Callawayasaurus
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''Callawayasaurus'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of
plesiosaur The Plesiosauria (; Greek: πλησίος, ''plesios'', meaning "near to" and ''sauros'', meaning "lizard") or plesiosaurs are an order or clade of extinct Mesozoic marine reptiles, belonging to the Sauropterygia. Plesiosaurs first appea ...
from the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Elasmosauridae Elasmosauridae is an extinct family of plesiosaurs, often called elasmosaurs. They had the longest necks of the plesiosaurs and existed from the Hauterivian to the Maastrichtian stages of the Cretaceous, and represented one of the two groups of ...
. When the holotype was first described by
Samuel Paul Welles Samuel Paul Welles (November 9, 1907 – August 6, 1997) was an American palaeontologist. Welles was a research associate at the Museum of Palaeontology, University of California, Berkeley. He took part in excavations at the Placerias Quarry in ...
in 1962, it was described as ''Alzadasaurus colombiensis'' before being moved into its current genus by Kenneth Carpenter in 1999.Carpenter, K. 1999. "Revision of North American elasmosaurs from the Cretaceous of the Western Interior". ''Paludicola'' 2(2):148-173.


Etymology

''Callawayasaurus'' is named in honor of the
paleontologist Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of fossi ...
Jack M. Callaway, editor of ''Ancient Marine Reptiles'' who, as Carpenter put it, "in his brief career as a vertebrate paleontologist, did much to improve our understanding of marine reptiles." The familiar suffix, ''-saurus'' comes from the Greek ' (), meaning "lizard" or "reptile."


Description

''Callawayasaurus'' was a large plesiosaur, with a skull length of , body length of and body mass of . The nares of ''Callawayasaurus'' are elongated and positioned over the
maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. T ...
, which has 3-5 
teeth A tooth ( : teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, ...
. The
neck The neck is the part of the body on many vertebrates that connects the head with the torso. The neck supports the weight of the head and protects the nerves that carry sensory and motor information from the brain down to the rest of the body. In ...
contains 56 
vertebrae The spinal column, a defining synapomorphy shared by nearly all vertebrates, Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristi ...
which are relatively short compared to other elasmosaurids. ''Callawayasaurus''
fossils A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
have no pectoral bars; in common with other plesiosaurs such as ''
Terminonatator ''Terminonatator'' (meaning "last swimmer") is a genus of elasmosaurid plesiosaur from the Late Cretaceous of Saskatchewan, Canada. It is known from a skull and partial skeleton from a young adult, found in the Campanian-age Bearpaw Form ...
''. They also lack postaxial accessory
facets A facet is a flat surface of a geometric shape, e.g., of a cut gemstone. Facet may also refer to: Arts, entertainment, and media * ''Facets'' (album), an album by Jim Croce * ''Facets'', a 1980 album by jazz pianist Monty Alexander and his tri ...
. Another nearly complete
skeleton A skeleton is the structural frame that supports the body of an animal. There are several types of skeletons, including the exoskeleton, which is the stable outer shell of an organism, the endoskeleton, which forms the support structure inside ...
was found to be slightly more robust than the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...
specimen Specimen may refer to: Science and technology * Sample (material), a limited quantity of something which is intended to be similar to and represent a larger amount * Biological specimen or biospecimen, an organic specimen held by a biorepository f ...
. This subtle change may indicate
sexual dimorphism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most an ...
.


Distribution

The first ''Callawayasaurus'' remains were found in the Paja Formation near Leiva, Boyaca
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the ...
. The specific name for the type, "''columbiensis''", means "from Colombia". ''Callawayasaurus'' are known from the
Aptian The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ag ...
faunal stage In chronostratigraphy, a stage is a succession of rock strata laid down in a single age on the geologic timescale, which usually represents millions of years of deposition. A given stage of rock and the corresponding age of time will by convent ...
of the early
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...
period, which extended from 125 to 112 million years ago.


See also

*
List of plesiosaur genera This list of plesiosaurs is a comprehensive listing of all genera that have ever been included in the order Plesiosauria, excluding purely vernacular terms. The list includes all commonly accepted genera, but also genera that are now considered inv ...
* Timeline of plesiosaur research


References


External links


Information on plesiosaurs, including images of a ''Callawayasaurus'' skull
{{Taxonbar, from=Q2634949 Elasmosaurids Early Cretaceous plesiosaurs Early Cretaceous reptiles of South America Plesiosaurs of South America Cretaceous Colombia Fossils of Colombia Altiplano Cundiboyacense Paja Formation Fossil taxa described in 1999 Taxa named by Kenneth Carpenter Sauropterygian genera