scare off horses at close range (horses are afraid of the camels' scent), a quality famously employed by the
Achaemenid The Achaemenid Empire or Achaemenian Empire (; peo, 𐎧𐏁𐏂, , ), also called the First Persian Empire, was an ancient Iranian empire founded by Cyrus the Great in 550 BC. Based in Western Asia, it was contemporarily the largest emp ...
Persians when fighting Lydia in the
Battle of Thymbra The Battle of Thymbra was the decisive battle in the war between Croesus of the Lydian Kingdom and Cyrus the Great of the Achaemenid Empire. Cyrus, after he had pursued Croesus into Lydia after the drawn Battle of Pteria, met the remains of ...
(547 BC).


19th and 20th centuries

The
United States Army The United States Army (USA) is the land warfare, land military branch, service branch of the United States Armed Forces. It is one of the eight Uniformed services of the United States, U.S. uniformed services, and is designated as the Army o ...
established the
U.S. Camel Corps The United States Camel Corps was a mid-19th-century experiment by the United States Army in using camels as pack animals in the Southwestern United States. Although the camels proved to be hardy and well suited to travel through the region, th ...
, stationed in
California California is a state in the Western United States, located along the Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the most populous U.S. state and the 3rd largest by area. It is also the m ...
, in the 19th century. One may still see stables at the Benicia Arsenal in Benicia, California, where they nowadays serve as the Benicia Historical Museum. Though the experimental use of camels was seen as a success (
John B. Floyd John Buchanan Floyd (June 1, 1806 – August 26, 1863) was the 31st Governor of Virginia, U.S. Secretary of War, and the Confederate general in the American Civil War who lost the crucial Battle of Fort Donelson. Early family life John Buch ...
,
Secretary of War The secretary of war was a member of the U.S. president's Cabinet, beginning with George Washington's administration. A similar position, called either "Secretary at War" or "Secretary of War", had been appointed to serve the Congress of the ...
in 1858, recommended that funds be allocated towards obtaining a thousand more camels), the outbreak of the
American Civil War The American Civil War (April 12, 1861 – May 26, 1865; also known by other names) was a civil war in the United States. It was fought between the Union ("the North") and the Confederacy ("the South"), the latter formed by states ...
in 1861 saw the end of the Camel Corps: Texas became part of the Confederacy, and most of the camels were left to wander away into the desert. France created a ''
méhariste Méhariste is a French word that roughly translates to camel cavalry. The word is most commonly used as a designation of military units. French camel corps Origins France created a corps of méhariste camel companies (''Compagnies Méharistes ...
'' camel corps in 1912 as part of the Armée d'Afrique in the Sahara in order to exercise greater control over the camel-riding
Tuareg The Tuareg people (; also spelled Twareg or Touareg; endonym: ''Imuhaɣ/Imušaɣ/Imašeɣăn/Imajeɣăn'') are a large Berber ethnic group that principally inhabit the Sahara in a vast area stretching from far southwestern Libya to southern Alg ...
and Arab insurgents, as previous efforts to defeat them on foot had failed. The Free French Camel Corps fought during
World War II World War II or the Second World War, often abbreviated as WWII or WW2, was a world war that lasted from 1939 to 1945. It involved the vast majority of the world's countries—including all of the great powers—forming two opposing ...
, and camel-mounted units remained in service until the end of French rule over Algeria in 1962. In 1916, the British created the
Imperial Camel Corps The Imperial Camel Corps Brigade (ICCB) was a camel-mounted infantry brigade that the British Empire raised in December 1916 during the First World War for service in the Middle East. From a small beginning the unit eventually grew to a brigad ...
. It was originally used to fight the Senussi, but was later used in the Sinai and Palestine Campaign in
World War I World War I (28 July 1914 11 November 1918), often abbreviated as WWI, was one of the deadliest global conflicts in history. Belligerents included much of Europe, the Russian Empire, the United States, and the Ottoman Empire, with fightin ...
. The Imperial Camel Corps comprised infantrymen mounted on camels for movement across desert, though they dismounted at battle sites and fought on foot. After July 1918, the Corps began to become run down, receiving no new reinforcements, and was formally disbanded in 1919. In World War I, the British Army also created the Egyptian Camel Transport Corps, which consisted of a group of Egyptian camel drivers and their camels. The Corps supported British war operations in Sinai Peninsula, Sinai, Palestine, and Syria by transporting supplies to the troops. The Somaliland Camel Corps was created by colonial authorities in British Somaliland in 1912; it was disbanded in 1944. Bactrian camels were used by Romanian forces during
World War II World War II or the Second World War, often abbreviated as WWII or WW2, was a world war that lasted from 1939 to 1945. It involved the vast majority of the world's countries—including all of the great powers—forming two opposing ...
in the Caucasian region. At the same period the Soviet units operating around Astrakhan in 1942 adopted local camels as draft animals due to shortage of trucks and horses, and kept them even after moving out of the area. Despite severe losses, some of these camels ended up as far west as to Battle of Berlin, Berlin itself. The Bikaner Camel Corps of British India fought alongside the British Indian Army in World Wars I and II. The ''Tropas Nómadas'' (Nomad Troops) were an auxiliary regiment of Sahrawi people, Sahrawi tribesmen serving in the colonial army in Spanish Sahara (today Western Sahara). Operational from the 1930s until the end of the Spanish presence in the territory in 1975, the ''Tropas Nómadas'' were equipped with small arms and led by Spanish officers. The unit guarded outposts and sometimes conducted patrols on camelback.


21st century competition

At the King Abdulaziz Camel Festival, in Saudi Arabia, thousands of camels are paraded and are judged on their lips and humps. The festival also features camel racing and camel milk tasting and has combined prize money of $57m (£40m). In 2018, 12 camels were disqualified from the beauty contest after it was discovered their owners had tried to improve their camel's good looks with injections of botox, into the animals' lips, noses and jaws. In 2021 over 40 camels were disqualified for acts of tampering and deception in beautifying camels.


Food uses


Dairy

Camel milk is a staple food of desert nomad tribes and is sometimes considered a meal itself; a nomad can live on only camel milk for almost a month. Camel milk can readily be made into yogurt, but can only be made into butter if it is soured first, churned, and a clarifying agent is then added. Until recently, camel milk could not be made into camel milk, camel cheese because rennet was unable to coagulate the milk proteins to allow the collection of curds. Developing less wasteful uses of the milk, the Food and Agriculture Organization, FAO commissioned Professor J.P. Ramet of the École Nationale Supérieure d'Agronomie et des Industries Alimentaires, who was able to produce curdling by the addition of calcium phosphate and vegetable rennet in the 1990s. The cheese produced from this process has low levels of cholesterol and is easy to digest, even for the lactose intolerant. Camel milk can also be made into ice cream.


Meat

They provide food in the form of meat and milk. Approximately 3.3 million camels and camelids are slaughtered each year for meat worldwide. A camel carcass can provide a substantial amount of meat. The male dromedary carcass can weigh , while the carcass of a male Bactrian can weigh up to . The carcass of a female dromedary weighs less than the male, ranging between . The brisket, ribs and loin are among the preferred parts, and the hump is considered a delicacy. The hump contains "white and sickly fat", which can be used to make the ''khli'' (preserved meat) of mutton, beef, or camel. On the other hand, camel milk and meat are rich in protein, vitamins, glycogen, and other nutrients making them essential in the diet of many people. From chemical composition to meat quality, the dromedary camel is the preferred breed for meat production. It does well even in arid areas due to its unusual physiological behaviors and characteristics, which include tolerance to extreme temperatures, radiation from the sun, water paucity, rugged landscape and low vegetation. Camel meat is reported to taste like coarse beef, but older camels can prove to be very tough, although camel meat becomes tenderer the more it is cooked. Camel is one of the animals that can be ritually slaughtered and divided into three portions (one for the home, one for extended family/social networks, and one for those who cannot afford to slaughter an animal themselves) for the Qurban (Islamic ritual sacrifice), qurban of Eid al-Adha. The Abu Dhabi Officers' Club serves a camel burger mixed with beef or lamb fat in order to improve the texture and taste. In Karachi, Pakistan, some restaurants prepare nihari from camel meat. Specialist camel butchers provide expert cuts, with the hump considered the most popular. Camel meat has been eaten for centuries. It has been recorded by Ancient Greece, ancient Greek writers as an available dish at banquets in ancient Persia, usually roasted whole. The Ancient Rome, Roman emperor Elagabalus, Heliogabalus enjoyed camel's heel. Camel meat is mainly eaten in certain regions, including Eritrea,
Somalia Somalia, , Osmanya script: 𐒈𐒝𐒑𐒛𐒐𐒘𐒕𐒖; ar, الصومال, aṣ-Ṣūmāl officially the Federal Republic of SomaliaThe ''Federal Republic of Somalia'' is the country's name per Article 1 of thProvisional Constituti ...
, Djibouti, Saudi Arabia, Egypt, Syria, Libya, Sudan, Ethiopia, Kazakhstan, and other arid regions where alternative forms of protein may be limited or where camel meat has had a long cultural history. Camel blood is also consumable, as is the case among pastoralists in northern Kenya, where camel blood is drunk with milk and acts as a key source of iron, vitamin D, salts and minerals. A 2005 report issued jointly by the Saudi Ministry of Health (Saudi Arabia), Ministry of Health and the United States Centers for Disease Control and Prevention details four cases of human bubonic plague resulting from the ingestion of raw camel liver.


Australia

Camel meat is also occasionally found in Australian cuisine: for example, a camel lasagna is available in Alice Springs. Australia has exported camel meat, primarily to the Middle East but also to Europe and the US, for many years. The meat is very popular among African Australians, East African Australians, such as Somalis, and other Australians have also been buying it. The feral nature of the animals means they produce a different type of meat to farmed camels in other parts of the world, and it is sought after because it is disease-free, and a unique genetic group. Demand is outstripping supply, and governments are being urged not to cull the camels, but redirect the cost of the cull into developing the market. Australia has seven camel dairies, which produce milk, cheese and skincare products in addition to meat.


Religion


Islam

Muslims consider camel meat ''halal'' ( ar, حلال, 'allowed'). However, according to some Islamic schools and branches, Islamic schools of thought, a state of impurity is brought on by the consumption of it. Consequently, these schools hold that Muslims must perform ''wudhu'' (ablution) before the next time they Salat, pray after eating camel meat. Also, some Islamic schools of thought consider it ''haram'' ( ar, حرام, 'forbidden') for a Muslim to perform ''Salat'' in places where camels lie, as it is said to be a dwelling place of the ''Devil (Islam), Shaytan'' ( ar, شيطان, 'Devil'). According to Abu Yusuf (d.798), the camel urine, urine of camel may be used for medical treatment if necessary, but according to Abū Ḥanīfah, the drinking of camel urine is discouraged. The Islamic texts contain several stories featuring camels. In the story of the people of Thamud, the Prophet Salih miraculously brings forth a ''She-Camel of God, naqat'' ( ar, ناقة, 'milch-camel') out of a rock. After the Prophet Muhammad Hegira, migrated from Mecca to Medina, he allowed Qaswa, his she-camel to roam there; the location where the camel stopped to rest determined the location where he would build his house in Medina.


Judaism

According to Jewish tradition, camel meat and milk are not kosher. Camels possess only one of the two kosher foods, kosher criteria; although they Ruminant, chew their cud, they do not possess Cloven hoof, cloven hooves: "But these you shall not eat among those that bring up the cud and those that have a cloven hoof: the camel, because it brings up its cud, but does not have a [completely] cloven hoof; it is unclean for you."


Cultural depictions

What may be the oldest carvings of camels were discovered in 2018 in Saudi Arabia. They were analysed by researchers from several scientific disciplines and, in 2021, were estimated to be 7,000 to 8,000 years old. The dating of rock art is made difficult by the lack of organic material in the carvings that may be tested, so the researchers attempting to date them tested animal bones found associated with the carvings, assessed erosion patterns, and analysed tool marks in order to determine a correct date for the creation of the sculptures. This Neolithic dating would make the carvings significantly older than Stonehenge (5,000 years old) and the Egyptian pyramids at Giza (4,500 years old) and it predates estimates for the domestication of camels. File:Shadda (detail), Karabagh region, southwest Caucasus.jpeg, Shadda (cover,detail), Karabagh region, southwest Caucasus, early 19th century File:Vessel in the Form of a Recumbent Camel with Jugs, 2015.65.15.jpg, Vessel in the form of a recumbent camel with jugs, 250 BC – 224 AD, Brooklyn Museum File:Brooklyn Museum - Maru Ragini (Dhola and Maru riding on a Camel).jpg, ''Maru Ragini'' (''Dhola and Maru Riding on a Camel)'', c. 1750, Brooklyn Museum File:Brooklyn Museum - The Magi Journeying (Les rois mages en voyage) - James Tissot - overall.jpg, ''The Magi Journeying'' (''Les rois mages en voyage'')—James Tissot, c. 1886, Brooklyn Museum File:KiplingCamel3.gif, ''How the Camel Got His Hump'' (From Rudyard Kipling's ''Just So Stories'')


Distribution and numbers

There are approximately 14 million camels alive , with 90% being dromedaries. Dromedaries alive today are List of domesticated animals, domesticated animals (mostly living in the Horn of Africa, the Sahel, Maghreb, Middle East and South Asia). The Horn region alone has the largest concentration of camels in the world, where the dromedaries constitute an Nomadic pastoralism, important part of local nomadic life. They provide nomadic people in Somalia and Ethiopia with milk, food, and transportation.Plain text version.
Over one million dromedary camels are estimated to be Australian feral camel, feral in Australia, descended from those introduced as a method of transport in the 19th and early 20th centuries. This population is growing about 8% per year; it was estimated at around 700,000 in 2008. Representatives of the Australian government have culled more than 100,000 of the animals in part because the camels use too much of the limited resources needed by sheep farmers. A small population of introduced camels, dromedaries and Bactrians, wandered through Southwestern United States after having been imported in the 19th century as part of the
U.S. Camel Corps The United States Camel Corps was a mid-19th-century experiment by the United States Army in using camels as pack animals in the Southwestern United States. Although the camels proved to be hardy and well suited to travel through the region, th ...
experiment. When the project ended, they were used as draft animals in mines and escaped or were released. Twenty-five U.S. camels were bought and exported to Canada during the Cariboo Gold Rush. The Bactrian camel is, , reduced to an estimated 1.4 million animals, most of which are domesticated. The
Wild Bactrian camel The wild Bactrian camel (''Camelus ferus'') is a critically endangered species of camel living in parts of northwestern China and southwestern Mongolia. It is closely related to the Bactrian camel (''Camelus bactrianus''). Both are large, doub ...
is a separate species and is the only truly wild (as opposed to feral) camel in the world. The wild camels are critically endangered and number approximately 1400, inhabiting the Gobi and Taklamakan Deserts in China and Mongolia.


See also

* Afghan cameleers in Australia * Australian feral camel * Camel howdah * Camel milk * Camel racing * Camel train (caravan) * Camel urine * Camel wrestling * Camelops * ''Camelus moreli'' * Dromedary * List of animals with humps * Xerocole


Citations


General and cited references


Camels and Camel Milk. Report Issued by Food and Agriculture Organization of the United Nations. (1982)
* * * *


Further reading

*


External links




Six Green Reasons to Drink Camel's Milk



The Camel as a pet

"Could Emirati camels hold the key to treating venomous snake bites?"
{{Authority control Camels, African cuisine Arab cuisine Camelids Domesticated animals Halal meat Livestock Middle Eastern cuisine Taxa named by Carl Linnaeus