Caltha appendiculata
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''Caltha appendiculata'' is a
perennial In horticulture, the term perennial ('' per-'' + '' -ennial'', "through the year") is used to differentiate a plant from shorter-lived annuals and biennials. It has thus been defined as a plant that lives more than 2 years. The term is also ...
herb Herbs are a widely distributed and widespread group of plants, excluding vegetables, with savory or aromatic properties that are used for flavoring and garnishing food, for medicinal purposes, or for fragrances. Culinary use typically distingu ...
plant found in several countries of South America.


Description

''Caltha appendiculata'' is a low (2½-7 cm high)
dioecious Dioecy ( ; ; adj. dioecious, ) is a characteristic of certain species that have distinct unisexual individuals, each producing either male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproduction is ...
, perennial herb, often growing in dense clusters over considerable areas, with thick
rhizome In botany and dendrology, a rhizome ( ) is a modified subterranean plant stem that sends out roots and Shoot (botany), shoots from its Node (botany), nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from ...
s. The strong stems are sparingly branched, somewhat elongated, and covered by the remains of old sheaths. Its thick and fleshy leaves consist of a short leafstalk and a blade of between six and ten mm long, which is ovate, elliptical or oblong in outline, has an entire margin or is split into three lobes, and usually i
retuse
at the tip. Near the midrib on the upward leaf face are two or three narrow leaflike outgrowths. This makes the leaf blade in its entirety reminiscent of an opened book. These appendages are in fact homologous with the ears at the base of the leaf blade of Northern Hemisphere ''Caltha'' species. The faintly sweet scented
actinomorphic Floral symmetry describes whether, and how, a flower, in particular its perianth, can be divided into two or more identical or mirror-image parts. Uncommonly, flowers may have no axis of symmetry at all, typically because their parts are spirall ...
solitary flowers of between one and two cm across have five spreading, lanceolate, pale yellow to creamy-green
sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 Etymology The term ''sepalum'' ...
s with a purplish margin, and which are gradually narrowing towards the tip. Male flowers have thick, grooved peduncles. There are usually nine stamens. The five ovaries do not develop seeds. Female flowers have rudimentary
staminode In botany, a staminode is an often rudimentary, sterile or abortive stamen, which means that it does not produce pollen.Jackson, Benjamin, Daydon; ''A Glossary of Botanic Terms with their Derivation and Accent''; Published by Gerald Duckworth & Co ...
s, and five to nine compressed
ovaries The ovary () is a gonad in the female reproductive system that produces ova; when released, an ovum travels through the fallopian tube/oviduct into the uterus. There is an ovary on the left and the right side of the body. The ovaries are endocr ...
, with a papillose exterior. The carpels each contain up to seven or eight
ovule In seed plants, the ovule is the structure that gives rise to and contains the female reproductive cells. It consists of three parts: the ''integument'', forming its outer layer, the ''nucellus'' (or remnant of the sporangium, megasporangium), ...
s, of which usually one or two develop into pale brown, shiny seeds. Flowers are present from late spring to summer. There are forty eight chromosomes (2n=48).


Distribution

The species occurs on the
Hermite Islands __NOTOC__ The Hermite Islands () are the islands ''Hermite'', ''Herschel'', ''Deceit'' and ''Hornos'' as well as the islets ''Maxwell'', ''Jerdán'', ''Arrecife'', ''Chanticleer'', ''Hall'', ''Deceit (islet)'', and ''Hasse'' at almost the southe ...
, the
Falklands The Falkland Islands (; ), commonly referred to as The Falklands, is an archipelago in the South Atlantic Ocean on the Patagonian Shelf. The principal islands are about east of South America's southern Patagonian coast and from Cape Du ...
,
Tierra del Fuego Tierra del Fuego (, ; Spanish for "Land of Fire", rarely also Fireland in English) is an archipelago off the southernmost tip of the South America, South American mainland, across the Strait of Magellan. The archipelago consists of the main is ...
and the Southern Andes, in trickling melt water, marshes and other wet areas, from sea level up to about 2000 m.


References

{{Taxonbar, from=Q17812106 appendiculata Plants described in 1823 Flora of Chile Flora of Argentina Flora of the Falkland Islands Dioecious plants